Intermediate Cave Beetle
Pseudanophthalmus intermedius (Valentine, 1931)
Taxonomy
| Kingdom | Animalia |
|---|---|
| Phylum | Arthropoda |
| Subphylum | Hexapoda |
| Class | Insecta |
| Order | Coleoptera |
| Family | Carabidae |
| Genus | Pseudanophthalmus |
| Species | Pseudanophthalmus intermedius (Valentine, 1931) |
Synonyms
- Neaphaenops intermedius Valentine, 1931 (prior combination, CoL)
Description
Diagnostic characters
- Ecological class
- Troglobiont
- Realm
- Terrestrial
- Troglomorphy
- Troglomorphic
Distribution
Type locality
Wonder Cave, Grundy Co. TN
Range
Known from three counties in Tennessee in the Interior Low Plateau karst region.
Continent
North America
Country
United States
States and provinces
Tennessee
Counties
- Tennessee
- Franklin, Grundy, Marion
Karst regions
Interior Low Plateau
Specimens
Type material
- Lectotype
- National Museum of Natural History, Smithsonian Institution
Collections holding material
| Institution | Lots |
|---|---|
| Museum of Comparative Zoology, Harvard | 1 |
| National Museum of Natural History, Smithsonian Institution | 1 |
2 cataloged lots in 2 collections. Compiled from GBIF and not an exhaustive census: many collections are not yet digitised, and holdings are added continually. Search GBIF for specimen records.
Download all 2 specimen records (CSV) Institution, catalog number, type status and preparation. No coordinates.
Conservation and management
NatureServe
G2 Imperiled
Global rank last reviewed 30 June 2005.
Subnational ranks
- Tennessee
- S2 Imperiled
Ranks looked up 23 August 2026. Conservation status changes, and a rank here is as current as that date and no more.
State and federal status
Not listed under the U.S. Endangered Species Act.
Other resources
References
- Barr, T.C., Jr. 2004. A classification and checklist of the genus Pseudanophthalmus Jeannel (Coleoptera: Carabidae: Trechinae). Special Publication 11. Virginia Museum of Natural History, Martinsville, Virginia, USA. 52 pp.
- Benito, J.B., Ober, K.A., Philips, T.K., Orndorff, W. & Niemiller, M.L. 2025. Phylogenomics and biogeography of North American trechine cave beetles (Coleoptera:Carabidae) inferred from ultraconserved elements. Evolution 79: 1622–1642. doi:10.1093/evolut/qpaf103
- Bousquet, Y. 2012. Catalogue of Geadephaga (Coleoptera: Adephaga) of America, north of Mexico. ZooKeys 245: 1–1. doi:10.3897/zookeys.245.3416
- Dixon, G.B. & Zigler, K.S. 2011. Cave-obligate biodiversity on the campus of Sewanee: The University of the South, Franklin County, Tennessee. Southeastern Naturalist 10: 251–266. https://www.jstor.org/stable/41263008. details
- Erwin, T.L. & House, G.N. 1978. A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera). The Coleopterists Bulletin 32: 231–256. https://www.jstor.org/stable/3999875. details
- Lewis, J.J. 2005a. A biological inventory of caves of the Cumberland Plateau. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 156 pp.
- Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
- Niemiller, M.L. & Zigler, K.S. 2013. Patterns of cave biodiversity and endemism in the Appalachians and Interior Plateau of Tennessee, USA. PLOS One 8: e64177. doi:10.1371/journal.pone.0064177
- Niemiller, M.L., Zigler, K.S., Hinkle, A., Stephen, C.D.R., Cramphorn, B., Higgs, J., Mann, N., Miller, B.T., Niemiller, K.D.K., Smallwood, K. & Hardy, J. 2023b. The Crystal-Wonder Cave System: a new hotspot of subterranean biodiversity in the southern Cumberland Plateau of south-central Tennessee, USA. Diversity 15: 801. doi:10.3390/d15070801
- Valentine, J.M. 1931. New cavernicole Carabidae of the subfamily Trechinae Jeannel. Journal of the Elisha Mitchell Scientific Society 46: 247–258.
- Valentine, J.M. 1932b. A classification of the genus Pseudanophthalmus Jeannel (Fam. Carabidae) with descriptions of new species and notes on distribution. Journal of the Elisha Mitchell Scientific Society 47: 261–280. https://www.jstor.org/stable/24332003. details
- Valentine, J.M. 1937. Anophthalmid beetles (fam. Carabidae) from Tennessee caves. Journal of the Elisha Mitchell Scientific Society 53: 93–100.
- Valentine, J.M. 1945. Speciation and raciation in Pseudanophthalmus (cavernicolous Carabidae). Transactions of the Connecticut Academy of Arts and Sciences 36: 631–672.
- Valentine, J.M. 1948. New Anophthalmid beetles (fam. Carabidae) from the Appalachian Region. Geological Survey of Alabama Museum Paper 27: 1–20.
This account was last revised 28 August 2026. Conservation ranks were looked up 23 August 2026.
How to cite this account
Data sources and funding are listed in the Acknowledgments.
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