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Subterranean biodiversity of North America and around the globe

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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe

Browse species

2986 species

Species Common name Group Class Status States Photo Spec Seq
Abacion lactarium (Say, 1821) Callipodida Trogloxene G5 GA, TN ○ ○ ○
Abacion magnum (Loomis, 1943) Callipodida Trogloxene G5 AL, GA, VA ○ ○ ○
Acanthocyclops brevispinosus (Herrick, 1884) Cyclopoida Stygoxene IL ○ ○ ○
Acanthocyclops einslei Mirabdullayev & Defaye, 2004 Cyclopoida Stygoxene IL, IN ○ ○ ○
Acanthocyclops exilis (Coker, 1934) Cyclopoida Stygoxene KY, TN ○ ○ ○
Acanthocyclops parasensitivus Reid, 1998 Cyclopoida Stygophile G2 IN, KY, TN ○ ○ ○
Acanthocyclops robustus (Sars G.O., 1863) Robusthops Cyclopoida Stygophile GA, IL, IN, KY, TN ○ ○ ○
Acanthocyclops venustoides (Coker, 1934) Cyclopoida Stygoxene IL, IN ○ ○ ○
Acanthocyclops vernalis (Fischer, 1853) Survannshops Cyclopoida Stygophile GA, IA, IL, IN, KY, NM, TN, TX ○ ○ ○
Achemenides pectinatus (Causey, 1952) Chordeumatida Troglophile G3 IA, IL, MN, WI ○ ○ ○
Aclodes nicuesa Hebard, 1928 Orthoptera Trogloxene ○ ○ ○
Acris blanchardi Harper, 1947 Blanchard's Cricket Frog Anura Accidental G5 LC OK ○ ○ ○
Acris crepitans Baird, 1854 Northern Cricket Frog Anura Accidental G4 LC KY ○ ○ ○
Acris gryllus (LeConte, 1825) Southern Cricket Frog Anura Accidental G5 LC FL ○ ○ ○
Acroneuria abnormis (Newman, 1838) Common Stone Plecoptera Stygoxene G5 ○ ○ ○
Acrotrichis lucidula Rosskothen, 1935 Coleoptera Troglophile G2 TN ○ ○ ○
Actia diffidens Curran, 1933 Diptera Accidental GNR ○ ○ ○
Adelocosa anops Gertsch, 1973 Kauai Cave Wolf Spider Araneae Troglobiont G1G2 EN ESA HI ○ 1 ○
Aedes abserratus (Felt & Young, 1904) Serrated Mosquito Diptera Trogloxene G5 ○ ○ ○
Aedes aurifer (Coquillett, 1903) Golden Bog Mosquito Diptera Trogloxene GNR ○ ○ ○
Aedes canadensis (Theobald, 1901) Woodland Pool Mosquito Diptera Trogloxene G5 ○ ○ ○
Aedes cinereus Wiedemann, 1818 Diptera Trogloxene ○ ○ ○
Aedes communis (De Geer, 1776) Snow Mosquito Diptera Trogloxene G5 ○ ○ ○
Aedes fitchii (Felt & Young, 1904) Diptera Trogloxene G5 ○ ○ ○
Aedes provocans (Walker, 1848) Provoking Mosquito Diptera Trogloxene G5 ○ ○ ○
Aedes punctor (Kirby, 1837) Moustique Pointu Provocateur Canadien Diptera Trogloxene G5 ○ ○ ○
Aedes sticticus (Meigen, 1835) Floodwater Mosquito Diptera Trogloxene G5 ○ ○ ○
Aedes stimulans (Walker, 1848) Woodland Mosquito Diptera Trogloxene G5 ○ ○ ○
Aedes triseriatus (Say, 1823) Eastern Treehole Mosquito Diptera Trogloxene G5 TN ○ ○ ○
Aedes vexans (Meigen, 1830) Vexans Mosquito Diptera Trogloxene G5 ○ ○ ○
Agastoschizomus texanus Monjarez-Ruedas et al., 2016 Schizomida Troglobiont TX ○ 3 ○
Agelenopsis pennsylvanica (C. L. Koch, 1843) Pennsylvania Grass Spider Araneae Trogloxene GNR IL ○ ○ ○
Agkistrodon conanti Gloyd, 1969 Green-Tailed Moccasin Squamata Accidental G5 FL ○ ○ ○
Agkistrodon contortrix (Linnaeus, 1766) Copperhead Squamata Accidental G5 LC AR, KY, TN ○ ○ ○
Agonum extensicolle (Say, 1823) Extensive Harp Ground Beetle Coleoptera Trogloxene G5 AL, MO ○ ○ ○
Agonum punctiforme (Say, 1823) Punctiform Harp Ground Beetle Coleoptera Trogloxene GNR AL, IN ○ ○ ○
Agyneta gagnei (Gertsch, 1973) Araneae Troglobiont G1G2 HI ○ 1 ○
Agyneta hedini Duperre & Paquin, 2009 Araneae Troglobiont CO, NM ○ 3 ○
Agyneta unimaculata (Banks, 1892) Araneae Trogloxene GNR IL ○ ○ ○
Alabamocreagris mortis (Muchmore, 1969) Alabama pseudoscorpion Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Alabamocreagris pecki (Muchmore, 1969) Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Albiorix anophthalmus Muchmore & Pape, 1998 Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Alectorobius concanensis Camicas et al., 1998 Ixodida Trogloxene OK ○ ○ ○
Alectorobius kelleyi Camicas et al., 1998 Ixodida Trogloxene GA ○ ○ ○
Aleochara castaneimarmotae Klimaszewski, Webster & Brunke, 2017 Coleoptera Troglophile GNR AR, IL, KY, MO, TN ○ ○ ○
Aleochara castaneipennis Mannerheim, 1843 Coleoptera Troglophile G5 AL ○ ○ ○
Aleochara fumata Gravenhorst, 1802 Rove Beetle Coleoptera Troglophile GNR AL, TN ○ ○ ○
Aleochara lucifuga (Casey, 1893) Coleoptera Troglophile GNR AL, IA, IL, IN, KY, TN ○ ○ ○
Aleochara ocularis Klimaszewski, 1984 Coleoptera Troglophile GNR IA ○ ○ ○
Alligator mississippiensis (Daudin, 1801) Louisiana Alligator Crocodylia Stygoxene G5 LC FL ○ ○ ○
Allocosa funerea (Hentz, 1844) Araneae Trogloxene GNR TN ○ ○ ○
Allocosa sublata (Montgomery, 1902) Araneae Accidental GNR AL ○ ○ ○
Allocrangonyx hubrichti Holsinger, 1971 Hubricht's Long-tailed Cave Amphipod Amphipoda Stygobiont G3G4 VU AR, MO ○ 18 ○
Allocrangonyx pellucidus (Mackin, 1935) Oklahoma Cave Amphipod Amphipoda Stygobiont G2G3 VU OK, MO ○ 12 ○
Allolobophora chlorotica (Savigny in Cuvier, 1826) Green Worm Crassiclitellata Troglophile VA ○ ○ ○
Allomideopsis wichitaensis (Smith, 1990) Trombidiformes Stygobiont TX ○ ○ ○
Allotexiweckelia hirsuta Holsinger, 1980 Amphipoda Stygobiont G2G3 TX ○ 43 ○
Almuerzothyas comalensis Smith & Cook, 2009 Trombidiformes Stygobiont GNR TX ○ ○ ○
Alocodesmus angustatus Silvestri, 1896 Polydesmida Trogloxene ○ ○ ○
Aloconota diversiseta Klimaszewski & Peck, 1986 Coleoptera Troglophile G1 TN ○ ○ ○
Aloconota insecta (Thomson, C. G., 1856) Staph Beetle Coleoptera Troglophile GNR AL, IL, TN ○ ○ ○
Aloconota neospelea Klimaszewski & Peck, 1986 Coleoptera Troglophile GNR AL ○ ○ ○
Aloconota sulcifrons (Stephens, 1832) Sulcate-Faced Minute Rove Beetle Coleoptera Troglophile GNR TN ○ ○ ○
Alucita huebneri Wallengren, 1859 Lepidoptera Accidental ○ ○ ○
Amara musculis (Say, 1823) Coleoptera Accidental G5 AL, KY ○ ○ ○
Ambloplites rupestris (Rafinesque, 1817) Rock Bass Centrarchiformes Stygoxene G5 LC KY ○ ○ ○
Amblyopsis hoosieri Niemiller, Prejean & Chakrabarty, 2014 Hoosier Cavefish Percopsiformes Stygobiont G2 IN ○ 63 1
Amblyopsis spelaea DeKay, 1842 Northern Cavefish Percopsiformes Stygobiont G2G3 NT KY ○ 93 384
Ambystoma maculatum (Shaw, 1802) Spotted Salamander Caudata Accidental G5 LC AL, AR, OH ○ ○ ○
Ambystoma opacum (Gravenhorst, 1807) Marbled Salamander Caudata Accidental G4 LC FL, OH ○ ○ ○
Ambystoma tigrinum (Green, 1825) Eastern Tiger Salamander Caudata Accidental G5 LC FL, GA, IL ○ ○ ○
Ameiurus catus (Linnaeus, 1758) White Catfish Siluriformes Stygoxene G5 LC FL ○ ○ ○
Ameiurus melas (Rafinesque, 1820) Black Bullhead Siluriformes Stygoxene G5 LC AL, IL ○ ○ ○
Ameiurus natalis (Lesueur, 1819) Yellow Bullhead Siluriformes Stygoxene G5 LC AL, FL, KY ○ ○ ○
Ameiurus nebulosus (Lesueur, 1819) Brown Bullhead Siluriformes Stygoxene G5 LC FL, GA ○ ○ ○
Ameractis satis Causey, 1959 Julida Troglobiont G2G4 AL, GA, TN, VA ○ 1 ○
