Pennsylvania
40 species recorded, 21 of them obligate subterranean

40 species 21 obligate subterranean 9 ranked G1 or G2 3 federally listed 3 known from here only 28 counties with a record

3 of these have been recorded nowhere else, which is usual for subterranean fauna: a species is often known from a single locality.

By county

35 of 40 species carry a county. The remaining 5 are recorded to the state only.

Adams 1 species
Armstrong 4 species
Bedford 1 species
Berks 7 species
Berkshire 1 species
Blair 4 species
Bucks 6 species
Cameron 1 species
Center 1 species
Centre 8 species
Chester 1 species
Clinton 2 species
Cumberland 2 species
Dauphin 6 species
Fayette 5 species
Franklin 5 species
Huntingdon 2 species
Lancaster 6 species
Lebanon 1 species
Lehigh 1 species
Luzerne 1 species
Lycoming 5 species
Mifflin 16 species
Monroe 1 species
Montgomery 1 species
Somerset 6 species
Westmoreland 1 species
York 3 species

County is the finest resolution published. Coordinates are withheld as a matter of policy, not because they are missing.

What is here

Amphipoda: 12 species
Chiroptera: 7 species
Polydesmida: 5 species
Araneae: 4 species
Isopoda: 3 species
Tricladida: 2 species
Entomobryomorpha: 2 species
Diptera: 2 species
Coleoptera: 1 species
Chordeumatida: 1 species
Poduromorpha: 1 species

How they use caves

Stygobiont: 14 species
Trogloxene: 10 species
Troglobiont: 7 species
Troglophile: 5 species
Stygophile: 3 species
Accidental: 1 species

What material exists

Museum material: 18 of 40 species
Published sequences: 6 of 40 species

Species

Open these in Browse to filter by habitat, rank, or group.

Species	Common name	Group	Class	Status
Bathyphantes weyeri	Wide Shield Sheetweaver	Araneae	Troglobiont	G4
Brachydesmus superus	Compact Flat-Backed Millipede	Polydesmida	Trogloxene
Caecidotea franzi	Franz's Cave Isopod	Isopoda	Stygobiont	G2G4
Catops gratiosus		Coleoptera	Troglophile	G5
Conasellus kenki		Isopoda	Stygobiont	G3
Conasellus pricei	Price's Cave Isopod	Isopoda	Stygobiont	G5
Conotyla blakei		Chordeumatida	Troglophile	G4
Corynorhinus townsendii	Pacific Townsend's Big-Eared Bat	Chiroptera	Trogloxene	G4
Crangonyx dearolfi	Pennsylvania Cave Amphipod	Amphipoda	Stygobiont	G2
Crangonyx floridanus	Florida Crangonyctid	Amphipoda	Stygophile	G5
Eidmannella pallida	Pallid Cave Spider	Araneae	Troglophile	GNR
Eptesicus fuscus	Big Brown Bat	Chiroptera	Trogloxene	G5
Gammarus cohabitus		Amphipoda	Stygobiont	G1G2
Gammarus minus		Amphipoda	Stygophile	GNR
Myotis leibii	Eastern Small-Footed Bat	Chiroptera	Trogloxene	G4
Myotis lucifugus	Little Brown Bat	Chiroptera	Trogloxene	G3
Myotis septentrionalis	Northern Long-eared Bat	Chiroptera	Trogloxene	G2 ESA
Myotis sodalis	Cluster Bat	Chiroptera	Trogloxene	G2 ESA
Oreonetides beattyi	Beatty's Sheetweb Cave Spider	Araneae	Troglobiont	GNR
Oxidus gracilis	Greenhouse Millipede	Polydesmida	Troglophile	G5
Perimyotis subflavus	Tricolored Bat	Chiroptera	Trogloxene	G3
Phagocata gracilis	Slender Spring Flatworm	Tricladida	Stygophile	G5
Polydesmus inconstans	European Sculpted Millipede	Polydesmida	Accidental	G4
Porrhomma cavernicola	Appalachian Cave Spider	Araneae	Troglobiont	G5
Pseudopolydesmus serratus	Common Pink Flat-Back	Polydesmida	Trogloxene	G5
Scytonotus granulatus	Granulated Millipede	Polydesmida	Trogloxene	G5
Sinella cavernarum		Entomobryomorpha	Troglobiont	G5
Sinella hoffmani	Hoffman's Cave Springtail	Entomobryomorpha	Troglobiont	G5
Spelobia tenebrarum	Cave dung fly	Diptera	Troglobiont	G5
Sphalloplana pricei	Refton Cave Flatworm	Tricladida	Stygobiont	G2G3
Stygobromus allegheniensis		Amphipoda	Stygobiont	G5
Stygobromus biggersi	Biggers' Cave Amphipod	Amphipoda	Stygobiont	G2G4
Stygobromus franzi	Franz's Cave Amphipod	Amphipoda	Stygobiont	G3G4
Stygobromus gracilipes		Amphipoda	Stygobiont	G3G4
Stygobromus hayi	Hay's Spring Amphipod	Amphipoda	Stygobiont	G1G2 ESA
Stygobromus pizzinii	Pizzini's Cave Amphipod	Amphipoda	Stygobiont	G3G4
Stygobromus stellmacki	Stellmack's Cave Amphipod	Amphipoda	Stygobiont	G1G2
Stygobromus tenuis		Amphipoda	Stygobiont	G4
Terrilimosina racovitzai		Diptera	Troglophile	GNR
Typhlogastrura helleri	Heller Cave Springtail	Poduromorpha	Troglobiont	GNR

About this list

Compiled from published records, museum material, and survey reports, and recorded to 28 counties. Coordinates are never published; distribution is published generalized to county and karst region. See the Data Sensitivity Policy.

Absence from this list does not mean absence from the ground. Most caves have never been surveyed, and subterranean fauna is consistently under-recorded. If you know of a species here that is missing, please tell us.