Skip to content

Cave Biology Data Portal

Subterranean biodiversity of North America and around the globe

  • ExploreExpand
    • Species
    • Taxonomy
    • States and Provinces
    • Karst Regions
    • References
    • Caves You Can Visit
  • ResourcesExpand
    • Funding Opportunities
    • Employment Opportunities
    • Meeting and Events
    • Public Notices
    • News and Updates
  • ContributeExpand
    • Report an Observation
    • Add a Photograph or Image
    • Post an Event
    • Post an Opportunity
    • Add a Reference
    • Add a New Species
    • Report a Correction
  • AboutExpand
    • Acknowledgments
    • Data Sensitivity Policy
    • Citation and Licensing
  • ContactExpand
    • Data Requests
Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe
AnimaliaArthropodaMyriapodaDiplopodaChordeumatidaCleidogonidaePseudotremia

Pseudotremia nodosa Loomis, 1939

Taxonomy

Kingdom Animalia
Phylum Arthropoda
Subphylum Myriapoda
Class Diplopoda
Order Chordeumatida
Family Cleidogonidae
Genus Pseudotremia
Species Pseudotremia nodosa Loomis, 1939

Description

Diagnostic characters

Ecological class
Troglobiont
Realm
Terrestrial
Troglomorphy
Troglomorphic

Distribution

Type locality

English Cave, Claiborne Co. TN

Range

Known from five counties across two states in the Appalachians karst region.

Continent

North America

Country

United States

States and provinces

Tennessee, Virginia

Counties

Tennessee
Anderson, Claiborne, Hancock, Union
Virginia
Lee

Karst regions

Appalachians

Specimens

Type material

Holotype
88042 · Museum of Comparative Zoology, Harvard
Paratype
National Museum of Natural History, Smithsonian Institution

Collections holding material

InstitutionLots
Museum of Comparative Zoology, Harvard 1
National Museum of Natural History, Smithsonian Institution 1

2 cataloged lots in 2 collections. Compiled from GBIF and not an exhaustive census: many collections are not yet digitised, and holdings are added continually. Search GBIF for specimen records.

Download all 2 specimen records (CSV) Institution, catalog number, type status and preparation. No coordinates.

Conservation and management

NatureServe

G4 Apparently Secure

G1G2G3G4▲G5

Global rank last reviewed 21 August 2002.

Subnational ranks

Tennessee
S2S3 Imperiled to Vulnerable
Virginia
S4 Apparently Secure

Ranks looked up 23 August 2026. Conservation status changes, and a rank here is as current as that date and no more.

State and federal status

Not listed under the U.S. Endangered Species Act.

Other resources

Catalogue of Life
4PDYK
GBIF
1022598
ITIS
570442
iNaturalist
229209

References

  • Barr, T.C., Jr. 1965. The Pseudanophthalmus of the Appalachian Valley (Coleoptera: Carabidae). The American Midland Naturalist 73: 41–72. doi:10.2307/2423320
  • Barr, T.C., Jr. 1981. Pseudanophthalmus from Appalachian caves (Coleoptera: Carabidae): the engelhardti complex. Brimleyana 5: 37–94.
  • Chamberlin, R.V. & Hoffman, R.L. 1958. Checklist of the millipedes of North America. United States National Museum Bulletin 212: 1–236.
  • Loomis, H.F. 1939. The millipeds collected in Appalachian caves by Mr. Kenneth Dearolf. Bulletin of the Museum of Comparative Zoology 86: 165–193. https://www.biodiversitylibrary.org/page/2775893. details
  • Loomis, H.F. 1943. New and epigen millipeds of the United States, with notes on some established species. Bulletin of the Museum of Comparative Zoology 92: 371–410. https://www.biodiversitylibrary.org/page/2825746. details
  • Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
  • Niemiller, M.L. & Zigler, K.S. 2013. Patterns of cave biodiversity and endemism in the Appalachians and Interior Plateau of Tennessee, USA. PLOS One 8: e64177. doi:10.1371/journal.pone.0064177
  • Shear, W.A. 1972. Studies in the milliped order Chordeumida (Diplopoda): A revision of the family Cleidogonidae and a reclassification of the order Chordeumida in the New World. Bulletin of the Museum of Comparative Zoology 144: 151–352. https://www.biodiversitylibrary.org/page/4227660. details
  • Trumbore, L.E., Reynolds, R.J., Thomas, S.C., Orndorff, W., Cramphorn, B., Gladstone, N.S., Harden, C.W., Lewis, J.J., Milne, M.A., Ober, K.A., Shear, W.A., Stephen, C.D.R. & Zigler, K.S. 2024. Cave biodiversity of Cumberland Gap National Historical Park. Journal of Cave and Karst Studies 86: 13–24. doi:10.4311/2022LSC0116

This account was last revised 28 August 2026. Conservation ranks were looked up 23 August 2026.

How to cite this account

Pseudotremia nodosa Loomis, 1939. Cave Biology Data Portal, account SP-0526. https://cavebiology.org/species/pseudotremia-nodosa/ (accessed 18 October 2026).
Email Bluesky X Facebook

Data sources and funding are listed in the Acknowledgments.

Spotted something wrong on this page? Report a correction. Accounts are compiled from many sources and corrections from people who know the group are the most useful email we get.

Cave Biology Data Portal

Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

Text and data CC BY 4.0. Images all rights reserved unless stated.

Explore

  • Species
  • Taxonomy
  • States and Provinces
  • Karst Regions
  • References

Resources

  • Funding
  • Employment
  • Contribute

About this site

  • About Us
  • Acknowledgments
  • Contact Us
  • Data Sensitivity Policy
  • Citation and Licensing
  • Accessibility Statement
  • Privacy Policy
  • Cookies Policy
  • Terms and Conditions
  • Explore
    • Species
    • Taxonomy
    • States and Provinces
    • Karst Regions
    • References
    • Caves You Can Visit
  • Resources
    • Funding Opportunities
    • Employment Opportunities
    • Meeting and Events
    • Public Notices
    • News and Updates
  • Contribute
    • Report an Observation
    • Add a Photograph or Image
    • Post an Event
    • Post an Opportunity
    • Add a Reference
    • Add a New Species
    • Report a Correction
  • About
    • Acknowledgments
    • Data Sensitivity Policy
    • Citation and Licensing
  • Contact
    • Data Requests