Americina adusta (Loew, 1863) Diptera Trogloxene GNR ○ ○ ○
Americocreagris columbiana (Chamberlin, 1962) Pseudoscorpiones Troglobiont G2G3 OR, WA ○ ○ ○
Amerigoniscus centralis Vandel, 1978 Isopoda Troglobiont G1G2 OK ○ ○ ○
Amerigoniscus curvatus Vandel, 1978 Isopoda Troglobiont G1 GA ○ ○ ○
Amerigoniscus georgiensis Vandel, 1978 Isopoda Troglobiont G1 GA ○ ○ ○
Amerigoniscus gipsocolus (Vandel, 1965) Isopoda Troglobiont G1G2 TX, NM ○ 1 ○
Amerigoniscus henroti Vandel, 1950 Powell Valley Terrestrial Cave Isopod Isopoda Troglobiont G3 VA ○ 10 ○
Amerigoniscus malheurensis Schultz, 1982 Isopoda Troglobiont G1 OR ○ 2 ○
Amerigoniscus nicholasi (Vandel, 1965) Isopoda Troglobiont G1G2 GA, TN ○ 4 ○
Amerigoniscus proximus Vandel, 1978 Isopoda Troglobiont G1G2 GA ○ ○ ○
Amerizus wingatei (Bland, 1864) Coleoptera Trogloxene G5 IA, IL, KY, WI ○ ○ ○
Ameroduvalius jeanneli Valentine, 1952 Coleoptera Troglobiont G3 KY ○ 8 6
Amnestus subferrugineus (Westwood, 1837) Hemiptera Troglophile ○ ○ ○
Amnicola cora Hubricht, 1979 Foushee Cavesnail Littorinimorpha Stygobiont G1 VU AR, AZ ○ 9 ○
Amnicola stygius Hubricht, 1971 Stygian cavesnail Littorinimorpha Stygobiont G1 VU MO ○ 6 ○
Amoeba guttula Duj. Euamoebida Stygoxene AL, KY ○ ○ ○
Amoeba vespertilio Pen. Euamoebida Stygoxene AL ○ ○ ○
Amoebaleria defessa (Osten Sacken, 1877) Diptera Troglophile GNR AL, AR, GA, IL, IN, KY, MO, TN, WI ○ ○ ○
Amphasia interstitialis (Say, 1823) Red-Headed Ground Beetle Coleoptera Accidental G5 IL ○ ○ ○
Amphibiocapillaria texensis Moravec & Huffman, 2000 Trichocephalida Stygobiont TX ○ ○ ○
Amplaria flucticulus Shear, 2021 Chordeumatida Troglobiont CA ○ ○ ○
Amplaria shastae (Causey, 1958) Chordeumatida Troglobiont G1G2 CA ○ 1 ○
Amynthas diffringens (Baird, 1869) Japanese Jumper Crassiclitellata Troglophile GNR AL ○ ○ ○
Amynthas minimus (Horst, 1893) Crassiclitellata Troglophile GA ○ ○ ○
Anachemmis jungi Platnick & Ubick, 2005 Araneae Troglobiont CA ○ ○ ○
Anahita punctulata (Hentz, 1844) Southeastern Wandering Spider Araneae Trogloxene G4 AL, GA, TN ○ ○ ○
Anargyrtes annulata (Bilimek, 1867) Grillo De Cueva Orthoptera Trogloxene ○ ○ ○
Anasaitis canosus (Walckenaer, 1837) Twin-Flagged Jumping Spider Araneae Accidental AL ○ ○ ○
Anaxyrus americanus (Holbrook, 1836) American Toad Anura Accidental G5 LC AL, IL, KY, TN ○ ○ ○
Anaxyrus fowleri (Hinckley, 1882) Fowler's Toad Anura Accidental G5 LC GA, KY ○ ○ ○
Anaxyrus terrestris (Bonnaterre, 1789) Southern Toad Anura Accidental G5 LC FL, GA ○ ○ ○
Andrognathus corticarius Cope, 1869 Cope's Noodle Millipede Platydesmida Trogloxene G5 GA ○ ○ ○
Aneides aeneus (Cope & Packard, 1881) Green Salamander Caudata Trogloxene G3 NT AL, GA, TN ○ ○ ○
Anevrina olympiae (Aldrich, 1904) Diptera Trogloxene GNR IA ○ ○ ○
Anguilla rostrata (Lesueur, 1817) American Eel Anguilliformes Stygoxene G4 EN FL, GA ○ ○ ○
Anguispira alternata (Say, 1817) Stylommatophora Trogloxene AL, FL, TN ○ ○ ○
Anguispira mordax (Shuttleworth, 1852) Appalachian Tigersnail Stylommatophora Trogloxene G4 KY, TN ○ ○ ○
Anillaspis explanata (Horn, 1888) Coleoptera Troglobiont CA ○ 1 ○
Anillinus actuipennis Sokolov & Reddell, 2014 Coleoptera Troglobiont TX ○ ○ ○
Anillinus bexarensis Sokolov, 2022 Coleoptera Troglobiont TX ○ ○ ○
Anillinus cavicola Sokolov, 2012 Coleoptera Troglobiont AL ○ ○ ○
Anillinus clinei Sokolov, 2020 Coleoptera Troglobiont AL ○ ○ ○
Anillinus folkertsioides Sokolov, 2020 Coleoptera Troglophile AL ○ ○ ○
Anillinus forthoodensis Sokolov & Reddell, 2014 Coleoptera Troglobiont TX ○ 4 ○
Anillinus hildebrandti Sokolov, 2020 Coleoptera Troglobiont AL ○ ○ ○
Anillinus longiceps Jeannel, 1963 Coleoptera Troglobiont TN ○ 2 ○
Anillinus reddelli Sokolov, 2022 Coleoptera Troglobiont TX ○ ○ ○
Anillinus smokiensis Sokolov, 2011 Coleoptera Troglobiont TN ○ ○ 13
Anillinus tombarri Sokolov, 2012 Coleoptera Troglobiont AL ○ ○ ○
Anillinus valentinei (Jeannel, 1963) Coleoptera Troglobiont G3G4 AL ○ 2 ○
Anillinus wisemanensis Sokolov & Kavanaugh, 2014 Coleoptera Troglobiont TX ○ 4 ○
Anisodactylus opaculus (LeConte, 1863) Coleoptera Accidental MO ○ ○ ○
Anisolabis howarthi Brindle, 1980 Dermaptera Troglobiont G1G2 HI ○ 1 ○
Aniulus nigrans (Chamberlin, 1918) Julida Trogloxene TN ○ ○ ○
Ankyrodrilus legaeus Holt, 1965 Branchiobdellida Stygophile TN ○ ○ ○
Anolis carolinensis Voigt, 1832 Carolina Anole Squamata Accidental G5 LC FL, TN ○ ○ ○
Anopheles earlei Vargas, 1943 Diptera Trogloxene G5 ○ ○ ○
Anopheles punctipennis (Say, 1823) Woodland Malaria Mosquito Diptera Trogloxene G5 GA, IL, IN, KY, TN ○ ○ ○
Anopheles quadrimaculatus Say, 1824 Common Malaria Mosquito Diptera Trogloxene G5 IL, IN ○ ○ ○
Anotylus exiguus (Erichson, 1840) Rove Beetle Coleoptera Trogloxene AL, IA ○ ○ ○
Anotylus tetracarinatus (Block, 1799) Rove Beetle Coleoptera Trogloxene GNR AL ○ ○ ○
Anthicus cervinus LaFerté-Senéctère, 1848 Cloudy Flower Beetle Coleoptera Accidental G5 WI ○ ○ ○
Anthrobia coylei Miller, 2005 Araneae Troglobiont G2 VA, WV, KY, TN ○ ○ ○
Anthrobia monmouthia Tellkampf, 1844 Mammoth Cave Spider Araneae Troglobiont G5 AL, KY, TN, VA ○ 3 ○
Anticarsia gemmatalis Hübner, 1818 Velvetbean Caterpillar Lepidoptera Trogloxene G5 AL ○ ○ ○
Antrobia culveri Hubricht, 1971 Tumbling Creek cavesnail Littorinimorpha Stygobiont G1 EN ESA MO ○ 7 ○
Antrodiaetus unicolor (Hentz, 1842) Araneae Trogloxene GNR AL, GA, TN ○ ○ ○
Antrolana lira Bowman 1964 Isopoda Stygobiont G2G4 VU ESA VA, WV ○ 37 17
Antrorbis breweri Hershler & Thompson, 1990 Manitou cavesnail Littorinimorpha Stygobiont G1 VU AL ○ 7 1
Antrorbis tennesseensis Perez, Shoobs, Gladstone, & Niemiller, 2019 Tennessee Cavesnail Littorinimorpha Stygobiont GNR TN ○ 3 6
Antroselates spiralis Hubricht, 1963 Shaggy cavesnail Littorinimorpha Stygobiont G3 IN, KY ○ 34 3
Antroxestoblatta immaculata (Hebard, 1920) Blattodea Trogloxene ○ ○ ○
Anurida tullbergi Schött, H, 1891 Poduromorpha Accidental IA ○ ○ ○
Anyphaena celer (Hentz, 1847) Araneae Accidental GNR AL ○ ○ ○
Aphaenogaster tennesseensis (Mayr, 1862) Tennessee Collared Ant Hymenoptera Accidental G5 KY ○ ○ ○
Apheloria corrugata (Wood, 1864) Polydesmida Accidental KY ○ ○ ○
Apheloria montana (Bollman, 1887) Polydesmida Accidental G5 TN ○ ○ ○
Apheloria virginiensis Drury, 1770 Black-And-Gold Flat Millipede Polydesmida Accidental G5 KY ○ ○ ○
Aphrastochthonius grubbsi Muchmore, 1984 Grubb's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Aphrastochthonius pachysetus Muchmore, 1976 Pseudoscorpiones Troglobiont G1G2 NM ○ ○ ○
Aphrastochthonius pecki Muchmore, 1968 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Aphrastochthonius similis Muchmore, 1984 Carlow's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Aphrastochthonius tenax Chamberlin, 1962 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Aphredoderus gibbosus Lesueur, 1833 Percopsiformes Stygoxene FL ○ ○ ○
