Greenhouse Millipede
Oxidus gracilis (Koch, 1847)

Taxonomy

Description

Diagnostic characters

Ecological class

Troglophile

Realm

Terrestrial

Distribution

Range

Known from 32 counties across eight states.

Continent

North America

Country

United States

States and provinces

Alabama, Florida, Georgia, Kentucky, Oklahoma, Pennsylvania, Tennessee, Virginia

Counties

Alabama

Jackson, Morgan

Florida

Alachua

Georgia

Bartow, Catoosa, Chattooga, Dade, Decatur, Floyd, Grady, Polk, Walker, Washington

Kentucky

Larue, Pulaski

Oklahoma

Adair, Delaware, Sequoyah

Pennsylvania

Dauphin

Tennessee

Bledsoe, Claiborne, Cumberland, Franklin, Grundy, Marion, Montgomery, Overton, Pickett, Rutherford, Warren, White

Virginia

Lee

Conservation and management

NatureServe

G5 Secure

National ranks

United States

NNA

Canada

NNA

Subnational ranks

Montana

SNA Not Applicable

Indiana

SNA Not Applicable

Oklahoma

SNA Not Applicable

Virginia

SNA Not Applicable

Ontario

SNA Not Applicable

British Columbia

SNA Not Applicable

Nova Scotia

SNA Not Applicable

Newfoundland and Labrador

SNA Not Applicable

State and federal status

Not listed under the U.S. Endangered Species Act

References

• Fenolio, D.B. 2012. 2012 Ozark aquatic subterranean wildlife surveys. Report to Oklahoma Biological Survey. Atlanta Botanical Garden, Atlanta, Georgia, USA. 3 pp.
• Fenolio, D.B. 2014. 2014 Ozark aquatic subterranean wildlife surveys. Report to Oklahoma Biological Survey. San Antonio Zoo, San Antonio, Texas, USA. 21 pp.
• Fenolio, D.B., Niemiller, M.L., Slay, M.E. & Carter, E.T. 2017. Biological inventories of caves in the Ozarks of northeastern Oklahoma: December 2017 field report. Report to Oklahoma Biological Survey. San Antonio Zoo, San Antonio, Texas, USA. 17 pp.
• Fenolio, D.B. 2017. 2017 Ozark aquatic subterranean wildlife surveys. Report to Oklahoma Biological Survey. San Antonio Zoo, San Antonio, Texas, USA. 15 pp.
• Fenolio, D.B., Niemiller, M.L., Slay, M.E. & Carter, E.T. 2018. Biological inventories of caves in the Ozarks of northeastern Oklahoma: December 2018 field report. Report to Oklahoma Biological Survey. San Antonio Zoo, San Antonio, Texas, USA. 25 pp.
• Fenolio, D.B., Slay, M.E. & Niemiller, M.L. 2023. Biological inventories of caves in the Ozarks of northeastern Oklahoma: December 2023 field report. Report to Oklahoma Biological Survey. San Antonio Zoo, San Antonio, Texas, USA. 24 pp.
• Fenolio, D.B., Slay, M.E. & Niemiller, M.L. 2023. Biological inventories of caves in the Ozarks of northeastern Oklahoma: December 2022 field report. Report to Oklahoma Biological Survey. San Antonio Zoo, San Antonio, Texas, USA. 24 pp.
• Hobbs, III., H.H. 1994. Assessment of the ecological resources of the caves of Russell Cave National Monument, Jackson County, Alabama and of selected caves at the Lookout Mountain Unit of Chickamauga-Chattanooga National Military Park, Dade County, Georgia and Hamilton County, Tennessee. 200 pp.
• Lewis, J.J. 2004. Interstate 66 karst faunal study. Environmental Impact Statement. 17 pp.
• Lewis, J.J. 2005. A biological inventory of caves of the Cumberland Plateau. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 156 pp.
• Lewis, J.J. & Lewis, S.L. 2007. A biological reconnaissance of selected caves in the Highland Rim area of central Tennessee. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 42 pp.
• Lewis, J.J., Whitaker, J.O. & Krantz, G.W. 2010. A biological reconnaissance of the invertebrate fauna of twelve Tennessee caves with notes on the guanophilic mites of the genus Macrocheles. Journal of the Tennessee Academy of Science 85: 53–61.
• Loomis, H.F. 1939. The millipeds collected in Appalachian caves by Mr. Kenneth Dearolf. Bulletin of the Museum of Comparative Zoology 86: 165–193. https://www.biodiversitylibrary.org/page/2775893.
• Mays, J. 2018. Inventory of Florida's cave fauna with a focus on terrestrial invertebrate species. Progress Report for Florida Park Service. Florida Fish & Wildlife Conservation Commission, Tallahassee, Florida, USA. 9 pp.
• Niemiller, M.L., Maxwell, E. & Higgs, J.P. 2024. A biological inventory of Sinking Spring Cave at Abraham Lincoln Birthplace National Historical Park, Larue County, Kentucky, USA. U.S. National Park Service, Hodgenville, Kentucky, USA. 21 pp.
• Niemiller, M.L., Anderson, C.L., Maxwell, E.C., Midgette, H., Quering, C., Richards, M., Wilson, K. & Wirwa, N. 2025. A noteworthy range extension of the federally endangered Alabama Cave Shrimp (Palaemonias alabamae) south of the Tennessee River in northern Alabama. Southeastern Naturalist 24: N47-N58. doi:10.1656/058.024.0403
• Peck, S.B. 1973. A systematic revision and the evolutionary biology of the Ptomaphagus (Adelops) beetles of North America (Coleoptera; Leiodidae; Catopinae), with emphasis on cave-inhabiting species. Bulletin of the Museum of Comparative Zoology 145: 29–162. https://www.biodiversitylibrary.org/page/4336298.
• Reeves, W.K., Jensen, J.B. & Ozier, J.C. 2000. New faunal and fungal records from caves in Georgia, USA. Journal of Cave and Karst Studies 62: 169–179. https://caves.org/wp-content/uploads/Publications/JCKS/v62/v62n3-Reeves.pdf.
• Trumbore, L.E., Reynolds, R.J., Thomas, S.C., Orndorff, W., Cramphorn, B., Gladstone, N.S., Harden, C.W., Lewis, J.J., Milne, M.A., Ober, K.A., Shear, W.A., Stephen, C.D.R. & Zigler, K.S. 2024. Cave biodiversity of Cumberland Gap National Historical Park. Journal of Cave and Karst Studies 86: 13–24. doi:10.4311/2022LSC0116
• Zigler, K.S., Niemiller, M.L., Stephen, C.D.R., Ayala, B.N., Milne, M.A., Gladstone, N.S., Engel, A.S., Jensen, J.B., Camp, C.D., Ozier, J.C. & Cressler, A. 2020. Biodiversity from caves and other subterranean habitats of Georgia, USA. Journal of Cave and Karst Studies 82: 125–167. doi:10.4311/2019LSC0125

How to cite this account

Data sources and funding are listed in the Acknowledgments.

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