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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe
AnimaliaArthropodaHexapodaInsectaColeopteraCarabidaePseudanophthalmus

Beakley's Cave Beetle
Pseudanophthalmus beakleyi Valentine, 1937

Taxonomy

Kingdom Animalia
Phylum Arthropoda
Subphylum Hexapoda
Class Insecta
Order Coleoptera
Family Carabidae
Genus Pseudanophthalmus
Species Pseudanophthalmus beakleyi Valentine, 1937

Synonyms

  • Pseudanophthalmus lupus Barr, 1959 (subjective synonym, CoL)
  • Pseudanophthalmus robustus beakleyi (subjective synonym, CoL)
  • Pseudanophthalmus robustus lupus (subjective synonym, CoL)
  • Pseudanophthalmus beaklei Valentine, 1937 (subjective synonym, CoL)

Description

Diagnostic characters

Ecological class
Troglobiont
Realm
Terrestrial
Troglomorphy
Troglomorphic

Distribution

Type locality

Bunkum Cave, Pickett Co. TN

Range

Known from six counties across two states in the Interior Low Plateau karst region.

Continent

North America

Country

United States

States and provinces

Kentucky, Tennessee

Counties

Kentucky
Clinton, McCreary, Wayne
Tennessee
Fentress, Overton, Pickett

Karst regions

Interior Low Plateau

Conservation and management

NatureServe

G3G4 Vulnerable to Apparently Secure

G1G2G3▲G4▲G5

Global rank last reviewed 4 April 2008.

Subnational ranks

Kentucky
S3 Vulnerable
Tennessee
S3 Vulnerable

Ranks looked up 23 August 2026. Conservation status changes, and a rank here is as current as that date and no more.

State and federal status

Not listed under the U.S. Endangered Species Act.

Other resources

Catalogue of Life
6WCJR
GBIF
1035099
ITIS
931980
iNaturalist
228933

References

  • Barr, T.C., Jr. 1959a. New cave beetles (Carabidae, Trechini) from Tennessee and Kentucky. Journal of the Tennessee Academy of Science 34: 5–30.
  • Barr, T.C., Jr. 1962. The robustus group in the genus Pseudanophthalmus (Coleoptera: Carabidae: Trechini). The Coleopterists Bulletin 16: 109–118. https://www.jstor.org/stable/3998965. details
  • Barr, T.C., Jr. 2004. A classification and checklist of the genus Pseudanophthalmus Jeannel (Coleoptera: Carabidae: Trechinae). Special Publication 11. Virginia Museum of Natural History, Martinsville, Virginia, USA. 52 pp.
  • Benito, J.B., Ober, K.A., Philips, T.K., Orndorff, W. & Niemiller, M.L. 2025. Phylogenomics and biogeography of North American trechine cave beetles (Coleoptera:Carabidae) inferred from ultraconserved elements. Evolution 79: 1622–1642. doi:10.1093/evolut/qpaf103
  • Bousquet, Y. 2012. Catalogue of Geadephaga (Coleoptera: Adephaga) of America, north of Mexico. ZooKeys 245: 1–1. doi:10.3897/zookeys.245.3416
  • Erwin, T.L. & House, G.N. 1978. A catalogue of the primary types of Carabidae (incl. Cicindelinae) in the collections of the United States National Museum of Natural History (USNM) (Coleoptera). The Coleopterists Bulletin 32: 231–256. https://www.jstor.org/stable/3999875. details
  • Harker, D.F., Jr. & Barr, T.C., Jr. 1979. Caves and associated fauna of Eastern Kentucky, Volume 3. Kentucky Nature Preserves Commission, Frankfurt, Kentucky, USA. 130 pp.
  • Lewis, J.J. 2005a. A biological inventory of caves of the Cumberland Plateau. Tennessee Chapter Cave Program, The Nature Conservancy, Nashville, Tennessee, USA. 156 pp.
  • Lewis, J.J. 2009b. Eight new cavernicolous species of the milliped genus Pseudotremia (Diplopoda: Chordeumatida: Cleidogonidae). In: Roble, S.M. & Mithcell, J.C. (eds), A Lifetime of Contributions to Myriapodology and the Natural History of Virginia: A Festschrift in Honor of Richard L. Hoffman’s 80th Birthday. Virginia Museum of Natural History Special Publication No. 16, pp. 171–186. Virginia Museum of Natural History, Martinsville, Virginia, USA.
  • Nicholas, G. 1960. Checklist of macroscopic troglobitic organisms of the United States. The American Midland Naturalist 64: 123–160. doi:10.2307/2422897
  • Niemiller, M.L. & Zigler, K.S. 2013. Patterns of cave biodiversity and endemism in the Appalachians and Interior Plateau of Tennessee, USA. PLOS One 8: e64177. doi:10.1371/journal.pone.0064177
  • Valentine, J.M. 1937. Anophthalmid beetles (fam. Carabidae) from Tennessee caves. Journal of the Elisha Mitchell Scientific Society 53: 93–100.

This account was last revised 28 August 2026. Conservation ranks were looked up 23 August 2026.

How to cite this account

Pseudanophthalmus beakleyi Valentine, 1937. Cave Biology Data Portal, account SP-0666. https://cavebiology.org/species/pseudanophthalmus-beakleyi/ (accessed 18 October 2026).
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Data sources and funding are listed in the Acknowledgments.

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Cave Biology Data Portal

Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

Text and data CC BY 4.0. Images all rights reserved unless stated.

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