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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe
AnimaliaMolluscaGastropodaBasommatophoraCarychiidaeCarychium

Cave thorn
Carychium stygium Call, 1897

Taxonomy

Kingdom Animalia
Phylum Mollusca
Class Gastropoda
Order Basommatophora
Family Carychiidae
Genus Carychium
Species Carychium stygium Call, 1897

Description

Diagnostic characters

Ecological class
Troglobiont
Realm
Terrestrial
Troglomorphy
Troglomorphic

Distribution

Type locality

Mammoth Cave, Edmonson Co. KY

Range

Known from 24 counties across three states across two karst regions.

Continent

North America

Country

United States

States and provinces

Alabama, Kentucky, Tennessee

Counties

Kentucky
Allen, Barren, Edmonson, Edmonson Co, Hart, Jackson, Pulaski, Rockcastle, Simpson, Warren
Tennessee
Anderson, Coffee, Franklin, Grundy, Jackson, Knox, Loudon, Montgomery, Overton, Roane, Sullivan, Sumner, Van Buren, Warren

Karst regions

Appalachians, Interior Low Plateau

Genetics

Molecular resources

MarkerMoleculeSequences LengthExemplar
COI mitochondrial 6 616-649 bp HQ171569.1

6 sequences, retrieved 23 August 2026. Counts change as sequences are deposited. Search GenBank.

Download all 6 sequence records (CSV) Accession, marker, molecule type, length, GenBank definition and a direct link.

Specimens

Type material

Paratype
46538 · Carnegie Museum of Natural History

Collections holding material

InstitutionLots
Field Museum of Natural History 53
National Museum of Natural History, Smithsonian Institution 4
Senckenberg Forschungsinstitut und Naturmuseum Frankfurt 4
Delaware Museum of Nature and Science 2
Carnegie Museum of Natural History 2
Academy of Natural Sciences of Drexel University 2
and 5 other collections 5

72 cataloged lots in 11 collections. Compiled from GBIF and not an exhaustive census: many collections are not yet digitised, and holdings are added continually. Search GBIF for specimen records.

Download all 72 specimen records (CSV) Institution, catalog number, type status and preparation. No coordinates.

Conservation and management

NatureServe

G3 Vulnerable

G1G2G3▲G4G5

Global rank last reviewed 9 November 2006.

Subnational ranks

Kentucky
S3 Vulnerable
Tennessee
S2 Imperiled

Ranks looked up 23 August 2026. Conservation status changes, and a rank here is as current as that date and no more.

State and federal status

Not listed under the U.S. Endangered Species Act.

Other resources

Catalogue of Life
RHZV
GBIF
2291065
ITIS
76754
iNaturalist
216161
NCBI Taxonomy
884962

References

  • Bailey, V. 1933. Cave Life of Kentucky. US Department of Agriculture, Bureau of Biological Survey, Washington, DC, USA. 256 pp.
  • Barr, T.C., Jr. & Kuehne, R.A. 1971. Ecological studies in the Mammoth Cave system of Kentucky. II. The Ecosystem. Annales de Speleologie 26: 47–96. doi:10.5038/1827-806X.3.1.10
  • Giovannoli, L. 1933. Invertebrate life of Mammoth and other neighboring caves. In: Bailey, V. (ed), Cave Life of Kentucky, pp. 600–623.
  • Gladstone, N.S., Carter, E.T., McKinney, M.L. & Niemiller, M.L. 2018a. Status and distribution of the cave-obligate land snails in the Appalachians and Interior Low Plateau of the eastern United States. American Malacological Bulletin 36: 62–78.
  • Harker, D.F., Jr. & Barr, T.C., Jr. 1980. Caves and associated fauna of the Western Kentucky coal field, Volume 2. Kentucky Nature Preserves Commission, Frankfurt, Kentucky, USA. 114 pp.
  • Hubricht, L. 1962. New species of Helicodiscus from the eastern United States. The Nautilus 75: 102–107. https://www.biodiversitylibrary.org/page/8283022. details
  • Hubricht, L. 1964. Land snails from the caves of Kentucky, Tennessee and Alabama. National Speleological Society Bulletin 26: 33–36.
  • Hubricht, L. 1973. The land snails of Tennessee. Sterkiana 49: 11–17.
  • Lewis, J.J. 2004. Interstate 66 karst faunal study. Environmental Impact Statement. 17 pp.
  • Lewis, J.J. 2005a. A biological inventory of caves of the Cumberland Plateau. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 156 pp.
  • Lewis, J.J. & Lewis, S.L. 2005. Inventory of the subterranean fauna of Tims Ford State Park, Franklin County, Tennessee. Tennesee Natural Heritage Program, Nashville, Tennessee, USA. 28 pp.
  • Lewis, J.J. & Lewis, S.L. 2007. A biological reconnaissance of selected caves in the Highland Rim area of central Tennessee. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 42 pp.
  • Lewis, J.J., Whitaker, J.O. & Krantz, G.W. 2010. A biological reconnaissance of the invertebrate fauna of twelve Tennessee caves with notes on the guanophilic mites of the genus Macrocheles. Journal of the Tennessee Academy of Science 85: 53–61.
  • Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
  • Niemiller, M.L. & Zigler, K.S. 2013. Patterns of cave biodiversity and endemism in the Appalachians and Interior Plateau of Tennessee, USA. PLOS One 8: e64177. doi:10.1371/journal.pone.0064177
  • Niemiller, M.L., Zigler, K.S., Hinkle, A., Stephen, C.D.R., Cramphorn, B., Higgs, J., Mann, N., Miller, B.T., Niemiller, K.D.K., Smallwood, K. & Hardy, J. 2023b. The Crystal-Wonder Cave System: a new hotspot of subterranean biodiversity in the southern Cumberland Plateau of south-central Tennessee, USA. Diversity 15: 801. doi:10.3390/d15070801

This account was last revised 28 August 2026. Conservation ranks were looked up 23 August 2026.

How to cite this account

Carychium stygium Call, 1897. Cave Biology Data Portal, account SP-1375. https://cavebiology.org/species/carychium-stygium/ (accessed 18 October 2026).
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Data sources and funding are listed in the Acknowledgments.

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Cave Biology Data Portal

Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

Text and data CC BY 4.0. Images all rights reserved unless stated.

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