Aplexa hypnorum (Linnaeus, 1758) Moss Bladder Snail Basommatophora Stygoxene ○ ○ ○
Apocheiridium reddelli Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ 3
Apochthonius coecus (Packard, 1884) Pseudoscorpiones Troglobiont G1G2 VA ○ 2 ○
Apochthonius colecampi Muchmore, 1967 Pseudoscorpiones Troglobiont G1G2 MO ○ ○ ○
Apochthonius diabolus Muchmore, 1967 Pseudoscorpiones Troglobiont G1 AR ○ ○ ○
Apochthonius forbesi Benedict, 1979 Pseudoscorpiones Troglobiont G1G2 OR ○ ○ ○
Apochthonius grubbsi Muchmore, 1980 Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Apochthonius hobbsi Muchmore, 1994 Buckskin Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 OH ○ ○ ○
Apochthonius holsingeri Muchmore, 1967 Pseudoscorpiones Troglobiont G1G2 VA ○ ○ ○
Apochthonius indianensis Muchmore, 1967 Indiana Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 IN ○ ○ ○
Apochthonius malheuri Benedict & Malcolm, 1973 Malheur Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 OR ○ ○ ○
Apochthonius minor Muchmore, 1976 Pseudoscorpiones Troglobiont G1 GA, TN ○ ○ ○
Apochthonius moestus (Banks, 1891) Pseudoscorpiones Troglophile GNR KY ○ ○ ○
Apochthonius mysterius Muchmore, 1976 Mystery Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 MO ○ ○ ○
Apochthonius paucispinosus Muchmore, 1967 Dry Fork Valley Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Apochthonius russelli Muchmore, 1976 Russell Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Apochthonius titanicus Muchmore, 1976 Pseudoscorpiones Troglobiont G2 AR ○ ○ ○
Apochthonius typhlus Muchmore, 1967 Stone County Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 MO ○ ○ ○
Aporrectodea longa (Ude, 1885) Deep-Burrowing Longworm Crassiclitellata Troglophile GNR AL ○ ○ ○
Aporrectodea rosea (Savigny in Cuvier, 1826) Pink Soilworm Crassiclitellata Troglophile GNR KY, VA ○ ○ ○
Aporrectodea trapezoides (Duges, 1828) Crassiclitellata Troglophile AL, GA, IL, KY, TN, WV ○ ○ ○
Aporrectodea tuberculata (Eisen, 1874) Crassiclitellata Troglophile IA, MN ○ ○ ○
Aporrectodea turgida (Eisen, 1874) Crassiclitellata Troglophile VA ○ ○ ○
Appaleptoneta barrowsi (Gertsch, 1974) Araneae Troglobiont G1 AL ○ 1 2
Appaleptoneta credula (Gertsch, 1974) Araneae Troglobiont G1 AL ○ 5 2
Appaleptoneta fiskei (Gertsch, 1974) Araneae Troglobiont GNR GA ○ 13 3
Appaleptoneta jonesi (Gertsch, 1974) Araneae Troglobiont G1 AL ○ ○ ○
Aquarius conformis (Uhler, 1878) Hemiptera Stygoxene GNR OK ○ ○ ○
Aquarius remigis (Say, 1832) North American Common Water Strider Hemiptera Stygoxene GNR AR, IL, MO, OK ○ ○ ○
Araneus cavaticus (Keyserling, 1881) Barn Orbweaver Araneae Trogloxene GNR AL, KY ○ ○ ○
Araneus diadematus Clerck, 1757 Cross Orbweaver Araneae Trogloxene GNR ○ ○ ○
Araneus marmoreus Clerck, 1757 Marbled Orbweaver Araneae Accidental G5 TN ○ ○ ○
Arcteonais lomondi (Martin, 1907) Aquatic Oligochaete Worm Tubificida Stygophile GNR GA ○ ○ ○
Arenophilus bipuncticeps Wood (1862) Northern Short-Clawed Centipede Geophilomorpha Trogloxene GNR GA ○ ○ ○
Argyrodes nephilae Taczanowski, 1873 Araneae Accidental AL ○ ○ ○
Arianops cavernensis Park, 1951 Coleoptera Troglobiont GNR AL ○ 7 ○
Arianops gorei Carlton, 2008 Coleoptera Troglobiont TN ○ 1 ○
Arianops jeanneli Park, 1956 Coleoptera Troglobiont G1 VA ○ ○ ○
Arianops pecki Barr, 1974 Coleoptera Troglobiont TN ○ 1 ○
Arianops steevesi Barr, 1974 Coleoptera Troglobiont GNR AL ○ 1 ○
Arianops stygica Park, 1960 Coleoptera Troglobiont G1 TN ○ ○ ○
Arion fasciatus (Nilsson, 1823) Banded Slug Stylommatophora Trogloxene G5 LC ○ ○ ○
Armadillidium nasatum Budde-Lund, 1885 Nosy Pill Woodlouse Isopoda Troglophile GNR AR, IL, KY, OK, TN ○ ○ ○
Armadillidium vulgare (Latreille, 1804) Common Pill Woodlouse Isopoda Troglophile GNR AL, AR, GA, IL, MO, TN ○ ○ ○
Arrhopalites benitus (Folsom, 1896) Symphypleona Trogloxene IA ○ ○ ○
Arrhopalites caecus Barjadze & Djanashvili, 2008 Symphypleona Troglophile IA, TN ○ ○ ○
Artesia subterranea Holsinger, 1980 Amphipoda Stygobiont G1G2 TX ○ 94 ○
Artesia welbourni Holsinger, 1992 Amphipoda Stygobiont G1G2 TX ○ 2 ○
Atheta alabama Klimaszewski & Peck, 1986 Coleoptera Troglophile GNR AL ○ ○ ○
Atheta annexa Casey, 1910 Coleoptera Troglophile G5 AL, FL, GA, IA, TN ○ ○ ○
Atheta klagesi Bernhauer, 1909 Coleoptera Troglophile G5 GA ○ ○ ○
Atheta lucifuga Klimaszewski & Peck, 1986 Coleoptera Troglophile G4 AL, FL, GA, TN ○ ○ ○
Atheta troglophila Klimaszewski & Peck, 1986 Coleoptera Troglophile G4 AL, FL, GA, IA, OK, TN ○ ○ ○
Atranus pubescens (Dejean, 1828) Coleoptera Troglophile GNR AL, AR, GA, IL, IN, KY, MO, TN ○ ○ ○
Attagenus brunneus Faldermann, 1835 Coleoptera Trogloxene GNR IA ○ ○ ○
Attheyella illinoisensis (Forbes, 1876) Cyclopoida Stygoxene GA, IN, TN ○ ○ ○
Attheyella nordenskioldii (Borutzky, 1952) Cyclopoida Stygoxene IL, IN ○ ○ ○
Attheyella pilosa (Chappuis, 1929) Cyclopoida Stygoxene GA ○ ○ ○
Australinocreagris grahami (Muchmore, 1969) Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Austrotyla awishoshola Wynne & Shear, 2016 Chordeumatida Troglobiont NM ○ ○ ○
Austrotyla coloradensis (Chamberlin, 1910) Chordeumatida Troglobiont CO ○ 47 ○
Austrotyla montivaga (Loomis, 1943) Chordeumatida Accidental NM ○ ○ ○
Austrotyla specus (Loomis, 1939) Chordeumatida Troglophile IL, MO ○ ○ ○
Austrotyla stephensoni (Chamberlin, 1910) Chordeumatida Troglobiont CO ○ 1 ○
Axonopsella bakeri Smith & Cook, 1998 Trombidiformes Stygobiont FL, TX ○ ○ ○
Azilia affinis O. Pickard-Cambridge, 1893 Longjawed Orb Weavers Araneae Trogloxene AL, FL ○ ○ ○
Bactrurus angulus Koenemann & Holsinger, 2001 Cumberland Gap Cave Amphipod Amphipoda Stygobiont G1 TN, VA ○ 4 ○
Bactrurus brachycaudus Hubricht & Mackin, 1940 Amphipoda Stygobiont G4 IL, MO ○ 98 8
Bactrurus cellulanus Koenemann & Holsinger, 2001 Amphipoda Stygobiont GX IN ○ 4 ○
Bactrurus hubrichti Shoemaker, 1945 Kansas Well Amphipod Amphipoda Stygobiont G4 KS, MO, OK, VA ○ 26 ○
Bactrurus mucronatus (Forbes, 1876) Amphipoda Stygobiont G5 IA, IL, IN, MI, OH, AR, MO ○ 126 6
Bactrurus pseudomucronatus Koenemann & Holsinger, 2001 False Sword-tailed Cave Amphipod Amphipoda Stygobiont G2G3 AR, MO ○ 25 5
Bactrurus speleopolis Holsinger, Sawicki, & Graening, 2006 Cave City Cave Amphipod Amphipoda Stygobiont G1 AR ○ 4 ○
Bactrurus wilsoni Koenemann & Holsinger, 2001 Amphipoda Stygobiont G1G2 AL ○ 4 ○
Balantiopteryx plicata Peters, 1867 Gray Sac-Winged Bat Chiroptera Trogloxene LC ○ ○ ○
Balconorbis coronae (Hershler & Longley, 1987) Littorinimorpha Stygobiont TX ○ ○ ○
Balconorbis uvaldensis Hershler & Longley, 1986 Balcones ghostsnail Littorinimorpha Stygobiont G1G2 TX ○ 47 9
Banksula californica Briggs, 1974 Alabaster Cave Harvestman Opiliones Troglobiont GH CA ○ 3 ○
