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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe
AnimaliaChordataMammaliaChiropteraVespertilionidaeMyotis

Cluster Bat
Myotis sodalis Miller & G. M. Allen, 1928

Taxonomy

Kingdom Animalia
Phylum Chordata
Class Mammalia
Order Chiroptera
Family Vespertilionidae
Genus Myotis
Species Myotis sodalis Miller & G. M. Allen, 1928

Description

Diagnostic characters

Ecological class
Trogloxene
Realm
Terrestrial

Distribution

Range

Known from 60 counties across nine states.

Continent

North America

Country

United States

States and provinces

Alabama, Arkansas, Florida, Georgia, Illinois, Kentucky, Pennsylvania, Tennessee, Virginia

Counties

Alabama
Jackson, Marshall, Morgan, Shelby
Arkansas
Baxter, Benton, Garland, Independence, Madison, Newton, Searcy, Stone, Washington
Florida
Jackson
Georgia
Chattooga, Dade
Illinois
Alexander, Hardin, Jackson, Monroe, Pope
Kentucky
Adair, Barren, Breckinridge, Caldwell, Carter, Christian, Edmonson, Elliott, Estill, Hardin, Hart, Jackson, Jessamine, Lee, Letcher, Livingston, Meade, Menifee, Powell, Pulaski, Rockcastle, Taylor, Trigg, Warren, Wayne, Whitley
Pennsylvania
Armstrong, Blair, Centre, Luzerne, Mifflin, Somerset
Tennessee
Blount, Campbell, Fentress, Montgomery, Van Buren, White
Virginia
Lee

Conservation and management

IUCN Red List

NT Near Threatened

EXEWCRENVUNT▲LC

Assessed in 2016.

NatureServe

G2 Imperiled

G1G2▲G3G4G5

National ranks

United States
N2 Imperiled

Subnational ranks

Tennessee
S1 Critically Imperiled
Arkansas
S1 Critically Imperiled
Michigan
S1 Critically Imperiled
Alabama
S2 Imperiled
West Virginia
S1 Critically Imperiled
Vermont
S1 Critically Imperiled
Missouri
S1 Critically Imperiled
NJ
S1 Critically Imperiled
Ohio
S1 Critically Imperiled
Oklahoma
S1 Critically Imperiled
Virginia
S1 Critically Imperiled
Maryland
S1 Critically Imperiled
Indiana
S1 Critically Imperiled
Kentucky
S1 Critically Imperiled
Georgia
S1 Critically Imperiled
Iowa
S1 Critically Imperiled
Illinois
S1 Critically Imperiled
Connecticut
SHN (non-breeding)
Mississippi
S1B Critically Imperiled (breeding)
Pennsylvania
S1 Critically Imperiled
South Carolina
SU Unrankable
Massachusetts
SH Possibly Extirpated
New York
S1 Critically Imperiled
North Carolina
S1 Critically Imperiled

