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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe
AnimaliaArthropodaMyriapodaDiplopodaChordeumatidaCleidogonidaePseudotremia

Ellett Valley Pseudotremia Millipede
Pseudotremia cavernarum Cope, 1869

Taxonomy

Kingdom Animalia
Phylum Arthropoda
Subphylum Myriapoda
Class Diplopoda
Order Chordeumatida
Family Cleidogonidae
Genus Pseudotremia
Species Pseudotremia cavernarum Cope, 1869

Description

Diagnostic characters

Ecological class
Troglobiont
Realm
Terrestrial
Troglomorphy
Troglomorphic

Distribution

Type locality

Erhart Cave, Montgomery Co. VA

Range

Known from five counties across four states in the Appalachians karst region.

Continent

North America

Country

United States

States and provinces

Indiana, Kentucky, Tennessee, Virginia

Counties

Indiana
Crawford
Kentucky
Carter
Tennessee
Marion
Virginia
Giles, Montgomery

Karst regions

Appalachians

Specimens

Type material

Syntype ×5
Museum of Comparative Zoology, Harvard

Collections holding material

InstitutionLots
Museum of Comparative Zoology, Harvard 5
Field Museum of Natural History 1
Museum national d'Histoire naturelle, Paris 1
Virginia Polytechnic Institute and State University 1

8 cataloged lots in 4 collections. Compiled from GBIF and not an exhaustive census: many collections are not yet digitised, and holdings are added continually. Search GBIF for specimen records.

Download all 8 specimen records (CSV) Institution, catalog number, type status and preparation. No coordinates.

Conservation and management

NatureServe

G2G3 Imperiled to Vulnerable

G1G2▲G3▲G4G5

Global rank last reviewed 21 August 2002.

Subnational ranks

Virginia
S1 Critically Imperiled

Ranks looked up 23 August 2026. Conservation status changes, and a rank here is as current as that date and no more.

State and federal status

Not listed under the U.S. Endangered Species Act.

Other resources

Catalogue of Life
4PDXR
GBIF
1022563
ITIS
570424
iNaturalist
229190

References

  • Banta, A.M. 1907. The fauna of Mayfield's Cave. Carnegie Institution of Washington Publication 67: 1–114. https://www.biodiversitylibrary.org/item/60987. details
  • Causey, N.B. 1960. Speciation in North American cave millipeds. American Midland Naturalist 64: 116–122.
  • Chamberlin, R.V. & Hoffman, R.L. 1958. Checklist of the millipedes of North America. United States National Museum Bulletin 212: 1–236.
  • Cope, E.D. & Packard, A.S. 1881. The fauna of the Nickajack Cave. American Naturalist 15: 877–882.
  • Loomis, H.F. 1939. The millipeds collected in Appalachian caves by Mr. Kenneth Dearolf. Bulletin of the Museum of Comparative Zoology 86: 165–193. https://www.biodiversitylibrary.org/page/2775893. details
  • Loomis, H.F. 1943. New and epigen millipeds of the United States, with notes on some established species. Bulletin of the Museum of Comparative Zoology 92: 371–410. https://www.biodiversitylibrary.org/page/2825746. details
  • Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
  • Shear, W.A. 1972. Studies in the milliped order Chordeumida (Diplopoda): A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology 144: 151–352. https://www.biodiversitylibrary.org/page/4227660. details

This account was last revised 28 August 2026. Conservation ranks were looked up 23 August 2026.

How to cite this account

Pseudotremia cavernarum Cope, 1869. Cave Biology Data Portal, account SP-0494. https://cavebiology.org/species/pseudotremia-cavernarum/ (accessed 18 October 2026).
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Data sources and funding are listed in the Acknowledgments.

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Cave Biology Data Portal

Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

Text and data CC BY 4.0. Images all rights reserved unless stated.

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