Banksula galilei Briggs, 1974 Galile's Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Banksula grahami Briggs, 1974 Opiliones Troglobiont G2G3 CA ○ 1 ○
Banksula grubbsi Briggs & Ubick, 1981 Grubbs' Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Banksula martinorum Briggs & Ubick, 1981 Martins' Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Banksula melones Briggs, 1974 Melones Cave Harvestman Opiliones Troglobiont G1 VU CA ○ 1 ○
Banksula rudolphi Briggs & Ubick, 1981 Rudolph's Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Banksula tuolumne Briggs, 1974 Tuolumne Cave Harvestman Opiliones Troglobiont G2 CA ○ 1 ○
Banksula tutankhamen Ubick & Briggs, 2002 King Tut Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Basacallis tarachodes Dyar, 1914 Lepidoptera Accidental GNR FL ○ ○ ○
Basilia boardmani Rozeboom, 1934 Diptera Trogloxene G3 FL ○ ○ ○
Bathynella fraterna Cho & Kim, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Bathynella germanitas Cho & Kim, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Bathynella riparia Pennak & Ward, 1985 Bathynellacea Stygobiont CO ○ 1 ○
Bathyphantes alboventris (Banks, 1892) Notch-Tipped Shield Sheetweaver Araneae Troglophile GNR IL, TN ○ ○ ○
Bathyphantes nigrinus (Westring, 1851) Donker Wevertje Araneae Trogloxene GNR IN ○ ○ ○
Bathyphantes pallidus (Banks, 1892) Pale Sheetweb Weaver Araneae Troglophile G5 GA, IL, MO, TN ○ ○ ○
Bathyphantes reprobus (Kulczyński, 1916) Reprobus Sheetweb Weaver Araneae Troglophile GNR ○ ○ ○
Bathyphantes weyeri (Emerton, 1875) Wide Shield Sheetweaver Araneae Troglobiont G4 AR, IN, KY, ON, PA, QC, VA, WI, WV, CO, ME, OH ○ 9 ○
Batriasymmodes greeveri Park, 1965 Coleoptera Troglobiont G1G2 TN ○ 2 ○
Batriasymmodes monstrosus (LeConte, J. L., 1849) Coleoptera Troglophile VA ○ ○ ○
Batriasymmodes parki Barr, 1987 Coleoptera Troglobiont GH WV ○ 1 ○
Batriasymmodes quisnamus (Park, O., 1951) Coleoptera Troglobiont G3 KY, TN ○ 3 ○
Batriasymmodes spelaeus (Park, 1951) Coleoptera Troglobiont G3G4 AL, GA, TN ○ ○ 2
Batriasymmodes troglodytes Park, 1951 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Batrisodes barri Park, 1958 Coleoptera Troglobiont G1G2 TN ○ 2 ○
Batrisodes cavernosus Park, 1951 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes clypeospecus Park, 1960 Coleoptera Troglobiont G1G2 TN ○ ○ ○
Batrisodes cryptotexanus Chandler & Reddell, 2001 Coffin Cave Mold Beetle Coleoptera Troglobiont G2 TX ○ 28 ○
Batrisodes denticollis (Casey, 1884) Coleoptera Trogloxene VA ○ ○ ○
Batrisodes dentifrons Chandler & Reddell, 2009 Coleoptera Troglobiont G1G2 TX ○ 1 ○
Batrisodes fanti Chandler & Reddell, 2009 Coleoptera Troglobiont G1G2 TX ○ 1 ○
Batrisodes feminiclypeus Chandler & Reddell, 2001 Coleoptera Troglobiont G1G2 TX ○ 3 ○
Batrisodes ferulifer Park, 1960 Coleoptera Troglobiont G1 TN ○ ○ ○
Batrisodes gemmoides Park, 1960 Coleoptera Troglobiont G1 TN ○ ○ ○
Batrisodes gemmus Park, 1956 Coleoptera Troglobiont G1 TN ○ ○ ○
Batrisodes gravesi Chandler & Reddell, 2001 Coleoptera Troglobiont G3 TX ○ 14 ○
Batrisodes grubbsi Chandler, 1992 Coleoptera Troglobiont G1G2 TX ○ 3 ○
Batrisodes henroti Park, 1956 Coleoptera Troglobiont G2G3 KY ○ 2 ○
Batrisodes hubrichti Park, 1958 Coleoptera Troglobiont G1 KY ○ 2 ○
Batrisodes incispes Chandler & Reddell, 2009 Coleoptera Troglobiont G1 TX ○ ○ ○
Batrisodes jocuvestus Park, 1960 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes jonesi Park, O., 1951 Coleoptera Troglobiont G2G3 AL ○ ○ ○
Batrisodes krekeleri Park, 1960 Coleoptera Troglobiont IN ○ ○ ○
Batrisodes lineaticollis (Aubé, 1833) Coleoptera Troglophile G5 AL, GA, TN ○ ○ ○
Batrisodes pannosus Park, 1960 Coleoptera Troglobiont G1 TN ○ 2 ○
Batrisodes pekinsi Chandler & Reddell, 2009 Coleoptera Troglobiont G1 TX ○ 2 ○
Batrisodes profundus Park, 1956 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Batrisodes reyesi Chandler, 1992 Coleoptera Troglobiont G2G3 TX ○ 25 ○
Batrisodes rossi Park, O., 1947 Coleoptera Troglophile GNR IL ○ ○ ○
Batrisodes schmitti (Casey, 1897) Coleoptera Troglophile TN ○ ○ ○
Batrisodes shadeae Chandler & Reddell, 2009 Coleoptera Troglobiont G1 TX ○ 1 ○
Batrisodes specus Park, 1951 Coleoptera Troglobiont G3G4 AL, TN, TX ○ 4 ○
Batrisodes subterraneus Park, 1951 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes texanus Chandler, 1992 Inner Space Cavern Mold Beetle Coleoptera Troglobiont G1G2 ESA TX ○ 15 ○
Batrisodes tumoris Park, 1960 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes valentinei Park, 1951 Coleoptera Troglobiont G2G4 AL, TN ○ ○ ○
Batrisodes venyivi Chandler, 1992 Helotes Cave Mold Beetle Coleoptera Troglobiont G1 ESA TX ○ 4 ○
Batrisodes wartoni Chandler & Reddell, 2001 Coleoptera Troglobiont G1G2 TX ○ 1 ○
Bdellodrilus illuminatus (Moore, 1894) Branchiobdellida Stygophile TN, WV ○ ○ ○
Belba bulbipedata (Packard, 1888) Oribatida Troglobiont G1 KY ○ ○ ○
Bembidion affine Say, 1823 Coleoptera Trogloxene GNR AL ○ ○ ○
Bembidion lacunarium (Zimmermann in LeConte, 1869) Greenbrier Riverbank Ground Beetle Coleoptera Troglophile GNR AL, GA, IA, IL, KY, TN ○ ○ ○
Bembidion nigripes (Kirby, 1837) Coleoptera Trogloxene G5 AL ○ ○ ○
Bembidion texanum Chaudoir, 1868 Simple Riverbank Ground Beetle Coleoptera Troglophile GNR IL, MO, OK ○ ○ ○
Bertkauia crosbyana Chapman, 1930 Psocodea Trogloxene AL ○ ○ ○
Bibio xanthopus Wiedemann, 1828 Yellow-Footed March Fly Diptera Accidental GNR ○ ○ ○
Bicornucandona fineganensis Kulkoyluoglu et al., 2011 Podocopida Stygobiont GNR TX ○ ○ ○
Bimastos heimburgeri (Smith, 1928) Crassiclitellata Troglophile GNR IL ○ ○ ○
Bimastos longicinctus (Smith & Gittins, 1915) Crassiclitellata Troglophile IL ○ ○ ○
Bimastos rubidus (Savigny in Cuvier, 1826) Gilt-Tail Crassiclitellata Troglophile GNR AL, GA, IA, IL, KY, MO, NC, TN, VA ○ ○ ○
Bimastos tumidus (Eisen, 1874) Crassiclitellata Troglophile AL, GA, MO, TN, VA ○ ○ ○
Bimastos zeteki (Smith & Gittins, 1915) Crassiclitellata Troglophile GA, TN ○ ○ ○
Bishopella jonesi Goodnight & Goodnight, 1942 Opiliones Troglobiont G1 AL ○ ○ ○
Bishopella laciniosa (Crosby & Bishop, 1924) Bishop's Harvestman Opiliones Troglophile AL, FL, GA, TN ○ ○ ○
Bishopella marianna Goodnight & Goodnight, 1942 Opiliones Troglophile FL ○ ○ ○
Bittacomorpha clavipes (Fabricius, 1781) Phantom Crane Fly Diptera Trogloxene GNR ○ ○ ○
Blaberus giganteus (Linnaeus, 1758) Caribbean Giant Cockroach Blattodea Trogloxene ○ ○ ○
Blackburnia aaae (Samuelson & Liebherr, 1992) Coleoptera Troglobiont G1G2 HI ○ ○ ○
Blackburnia howarthi (Samuelson & Liebherr, 1992) Coleoptera Troglobiont G1G2 HI ○ ○ ○
Blancosoma scaturgo Shear & Hubbard, 1998 Chordeumatida Troglobiont G1G2 CO ○ ○ ○
Blaniulus guttulatus (Fabricius, 1798) Snake Millepede Julida Trogloxene G5 GA ○ ○ ○
Blarina brevicauda (Say, 1823) Eulipotyphla Accidental AL, KY ○ ○ ○