State and federal status

Endangered under the U.S. Endangered Species Act listed 1967

References

  • Bailey, V. 1933. Cave Life of Kentucky. US Department of Agriculture, Bureau of Biological Survey, Washington, DC, USA. 256 pp.
  • Barr, T.C., Jr. & Kuehne, R.A. 1971. Ecological studies in the Mammoth Cave system of Kentucky. II. The Ecosystem. Annales de Speleologie 26: 47–96. doi:10.5038/1827-806X.3.1.10
  • Beck, B.F. 1980. An Introduction to Caves and Cave Exploring in Georgia. Georgia Department of Natural Resources, Atlanta, Georgia, USA. 43 pp.
  • Cassidy, J.J., O'Hagan, G.G., Sanders, J.S. & Harvey, M.J. 1979. Swarming of bats at two southern Ozark caves. Proceedings of the Tennessee Academy of Science 54: 73.
  • Collins, J. & Sanders, C. 2016. A summary of bat-friendly gates and applications in Pennsylvania. In: Butchkoski, C.M., Reeder, D.M., Turner, G.G. & Whidden, H.P. (eds), Conservation and Ecology of Pennsylvania's Bats, pp. 201–208. The Pennsylvania Academy of Science, East Stroudsburg, Pennsylvania, USA.
  • Dusi, J.L. 1976. Mammals. In: Boshung, H. (ed), Endangered and threatened plants and animals of Alabama. Alabama Museum of Natural History Bulletin 2, pp. 88–92. Alabama Museum of Natural History, University of Alabama, Tuscaloosa, Alabama, USA.
  • Franz, R., Bauer, J. & Morris, T. 1994. Review of biologically significant caves and their faunas in Florida and south Georgia. Brimleyana 20: 1–109. https://www.biodiversitylibrary.org/part/229726.
  • Giovannoli, L. 1933. Invertebrate life of Mammoth and other neighboring caves. In: Bailey, V. (ed), Cave Life of Kentucky, pp. 600–623.
  • Godwin, J.C. 2000. Reassessment of the historical and search for new localities of the Tennessee cave salamander (Gyrinophilus palleucus) in Alabama. Alabama Department of Conservation and Natural Resources, Montgomery, Alabama, USA. 15 pp.
  • Harker, D.F., Jr. & Barr, T.C., Jr. 1979. Caves and associated fauna of Eastern Kentucky, Volume 3. Kentucky Nature Preserves Commission, Frankfurt, Kentucky, USA. 130 pp.
  • Harker, D.F., Jr. & Barr, T.C., Jr. 1980. Caves and associated fauna of the Western Kentucky coal field, Volume 2. Kentucky Nature Preserves Commission, Frankfurt, Kentucky, USA. 114 pp.
  • Hart, J.A. & Brenner, F.J. 2016. The history of bat research in Pennsylvania. In: Butchkoski, C.M., Reeder, D.M., Turner, G.G. & Whidden, H.P. (eds), Conservation and Ecology of Pennsylvania's Bats, pp. 49–61. The Pennsylvania Academy of Science, East Stroudsburg, Pennsylvania, USA.
  • Hobbs, III., H.H. 1985. A description of Bat Cave and the Saltpetre-Moon Cave system, Carter County, Kentucky. In: Hacker, C., Vore, D. & Vore, D. (eds), 1985 NSS Convention Guidebook no. 25: The Caves of South Eastern Kentucky, pp. 28–34. Ohio Valley Region, National Speleological Society.
  • Hudson, M.K. 2017. Indiana Myotis Myotis sodalis (Miller and Allen). In: Shelton-Nix, E. (ed), Alabama Wildlife, Volume 5, pp. 334–335. The University of Alabama Press, Tuscaloosa, Alabama, USA.
  • Humphrey, S.R. 1992. Indiana Bat Myotis sodalis. In: Humphrey, S.R. (ed), Rare and Endangered Biota of Florida. Volume I. Mammals, pp. 54–62. University Press of Florida, Gainesville, Florida, USA.
  • Jandebeur, T.S. 1999. Caves and springs of north Alabama in social context. Journal of the Alabama Academy of Science 70: 108–122.
  • Johnson, J.S., Scafini, M.R., Sewall, B.J. & Turner, G.G. 2016. Hibernating bat species in Pennsylvania use colder winter habitats following the arrival of white-nose syndrome. In: Butchkoski, C.M., Reeder, D.M., Turner, G.G. & Whidden, H.P. (eds), Conservation and Ecology of Pennsylvania's Bats, pp. 181–199. The Pennsylvania Academy of Science, East Stroudsburg, Pennsylvania, USA.
  • Johnson, J.S., Sharp, N.W., Monarchino, M.N., Lilley, T.M. & Edelman, A.J. 2021. No sign of infection in free-ranging Myotis austroriparius hibernating in the presence of Pseudogymnoascus destructans in Alabama. Southeastern Naturalist 20: 20–28. doi:10.1656/058.020.0102
  • Lamb, J. & Wyckoff, G. 2010. White-nose syndrome monitoring and response report for Tennessee. Tennessee Bat Working Group. 24 pp.
  • Lewis, J.J. 2005. A biological inventory of caves of the Cumberland Plateau. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 156 pp.
