Granulated Millipede
Scytonotus granulatus (Say, 1821)
Taxonomy
| Kingdom | Animalia |
|---|---|
| Phylum | Arthropoda |
| Subphylum | Myriapoda |
| Class | Diplopoda |
| Order | Polydesmida |
| Family | Polydesmidae |
| Genus | Scytonotus |
| Species | Scytonotus granulatus (Say, 1821) |
Description
Diagnostic characters
- Ecological class
- Trogloxene
- Realm
- Terrestrial
Distribution
Range
Known from nine counties across five states.
Continent
North America
Country
United States
States and provinces
Alabama, Indiana, Kentucky, Pennsylvania, Tennessee
Counties
- Alabama
- Jackson
- Indiana
- Lawrence, Monroe
- Kentucky
- Barren, Pulaski
- Pennsylvania
- Berks
- Tennessee
- Rutherford, Van Buren, White
Conservation and management
NatureServe
G5 Secure
National ranks
- United States
- NNR
- Canada
- N5 Secure
Subnational ranks
- Indiana
- S4 Apparently Secure
- Virginia
- S5 Secure
- Ontario
- S5 Secure
State and federal status
Not listed under the U.S. Endangered Species Act
References
- Banta, A.M. 1907. The fauna of Mayfield's Cave. Carnegie Institution of Washington Publication 67: 1–114. https://www.biodiversitylibrary.org/item/60987.
- Barr, T.C., Jr. & Kuehne, R.A. 1971. Ecological studies in the Mammoth Cave system of Kentucky. II. The Ecosystem. Annales de Speleologie 26: 47–96. doi:10.5038/1827-806X.3.1.10
- Causey, N.B. 1960. Speciation in North American cave millipeds. American Midland Naturalist 64: 116–122.
- Chamberlin, R.V. & Hoffman, R.L. 1958. Checklist of the millipedes of North America. United States National Museum Bulletin 212: 1–236.
- Lewis, J.J. 2004. Interstate 66 karst faunal study. Environmental Impact Statement. 17 pp.
- Lewis, J.J. 2005. A biological inventory of caves of the Cumberland Plateau. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 156 pp.
- Lewis, J.J. & Lewis, S.L. 2007. A biological reconnaissance of selected caves in the Highland Rim area of central Tennessee. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 42 pp.
- Lewis, J.J., Whitaker, J.O. & Krantz, G.W. 2010. A biological reconnaissance of the invertebrate fauna of twelve Tennessee caves with notes on the guanophilic mites of the genus Macrocheles. Journal of the Tennessee Academy of Science 85: 53–61.
- Loomis, H.F. 1939. The millipeds collected in Appalachian caves by Mr. Kenneth Dearolf. Bulletin of the Museum of Comparative Zoology 86: 165–193. https://www.biodiversitylibrary.org/page/2775893.
- Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
- Peck, S.B. 1988. A review of the cave fauna of Canada, and the composition and ecology of the invertebrate fauna of caves and mines in Ontario. Canadian Journal of Zoology 66: 1197–1213. doi:10.1139/z88-176
- Peck, S.B. 1989. The cave fauna of Alabama. Part I. The terrestrial invertebrates (excluding insects). National Speleological Society Bulletin 51: 11–33.
How to cite this account
Scytonotus granulatus (Say, 1821). Cave Biology Data Portal, account SP-2040. https://cavebiology.org/species/scytonotus-granulatus/ (accessed 7 September 2026).
Data sources and funding are listed in the Acknowledgments.
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