Blarina carolinensis (Bachman, 1837) Southern Short-Tailed Shrew Eulipotyphla Accidental G5 LC AR, FL ○ ○ ○
Bleptina caradrinalis Guenée, 1854 Bent-Winged Owlet Lepidoptera Trogloxene G5 AL ○ ○ ○
Bodo minimus Klebs Eubodonida Stygoxene AL, KY ○ ○ ○
Boisea trivittata (Say, 1825) Boxelder Bug Hemiptera Accidental GNR IL ○ ○ ○
Bolitophila cinerea Meigen & Wiedemann, 1818 Diptera Trogloxene GNR ○ ○ ○
Bolitophila hybrida (Meigen, 1804) Diptera Trogloxene GNR IN ○ ○ ○
Bolitophila perlata Garrett, 1925 Diptera Trogloxene GNR ○ ○ ○
Bos taurus Linnaeus, 1758 Taurine Cattle Artiodactyla Accidental G5 KY ○ ○ ○
Bosmina longirostris (O. F. Müller, 1776) Common Long-Nosed Waterflea Anomopoda Stygoxene GNR KY, TN ○ ○ ○
Bowmanasellus sequoiae (Bowman, 1975) Sequoia Cave Isopod Isopoda Stygobiont G2 CA ○ ○ ○
Brachinus fumans (Fabricius, 1781) American Bombardier Beetle Coleoptera Accidental GNR MO ○ ○ ○
Brachycybe lecontii Wood, 1864 Platydesmida Trogloxene G5 AR ○ ○ ○
Brachycybe petasata Loomis, 1936 Platydesmida Trogloxene TN ○ ○ ○
Brachycybe producta Loomis, 1936 Platydesmida Trogloxene CA ○ ○ ○
Brachydesmus superus Latzel, 1884 Compact Flat-Backed Millipede Polydesmida Trogloxene PA, VA ○ ○ ○
Brackenridgia ashleyi Lewis, 2004 Ashley's Terrestrial Cave Isopod Isopoda Troglobiont G1G3 AR, MO ○ 3 ○
Brackenridgia cavernarum Ulrich, 1902 Isopoda Troglobiont G3G4 TX, NM ○ 8 ○
Brackenridgia reddelli (Vandel, 1965) Isopoda Troglobiont G2G3 TX ○ 13 ○
Bradycellus rupestris (Say, 1823) Rock-Dwelling Harp Ground Beetle Coleoptera Trogloxene G5 AL ○ ○ ○
Bradysia forficulata (Bezzi, 1914) Diptera Troglophile GNR GA ○ ○ ○
Brathinus nitidus LeConte, J. L., 1852 Coleoptera Troglophile G4 AL, KY, TN ○ ○ ○
Bremia caricis (Felt, 1907) Diptera Trogloxene ○ ○ ○
Bryocamptus morrisoni (Chappuis, 1929) Morrison's Cave Copepod Harpacticoida Stygobiont G3G4 IN, KY, MD ○ 2 ○
Bryocamptus newyorkensis (Chappuis, 1927) Harpacticoida Stygoxene TN ○ ○ ○
Bryocamptus nivalis (Willey, 1925) Harpacticoida Stygoxene IN ○ ○ ○
Caconemobius albus Otte, 1994 Orthoptera Troglobiont HI ○ ○ ○
Caconemobius howarthi Gurney & Rentz, 1978 Howarth's Cave Cricket Orthoptera Troglobiont G1G2 VU HI ○ ○ ○
Caconemobius paralbus Otte, 1994 Orthoptera Troglobiont HI ○ ○ ○
Caconemobius uuku Otte, 1994 Orthoptera Troglobiont HI ○ 1 ○
Caconemobius varius Gurney & Rentz, 1978 Kuamana Cave Cricket Orthoptera Troglobiont G1G2 VU HI ○ 1 ○
Caecidotea acuticarpa Mackin & Hubricht, 1940 Isopoda Stygobiont G2G3 OK, KS ○ 26 ○
Caecidotea adenta Mackin & Hubricht, 1940 Isopoda Stygobiont G1G2 OK ○ 3 ○
Caecidotea alabamensis Stafford, 1911 Isopoda Stygobiont G3G4 AL, IL, IN, KY, TN ○ 128 ○
Caecidotea alleghenyensis Lewis, Bowman, & Feller, 2011 Allegheny Spring Isopod Isopoda Stygobiont G1G2 MD ○ 18 ○
Caecidotea ancyla (Fleming, 1972) Isopoda Stygobiont G3G4 AR, MO, OK ○ 34 ○
Caecidotea antricola Creaser, 1931 Isopoda Stygobiont G5 AR, MO, OK, IL, KY, OH ○ 191 10
Caecidotea barri (Steeves, 1965) Clifton Cave Isopod Isopoda Stygobiont G1 EN KY ○ 3 ○
Caecidotea beattyi Lewis & Bowman, 1981 Beatty's Groundwater Isopod Isopoda Stygobiont G3G4 IL, IN, MO ○ 16 13
Caecidotea bicrenata (Steeves, 1963) Two-toothed Cave Isopod Isopoda Stygobiont G5 AL, IL, KY, TN ○ 66 28
Caecidotea bilineata Lewis & Bowman, 1996 Isopoda Stygobiont G2G3 TX ○ 4 ○
Caecidotea cannula (Steeves, 1963) Cannulate Cave Isopod Isopoda Stygobiont G1 WV ○ 12 ○
Caecidotea carolinensis (Lewis & Bowman, 1977 Bennett's Mill Cave Isopod Isopoda Stygobiont G2G3 NC, SC ○ 4 ○
Caecidotea catachaetus (Fleming & Steeves, 1972) Isopoda Stygobiont GA, TN, AL ○ 10 ○
Caecidotea circulus (Steeves & Holsinger, 1968) Isopoda Stygobiont G1G2 TN ○ 3 ○
Caecidotea cumberlandensis Lewis, 2000 Cumberland Gap Cave Isopod Isopoda Stygobiont G1G2 VA ○ 2 3
Caecidotea cyrtorhynchus (Fleming & Steeves, 1972) Isopoda Stygobiont G1 GA ○ 4 ○
Caecidotea dauphina Modlin, 1986 Isopoda Stygobiont AL ○ 2 ○
Caecidotea dimorpha (Mackin & Hubricht, 1940) Isopoda Stygobiont G2G3 AR, MO, IN ○ 17 ○
Caecidotea dunlaporum Lewis, 2015 Dunlap's Groundwater Isopod Isopoda Stygobiont GNR IN ○ ○ 4
Caecidotea extensolinguala (Fleming, 1972) Isopoda Stygobiont MO ○ 3 ○
Caecidotea filicispeluncae Bowman & Hobbs, 1983 Fern Cave Isopod Isopoda Stygobiont G1 OH ○ 3 ○
Caecidotea fisherorum Lewis & Lewis, 2023 Isopoda Stygobiont GNR VA ○ ○ 15
Caecidotea fonticulus Lewis, 1983 Isopoda Stygobiont GNR AR ○ 5 ○
Caecidotea franzi (Holsinger & Steeves, 1971) Franz's Cave Isopod Isopoda Stygobiont G2G4 KY, MD, PA, WV ○ 15 ○
Caecidotea fustis Lewis, 1981 Fustis Cave Isopod Isopoda Stygobiont G3 MO ○ 15 ○
Caecidotea hobbsi (Maloney, 1939) Florida Cave Isopod Isopoda Stygobiont G1G2 FL, GA, VA ○ 17 8
Caecidotea insula Lewis, 2013 Kindt's Cave Isopod Isopoda Stygobiont G1 OH ○ 2 ○
Caecidotea jordani (Eberly, 1966) Jordan's Cave Isopod Isopoda Stygobiont G2G3 IN ○ 20 10
Caecidotea kendeighi (Steeves & Seidenberg, 1971) Kendeigh's Groundwater Isopod Isopoda Stygobiont G5 IA, IL, IN, MO, OH ○ 81 9
Caecidotea lesliei Lewis & Bowman, 1981 Isopoda Stygobiont G1G2 IL, MO ○ 3 ○
Caecidotea mackini Lewis, Graening, Fenolio & Bergey, 2006 Isopoda Stygobiont GNR OK ○ 3 ○
Caecidotea macropropoda Chase & Blair, 1937 Bat Cave Isopod Isopoda Stygobiont G3 VU AR, OK ○ 27 ○
Caecidotea metcalfi (Fleming, 1972) Isopoda Stygobiont G1G2 KS ○ 4 ○
Caecidotea nickajackensis Packard, 1881 Nickajack Cave Isopod Isopoda Stygobiont G1 VU AL, TN, AR, GA, KY, MO, VA ○ 9 ○
Caecidotea nordeni Lewis & Bowman, 2010 Norden's Groundwater Isopod Isopoda Stygobiont GH MD ○ 2 ○
Caecidotea nortoni (Steeves, 1966) Norton's Cave Isopod Isopoda Stygobiont G2G3 TN ○ 6 ○
Caecidotea ornatus Lewis & Lewis, 2023 Isopoda Stygobiont G1 VA ○ ○ 21
Caecidotea packardi Mackin & Hubricht, 1940 Isopoda Stygobiont G2G4 IL, MO, KY ○ 34 10
Caecidotea paurotrigona (Fleming, 1972) Isopoda Stygobiont G1G2 LA, MS ○ 3 ○
Caecidotea putea Lewis, 2009 American Waterslaters Isopoda Stygobiont G1G3 FL, GA ○ ○ ○
Caecidotea recurvata (Steeves, 1963) Southwest Virginia Cave Isopod Isopoda Stygobiont G5 TN, VA ○ 120 76
Caecidotea reddelli (Steeves, 1968) Isopoda Stygobiont G3G4 TX ○ 267 11
Caecidotea richardsonae Hay, 1901 Tennessee Valley Cave Isopod Isopoda Stygobiont G5 AL, GA, TN, VA ○ 59 ○
Caecidotea rotunda Bowm& & Lewis, 1984 Frost Cave Isopod Isopoda Stygobiont G2G4 IN, OH ○ 17 32
Caecidotea salemensis Lewis, 1981 Salem Cave Isopod Isopoda Stygobiont G4 AR, MO ○ 33 4
Caecidotea scyphus (Steeves & Holsinger, 1968) Isopoda Stygobiont G1G2 TN, WV ○ ○ ○
Caecidotea serrata (Fleming, 1972) Serrated Cave Isopod Isopoda Stygobiont G1G2 MO ○ 4 ○
Caecidotea simonini (Bresson, 1955) Isopoda Stygobiont G1 WV ○ 3 ○
Caecidotea simulator Lewis, 1999 Isopoda Stygobiont G2G3 AR, KS, OK ○ 7 ○