  • Lewis, J.J. & Lewis, S.L. 2005. Inventory of the subterranean fauna of Tims Ford State Park, Franklin County, Tennessee. Tennesee Natural Heritage Program, Nashville, Tennessee, USA. 28 pp.
  • Lewis, J.J. & Lewis, S.L. 2007. A biological reconnaissance of selected caves in the Highland Rim area of central Tennessee. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 42 pp.
  • Linzey, D.W. 1995. Mammals of Great Smoky Mountains National Park - 1995 update. Journal of the Elisha Mitchell Scientific Society 111: 1–81.
  • Linzey, D.W. 2016. Mammals of Great Smoky Mountains National Park: 2016 revision. Southeastern Naturalist 15, Monograph 8: 1–93.
  • McDaniel, V.R. & Gardner, J.E. 1977. Cave fauna of Arkansas: vertebrate taxa. Journal of the Arkansas Academy of Science 31: 68–71. https://scholarworks.uark.edu/jaas/vol31/iss1/22.
  • McGregor, S.W., Rheams, K.F., O'Neil, P.E., Moser, P.H. & Blackwood, R. 1994. Biological, geological, and hydrological investigations in Bobcat, Matthews, and Shelta caves and other selected caves in north Alabama. Geological Survey of Alabama, Tuscaloosa, Alabama, USA. 226 pp.
  • McGregor, S.W., O'Neil, P.E., Rheams, K.F., Moser, P.H. & Blackwood, R. 1997. Biological, geological, and hydrological investigations in Bobcat, Matthews, and Shelta caves and other selected caves in north Alabama. Geological Survey of Alabama Bulletin 166: 1–198.
  • McLaren, S.B. & Wible, J.R. 2016. Pennsylvania bats preserved as museum specimens: a storehouse of information. In: Butchkoski, C.M., Reeder, D.M., Turner, G.G. & Whidden, H.P. (eds), Conservation and Ecology of Pennsylvania's Bats, pp. 63–74. The Pennsylvania Academy of Science, East Stroudsburg, Pennsylvania, USA.
  • Nolfi, D.C. 2009. Review and assessment of known cavernicoles and rare epigean biology of karst and caves, Great Smoky Mountains National Park. Proceedings of the 15th International Congress of Speleology, Kerrville, Texas, USA, Volume 3: 1323–1328.
  • Peck, S.B. 1973. A systematic revision and the evolutionary biology of the Ptomaphagus (Adelops) beetles of North America (Coleoptera; Leiodidae; Catopinae), with emphasis on cave-inhabiting species. Bulletin of the Museum of Comparative Zoology 145: 29–162. https://www.biodiversitylibrary.org/page/4336298.
  • Poulson, T.L. 1961. Cave adaptation in amblyopsid cavefishes. University of Michigan, Ann Arbor, Michigan, USA. 185 pp. doi:10.7302/9847
  • Trumbore, L.E., Reynolds, R.J., Thomas, S.C., Orndorff, W., Cramphorn, B., Gladstone, N.S., Harden, C.W., Lewis, J.J., Milne, M.A., Ober, K.A., Shear, W.A., Stephen, C.D.R. & Zigler, K.S. 2024. Cave biodiversity of Cumberland Gap National Historical Park. Journal of Cave and Karst Studies 86: 13–24. doi:10.4311/2022LSC0116
  • Tuttle, M.D. 1977. Gating as a means of protecting cave dwelling bats. 1976 National Cave Management Symposium Proceedings: 77–82.
  • Wallace, R.L. 1984. Biological survey report of Great Smoky Mountains National Park caves. Great Smoky Mountains National Park, U.S. National Park Service, Gatlinburg, Tennessee, USA. 33 pp.
  • Wallace, R.L. 1990. Biological survey report of the Great Smoky Mountains National Park caves. Great Smoky Mountains National Park, U.S. National Park Service, Gatlinburg, Tennessee, USA. 34 pp.
  • Webb, D.W., Taylor, S.J. & Krejca, J.K. 1993. The biological resources of Illinois caves and other subterranean environments. Illinois Natural History Survey, Champaign, Illinois, USA. 143 pp.
  • Zigler, K.S., Niemiller, M.L., Stephen, C.D.R., Ayala, B.N., Milne, M.A., Gladstone, N.S., Engel, A.S., Jensen, J.B., Camp, C.D., Ozier, J.C. & Cressler, A. 2020. Biodiversity from caves and other subterranean habitats of Georgia, USA. Journal of Cave and Karst Studies 82: 125–167. doi:10.4311/2019LSC0125

How to cite this account

Myotis sodalis Miller & G. M. Allen, 1928. Cave Biology Data Portal, account SP-2690. https://cavebiology.org/species/myotis-sodalis/ (accessed 7 September 2026).

Data sources and funding are listed in the Acknowledgments.

Spotted something wrong on this page? Report a correction. Accounts are compiled from many sources and corrections from people who know the group are the most useful email we get.

Cave Biology Data Portal

Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

Text and data CC BY 4.0. Images all rights reserved unless stated.

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