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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe
AnimaliaArthropodaHexapodaInsectaColeopteraCarabidaeHorologion

Arbuckle Cave Beetle
Horologion speokites Valentine, 1932

Taxonomy

Kingdom Animalia
Phylum Arthropoda
Subphylum Hexapoda
Class Insecta
Order Coleoptera
Family Carabidae
Genus Horologion
Species Horologion speokites Valentine, 1932

Description

Diagnostic characters

Ecological class
Troglobiont
Realm
Terrestrial
Troglomorphy
Troglomorphic

Distribution

Type locality

Arbuckle Cave, Greenbrier Co. WV

Range

Known from one county in West Virginia in the Appalachians karst region.

Continent

North America

Country

United States

States and provinces

West Virginia

Counties

West Virginia
Greenbrier

Karst regions

Appalachians

Conservation and management

NatureServe

GH Possibly Extinct

G1G2G3G4G5

Global rank last reviewed 1997.

Subnational ranks

West Virginia
SH Possibly Extirpated

State and federal status

Not listed under the U.S. Endangered Species Act.

Other resources

Catalogue of Life
PK8HB
GBIF
1035676
ITIS
110216
iNaturalist
221502

References

  • Bousquet, Y. 2012. Catalogue of Geadephaga (Coleoptera: Adephaga) of America, north of Mexico. ZooKeys 245: 1–1. doi:10.3897/zookeys.245.3416
  • Erwin, T.L. & House, G.N. 1978. A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera). The Coleopterists Bulletin 32: 231–256. https://www.jstor.org/stable/3999875. details
  • Harden, C.W., Davidson, R.L., Malabad, T.E., Caterino, M.S. & Maddison, D.R. 2024. Phylogenetic systematics of the enigmatic genus Horologion Valentine, 1932 (Coleoptera, Carabidae, Trechinae, Horologionini), with description of a new species from Bath County, Virginia. Subterranean Biology 48: 1–49. doi:10.3897/subtbiol.48.114404
  • Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
  • Sisco, J.M. & Sawicki, T.R. 2023. Molecular and morphological analyses reveal a new hypogean species of amphipod in the genus Crangonyx Bate, 1859 (Crustacea: Crangonyctidae) within the floridanus species complex, from Suwannee County, Florida. Journal of Natural History 57: 21–24. doi:10.1080/00222933.2023.2247155
  • Valentine, J.M. 1932a. Horologion, a new genus of cave beetles. Annals of the Entomological Society of America 25: 1–11. doi:10.1093/aesa/25.1.1

This account was last revised 28 August 2026.

How to cite this account

Horologion speokites Valentine, 1932. Cave Biology Data Portal, account SP-0651. https://cavebiology.org/species/horologion-speokites/ (accessed 18 October 2026).
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Data sources and funding are listed in the Acknowledgments.

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Cave Biology Data Portal

Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

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