Caecidotea sinuncus (Steeves, 1965) Isopoda Stygobiont G1 WV ○ 3 ○
Caecidotea spatulata Mackin & Hubricht, 1940 Isopoda Stygobiont G3G4 IL, MO, VA ○ 6 ○
Caecidotea steevesi (Fleming, 1972) Steeves' Cave Isopod Isopoda Stygobiont G3G4 AR, KS, MO, OK ○ 10 ○
Caecidotea stiladactyla Mackin & Hubricht, 1940 Slender-fingered Cave Isopod Isopoda Stygobiont G3G4 AR, MO, OK ○ 64 ○
Caecidotea stygia Packard, 1871 Stygian Cave Isopod Isopoda Stygobiont G5 IL, IN, KY, MO, OH, TN, MS, VA ○ 190 111
Caecidotea teresae Lewis, 1982 Isopoda Stygobiont GX IN ○ 5 ○
Caecidotea tridentata Hungerford, 1922 Isopoda Stygobiont G1G2 IL, KS, AR, MO, OK ○ 38 ○
Calasellus californicus (Miller, 1933) Isopoda Stygobiont G2 CA, OR ○ 20 1
Calasellus longus Bowman, 1981 Isopoda Stygobiont G1 CA ○ 3 ○
Calathaemon holthuisi (Strenth, 1976) Purgatory Cave Shrimp Decapoda Stygobiont G1G2 EN TX ○ 11 ○
Calathus opaculus LeConte, 1854 Coleoptera Trogloxene GNR KY ○ ○ ○
Calicina cloughensis (Briggs & Hom, 1967) Clough Cave Harvestman Opiliones Troglobiont G1 CA ○ 5 7
Califobathynella noodti Cho, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Califobathynella teucherti Cho, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Californibathynella californica Schminke & Noodt, 1988 Bathynellacea Stygobiont CA ○ ○ ○
Calileptoneta briggsi Ledford, 2004 Briggs' Cave Spider Araneae Troglobiont G1 CA ○ 1 ○
Callinectes sapidus Rathbun, 1896 American Blue Crab Decapoda Stygoxene GNR FL ○ ○ ○
Calliphora terraenovae Hall, 1948 Diptera Trogloxene NM ○ ○ ○
Calliphora vicina Robineau-Desvoidy, 1830 Diptera Trogloxene GA, IL ○ ○ ○
Calliphora vomitoria (Linnaeus, 1758) Orange-Bearded Bluebottle Diptera Trogloxene GNR GA, IN ○ ○ ○
Callobius bennetti (Blackwall, 1846) Bennett's Laceweaver Araneae Trogloxene G5 IN ○ ○ ○
Caloptilia alnivorella (Chambers, 1875) Alder Leafminer Lepidoptera Accidental GNR ○ ○ ○
Calymmaria persica (Hentz, 1847) Araneae Troglophile GNR AL, GA, IL, KY, TN, VA ○ ○ ○
Calymmaria virginica Heiss & Draney, 2004 Araneae Trogloxene G1 TN ○ ○ ○
Cambala annulata (Say, 1821) Violet Ridged Millipede Spirostreptida Troglophile G5 AL, FL, GA, IN, KY, TN ○ ○ ○
Cambala hubrichti Hoffman, 1958 Spirostreptida Troglophile G5 GA, NC, TN ○ ○ ○
Cambala minor Bollman, 1888 Amber Ridged Millipede Spirostreptida Troglophile G5 AL, AR, GA, IL, IN, KY, MO, OK, TN, VA, WV ○ ○ ○
Cambala ochra Chamberlin, 1942 Spirostreptida Troglophile AL, GA ○ ○ ○
Cambala reddelli Causey, 1964 Spirostreptida Troglophile NM, TX ○ ○ ○
Cambala speobia Chamberlin, 1953 Spirostreptida Troglobiont G4 TX, Mexico ○ 1.3k ○
Cambarincola alienus Holt, 1963 Branchiobdellida Stygobiont G1G2 KY, TN ○ 9 ○
Cambarincola demissus Hoffman, 1963 Branchiobdellida Stygophile WV ○ ○ ○
Cambarincola dubius Holt, 1973 Branchiobdellida Stygobiont G1G2 IN ○ 8 ○
Cambarincola fallax Hoffman, 1963 Branchiobdellida Stygophile GNR TN, WV ○ ○ ○
Cambarincola heterognathus Hoffman, 1963 Branchiobdellida Stygophile WV ○ ○ ○
Cambarincola holostomus Hoffman, 1963 Branchiobdellida Stygophile TN ○ ○ ○
Cambarincola leoni Holt, 1973 Branchiobdellida Stygobiont G3 FL ○ 13 ○
Cambarincola leptadenus Holt, 1973 Branchiobdellida Stygophile G1 TN ○ ○ ○
Cambarincola manni Holt, 1973 Branchiobdellida Stygoxene FL ○ ○ ○
Cambarincola marthae Holt, 1973 Branchiobdellida Stygobiont G1G2 TN ○ 2 ○
Cambarincola philadelphicus (Leidy, 1851) Branchiobdellida Stygophile FL, IN, KY, TN, VA, WV ○ ○ ○
Cambarincola sheltensis Holt, 1973 Branchiobdellida Stygobiont G1G2 AL ○ 12 ○
Cambarincola steevesi Holt, 1973 Branchiobdellida Stygobiont G3 KY ○ 6 ○
Cambarincola susanae Holt, 1973 Branchiobdellida Stygophile ○ ○ ○
Cambarincola vitrea Ellis, 1918 Branchiobdellida Stygophile FL ○ ○ ○
Cambarus aculabrum Hobbs & Brown, 1987 Benton County Cave Crayfish Decapoda Stygobiont G1 CR ESA AR, MO ○ 11 4
Cambarus bartonii (Fabricius, 1798) Appalachian Brook Crayfish Decapoda Stygophile G5 LC GA, IN, KY, TN, VA, WV ○ ○ ○
Cambarus cryptodytes Hobbs, 1941 Dougherty Plain Cave Crayfish Decapoda Stygobiont G2G3 LC FL, GA ○ 47 6
Cambarus hamulatus (Cope & Packard, 1881) Prickly Cave Crayfish Decapoda Stygobiont G3G4 LC AL, TN, KY ○ 46 58
Cambarus hubbsi Creaser, 1931 Hubbs' Crayfish Decapoda Stygoxene G3 LC ○ ○ ○
Cambarus hubrichti Hobbs, 1952 Salem Cave Crayfish Decapoda Stygobiont G4 DD MO ○ 21 23
Cambarus jonesi Hobbs & Barr, 1960 Alabama Cave Crayfish Decapoda Stygobiont G2 VU AL ○ 70 36
Cambarus laconensis Buhay & Crandall, 2009 Lacon Exit Cave Crayfish Decapoda Stygobiont G1 CR AL ● 8 12
Cambarus latimanus (LeConte, 1856) Variable Crayfish Decapoda Stygoxene G5 LC GA, TN ○ ○ ○
Cambarus nerterius Hobbs, 1964 Greenbrier Cave Crayfish Decapoda Stygobiont G1 NT WV ○ 26 2
Cambarus parvoculus Hobbs & Shoup, 1947 Mountain Midget Crayfish Decapoda Accidental G5 LC KY ○ ○ ○
Cambarus pecki (Hobbs, 1967) Phantom Cave Crayfish Decapoda Stygobiont G1G2 EN AL ○ 16 10
Cambarus setosus Faxon, 1889 Bristly Cave Crayfish Decapoda Stygobiont G3 NT AR, MO, OK ○ 51 11
Cambarus sheltae (Cooper & Cooper, 1997) Shelta Cave Crayfish Decapoda Stygobiont G1 CR AL ○ ○ ○
Cambarus speleocoopi Buhay & Crandall, 2009 Sweet Home Alabama Cave Crayfish Decapoda Stygobiont G1 EN AL ○ 6 12
Cambarus striatus Hay, 1902 Ambiguous Crayfish Decapoda Stygophile G5 LC AL, GA, TN ○ ○ ○
Cambarus subterraneus Hobbs III, 1993 Delaware County Cave Crayfish Decapoda Stygobiont G1 CR OK, KY ○ 7 6
Cambarus tartarus Hobbs & Cooper, 1972 Oklahoma Cave Crayfish Decapoda Stygobiont G1 CR OK ○ 11 10
Cambarus tenebrosus Hay, 1902 Cavespring Crayfish Decapoda Stygophile G5 LC AL, GA, IL, IN, KY, TN ○ ○ ○
Cambarus veitchorum Cooper & Cooper, 1997 White Spring Cave Crayfish Decapoda Stygobiont G1 CR AL ○ 5 ○
Cambarus zophonastes Hobbs & Bedinger, 1964 Hell Creek Cave Crayfish Decapoda Stygobiont G1 CR ESA AR ○ 6 2
Camponotus sayi Emery, 1893 Say's Carpenter Ant Hymenoptera Accidental GNR AR ○ ○ ○
Campostoma anomalum (Rafinesque, 1820) Central Stoneroller Cypriniformes Stygoxene G5 LC KY ○ ○ ○
Campostoma oligolepis Hubbs & Greene, 1935 Largescale Stoneroller Cypriniformes Stygoxene G5 LC AL, KY ○ ○ ○
Canis dirus Leidy, 1858 Dire Wolf Carnivora Accidental ○ ○ ○
Canis familiaris Linnaeus, 1758 Domestic Dog Carnivora Accidental GNA AL ○ ○ ○
Canis latrans Say, 1823 Coyote Carnivora Accidental G5 LC KY, OK ○ ○ ○
Canis rufus Audubon & Bachman, 1851 Red Wolf Carnivora Accidental G1 CR AL ○ ○ ○
Canthon imitator Brown, 1946 Coleoptera Trogloxene NM ○ ○ ○
Carcinophora americana (Palisot de Beauvois) Dermaptera Trogloxene ○ ○ ○
Carebara urichi (Wheeler, 1922) Hymenoptera Accidental ○ ○ ○
Carnarimelita janstocki Bousfield, 1990 Amphipoda Stygobiont HI ○ 6 ○
Carphophis amoenus (Say, 1825) Common Wormsnake Squamata Accidental G5 LC AL, GA ○ ○ ○
Cartodere constricta (Gyllenhal, 1827) Plaster Beetle Coleoptera Accidental GNR IA ○ ○ ○
Carychium clappi Hubricht, 1959 Appalachian Thorn Basommatophora Troglophile G5 AL ○ ○ ○
Carychium exile H. C. Lea, 1842 Ice Thorn Basommatophora Troglophile G5 AL, IL, KY, TN ○ ○ ○
Carychium mexicanum Pilsbry, 1891 Southern Thorn Basommatophora Trogloxene G5 AL, IL ○ ○ ○
Carychium nannodes G. H. Clapp, 1905 File Thorn Basommatophora Trogloxene G5 IL, TN ○ ○ ○
Carychium riparium Hubricht, 1978 Floodplain Thorn Basommatophora Trogloxene G2 IL ○ ○ ○
Carychium stygium Call, 1897 Cave thorn Basommatophora Troglobiont G3 AL, KY, TN ○ 72 6
Castor canadensis Kuhl, 1820 American Beaver Rodentia Stygoxene G5 LC AL, FL, GA, IL, TN ○ ○ ○
Cathartes aura (Linnaeus, 1758) Turkey Vulture Accipitriformes Trogloxene G5 LC GA, IL, TN ○ ○ ○
Catopocerus alabamae Peck, 1975 Coleoptera Troglobiont GNR AL ○ 1 ○
Catopocerus appalachianus Peck, 1975 Coleoptera Troglophile GNR AL, TN ○ ○ ○
Catopocerus jonesi Peck, 1975 Coleoptera Troglobiont GNR AL ○ 1 ○
Catops gratiosus (Blanchard, F., 1915) Coleoptera Troglophile G5 AL, GA, IA, IL, IN, KY, MA, MD, NY, OH, PA, TN, VA, WV ○ ○ ○
Catostomus commersonii (Lacepède, 1803) White Sucker Cypriniformes Stygoxene G5 LC AL, KY, TN ○ ○ ○
Cauloxenus stygius Cope, 1872 Northern Cavefish Copepod Siphonostomatoida Stygobiont G1G2 IN, KY ○ ○ ○
Causeyella causeyae Shear, 2003 Chordeumatida Troglobiont AR ○ ○ ○
Causeyella dendropus (Loomis, 1939) Causeyella Cave Millipede Chordeumatida Troglobiont G3 AR, MO ○ 1 ○
Causeyella youngsteadtorum Shear, 2003 Chordeumatida Troglobiont AR ○ ○ ○
Cavaticovelia aaa (Gagne & Howarth, 1975) Heteroptera Troglobiont G1G2 NE HI ○ 8 ○
Cavenderia aureostipes (Cavender, Raper & Norberg) S.Baldauf, S.Sheikh & Thulin, 2018 Acytosteliales Stygophile GA ○ ○ ○
Cavernicola pilosa Barber, 1937 Barbeiro Heteroptera Trogloxene ○ ○ ○
Cemophora coccinea (Blumenbach, 1788) Scarlet Snake Squamata Accidental G5 LC GA ○ ○ ○
Centromerus cornupalpis (O. Pickard-Cambridge, 1875) Eastern Spurred Sheetweaver Araneae Troglophile GNR AL, IL ○ ○ ○
Centromerus denticulatus (Emerton, 1909) Araneae Troglophile GNR AL, GA ○ ○ ○
Centromerus latidens (Emerton, 1882) Araneae Troglophile G5 AL, FL, GA, IL, KY, MO, TN ○ ○ ○
Centromerus tennapex (Barrows, 1940) Araneae Trogloxene TN ○ ○ ○
Centropyxis aculeata (Ehrenberg, 1832) Stein, 1859 Arcellinida Stygoxene KY ○ ○ ○
Ceraticelus fissiceps (O. Pickard-Cambridge, 1874) Araneae Troglophile G5 AL ○ ○ ○
Ceraticelus minutus (Emerton, 1882) Araneae Troglophile G5 ○ ○ ○
Ceratophysella boletivora (Packard, AS, 1873) Springtail Poduromorpha Trogloxene GNR IL ○ ○ ○
Ceratophysella brevis (Christiansen, K & Bellinger, P, 1980) Poduromorpha Trogloxene GNR IL ○ ○ ○
Ceratophysella denticulata Ardanaz, 1986 Poduromorpha Troglophile IA, IL, TN ○ ○ ○
Ceratophysella engadinensis (Gisin, 1949) Poduromorpha Trogloxene IL ○ ○ ○
Ceratophysella lucifuga (Packard, 1888) Wyandotte Cave Springtail Poduromorpha Troglobiont GNR IN, IL ○ 2 ○
Ceratozetes gracilis (Michael, 1884) Sarcoptiformes Troglophile GNR IA, KY ○ ○ ○
Cerobasis guestfalica (Kolbe, 1880) Book Lice Psocodea Trogloxene KY ○ ○ ○
Ceuthophilus brevipes Scudder, 1862 Short-Legged Camel Cricket Orthoptera Trogloxene G5 ○ ○ ○
Ceuthophilus carlsbadensis Caudell, 1924 Orthoptera Trogloxene NM, TX ○ ○ ○
Ceuthophilus conicaudus Hubbell, 1936 Orthoptera Trogloxene NM ○ ○ ○
Ceuthophilus elegans Hubbell, 1934 Orthoptera Trogloxene IL, MO, WI ○ ○ ○
Ceuthophilus ensifer Packard, 1881 Orthoptera Trogloxene AL, GA, TN ○ ○ ○
Ceuthophilus gracilipes (Haldeman, 1850) Slender-Legged Camel Cricket Orthoptera Trogloxene GNR AL, FL, GA, IL, TN ○ ○ ○
Ceuthophilus isletae Hubbell, 1936 Orthoptera Trogloxene NM ○ ○ ○
Ceuthophilus latens Scudder, 1862 Black-Sided Camel Cricket Orthoptera Trogloxene GNR AL, VA ○ ○ ○
Ceuthophilus latibuli Scudder, 1894 Orthoptera Trogloxene FL ○ ○ ○
Ceuthophilus longipes Caudell, 1924 Orthoptera Trogloxene GNR NM ○ ○ ○
Ceuthophilus maculatus (Harris, 1835) Spotted Camel Cricket Orthoptera Trogloxene G5 ○ ○ ○
Ceuthophilus pallidipes Walker, 1905 Orthoptera Trogloxene KY ○ ○ ○
Ceuthophilus palmeri Scudder, 1894 Orthoptera Trogloxene ○ ○ ○
Ceuthophilus seclusus Scudder, 1894 Orthoptera Trogloxene GNR IL ○ ○ ○
Ceuthophilus stygius (Scudder, 1861) Cave Camel Cricket Orthoptera Trogloxene GNR AL, IN, KY, TN ○ ○ ○
Ceuthophilus virgatipes Rehn & Hebard, 1905 Orthoptera Trogloxene FL ○ ○ ○
Ceuthophilus williamsoni Hubbell, 1934 Orthoptera Trogloxene GNR IL ○ ○ ○
Chaetaspis aleyorum Lewis, 2002 Aley's Cave Milliped Polydesmida Troglobiont G1 MO ○ ○ ○
Chaetaspis attenuatus Lewis & Slay, 2013 Polydesmida Troglobiont AR ○ ○ ○
Chaetaspis debilis (Causey, 1959) Polydesmida Troglobiont TN ○ ○ ○
Chaetaspis fragilis (Loomis, 1943) Polydesmida Troglobiont GNR KY ○ 2 ○
Chaetaspis mollis (Causey, 1959) Polydesmida Troglobiont G1 TN, AL ○ ○ ○
Chaetogaster diaphanus (Gruithuisen, 1828) Aquatic Oligochaete Worm Tubificida Stygophile GNR SC ○ ○ ○
Chaoborus albatus Johnson, 1921 Diptera Trogloxene GNR ○ ○ ○
Chauliodes pectinicornis (Linnaeus, 1763) Summer Fishfly Megaloptera Accidental GNR ○ ○ ○
Cheiracanthium inclusum (Hentz, 1847) Yellow Sac Spider Araneae Accidental GNR AL ○ ○ ○
Cheiridium reyesi Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Chelydra serpentina (Linnaeus, 1758) North American Snapping Turtle Testudines Accidental G4 LC FL, KY ○ ○ ○
Cherokia georgiana (Bollman, 1889) Polydesmida Accidental G4 GA ○ ○ ○
Cheumatopsyche minuscula (Banks, 1907) Trichoptera Stygoxene G5 ○ ○ ○
Chilodonella caudata Stokes Chlamydodontida Stygoxene AL ○ ○ ○
Chinquipellobunus madlae (Goodnight & Goodnight, 1967) Opiliones Troglobiont G4 TX ○ 284 ○
Chinquipellobunus russelli (Goodnight & Goodnight, 1967) Opiliones Troglobiont G2G3 TX ○ 21 ○
Chionea scita Walker, 1848 Diptera Trogloxene GNR KY ○ ○ ○
Chironomus decorus Johannsen, 1905 Diptera Trogloxene GNR GA ○ ○ ○
Chironomus plumosus (Linnaeus, 1758) Chironome Plumosus Diptera Trogloxene ○ ○ ○
Chiroptonyssus robustipes (Ewing, 1925) Mesostigmata Trogloxene NM, TX ○ ○ ○
Chitrella archeri Malcolm & J. C. Chamberlin, 1960 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Chitrella cavicola (Packard, 1884) Pseudoscorpiones Troglophile G3 ○ ○ ○
Chitrella elliotti Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Chitrella major Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Chitrella regina Malcolm & Chamberline, 1960 Royal Syarinid Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Chitrella superba Muchmore, 1973 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Chitrella welbourni Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 NM ○ ○ ○
Chitrellina chiricahuae Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Chlaenius aestivus Say, 1823 Summer Harp Ground Beetle Coleoptera Accidental GNR IL ○ ○ ○
Chlamydotheca texasiensis Baird, 1862 Podocopida Stygophile TX ○ ○ ○
Choctella cumminsi Chamberlin, 1918 Spirostreptida Trogloxene TN ○ ○ ○
Chrosomus erythrogaster (Rafinesque, 1820) Southern Redbelly Dace Cypriniformes Accidental G5 LC AL ○ ○ ○
Chrysops univittatus Macquart, 1855 One-Striped Deer Fly Diptera Accidental G5 TN ○ ○ ○
Chthonius cryptus Chamberlin, 1962 Pseudoscorpiones Troglophile ○ ○ ○
Chthonius paludis (J. C. Chamberlin, 1929) Pseudoscorpiones Troglophile GA, TN ○ ○ ○
Cicurina arcata Chamberlin & Ivie, 1940 Araneae Troglophile G5 IL ○ ○ ○
Cicurina arcuata Keyserling, 1887 Araneae Troglophile G5 GA, IL, TN ○ ○ ○
Cicurina bandera Gertsch, 1992 Araneae Troglobiont G2G3 TX ○ 2 ○
Cicurina bandida Gertsch, 1992 Bandit Cave Meshweaver Araneae Troglobiont G2 TX ○ 12 ○
Cicurina baronia Gertsch, 1992 Robber Baron Cave Meshweaver Araneae Troglobiont G1 ESA TX ○ 12 ○
Cicurina barri Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina breviaria Bishop & Crosby, 1926 Araneae Troglophile AL, TN ○ ○ ○
Cicurina brevis (Emerton, 1890) Small-Eared Meshweaver Araneae Troglophile G5 IL ○ ○ ○
Cicurina browni Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina brunsi Cokendolpher, 2004 Araneae Troglobiont TX ○ 4 ○
Cicurina bullis Cokendolpher, 2004 Araneae Troglobiont TX ○ 14 ○
Cicurina buwata Chamberlin & Ivie, 1940 Araneae Troglobiont G1G2 TX ○ 1 107
Cicurina caliga Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 8 3
Cicurina cavealis Bishop & Crosby, 1926 Araneae Troglophile GNR IL ○ ○ ○
Cicurina caverna Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina coryelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 5 1
Cicurina delrio Gertsch, 1992 Araneae Troglobiont TX ○ ○ ○
Cicurina ezelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ 1
Cicurina gruta Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina holsingeri Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina hoodensis Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 35 11
Cicurina joya Gertsch, 1992 Araneae Troglobiont TX ○ ○ ○
Cicurina loftini Cokendolpher, 2004 Araneae Troglobiont TX ○ ○ ○
Cicurina machete Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina madla Gertsch, 1992 Madla Cave Meshweaver Araneae Troglobiont G1 ESA TX ○ 23 22
Cicurina mckenziei Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina medina Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina menardia Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina minima Chamberlin & Ivie, 1940 Araneae Troglophile GNR AL ○ ○ ○
Cicurina mirifica Gertsch, 1992 Araneae Troglobiont TX ○ ○ ○
Cicurina mixmaster Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 3 1
Cicurina neovespera Cokendolpher, 2004 Araneae Troglobiont TX ○ 4 ○
Cicurina obscura Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina orellia Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina pablo Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina pallida Keyserling, 1887 Araneae Troglophile GNR GA, IL, IN, KY ○ ○ ○
Cicurina pastura Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina patei Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina placida Banks, 1892 Araneae Troglophile GNR TN ○ ○ ○
Cicurina platypus Cokendolpher, 2004 Araneae Troglobiont TX ○ 1 ○
Cicurina porteri Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina puentecilla Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 4 7
Cicurina rainesi Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 ○
Cicurina reclusa Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina reddelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ 1
Cicurina russelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina sansaba Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina selecta Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina serena Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina sheari Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina sprousei Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 ○
Cicurina stowersi Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina suttoni Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina travisae Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 11 208
Cicurina troglobia Cokendolpher, 2004 Araneae Troglobiont GNR TX ○ 2 ○
Cicurina ubicki Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina uvalde Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 ○
Cicurina varians Gertsch & Mulaik, 1940 Araneae Troglophile GNR NM ○ ○ ○
Cicurina venefica Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina vespera Gertsch, 1992 Government Canyon Bat Cave Meshweaver Araneae Troglobiont G1 ESA TX ○ 6 ○
Cicurina vibora Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 4
Cicurina wartoni Gertsch, 1992 Warton Cave Meshweaver Araneae Troglobiont G1 TX ○ ○ ○
Cicurina watersi Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina wiltoni Gertsch, 1992 Araneae Troglobiont G1 AL ○ ○ ○
Cirolanides texensis Benedict, 1896 Isopoda Stygobiont G4 TX, Mexico ○ 431 130
Cirolanides wassenichae Schwartz et al., 2019 Isopoda Stygobiont TX ○ 76 38
Cladura flavoferruginea Osten Sacken, 1860 Diptera Trogloxene GNR IN ○ ○ ○
Cleidogona baroqua Shear, 1972 Chordeumatida Trogloxene ○ ○ ○
Cleidogona crucis (Chamberlin, 1942) Chordeumatida Trogloxene ○ ○ ○
Cleidogona crystallina Shear, 1972 Chordeumatida Trogloxene ○ ○ ○
Cleidogona hoffmani Shear, 1972 Chordeumatida Trogloxene G3 VA ○ ○ ○
Cleidogona mayapec Shear, 1972 Chordeumatida Trogloxene ○ ○ ○
Cleidogona pecki Shear, 1972 Chordeumatida Trogloxene ○ ○ ○
Cleidogona unita Causey, 1951 Chordeumatida Troglophile IL ○ ○ ○
Clethrionomys gapperi (Vigors, 1830) Rodentia Accidental ○ ○ ○
Climacia areolaris (Hagen, 1861) Neuroptera Accidental GNR ○ ○ ○
Coecobrya kukae (Christiansen & Bellinger, 1992) Entomobryomorpha Troglobiont G1G2 HI ○ 1 ○
Coecobrya lua (Christiansen & Bellinger, 1992) Entomobryomorpha Troglobiont G1G2 HI ○ 1 ○
Coecobrya nupa (Christiansen & Bellinger, 1992) Entomobryomorpha Troglobiont G1G2 HI ○ 1 ○
Coecobrya tenebricosa (Folsom, JW, 1902) Entomobryomorpha Troglophile GNR AL, IL ○ ○ ○
Colacium vesiculosum Ehrenb. Euglenales Stygoxene KY ○ ○ ○
Coloradesmus aquiliensis (Shear, 1984) Polydesmida Troglobiont G1G2 CO ○ 7 ○
Coloradesmus beckleyi Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 2 ○
Coloradesmus hopkinsae Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 3 ○
Coloradesmus manitou Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 6 ○
Coloradesmus warneri Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 7 ○
Coluber constrictor Linnaeus, 1758 North American Racer Squamata Accidental G5 LC FL ○ ○ ○
Columba livia J. F. Gmelin, 1789 Common Pigeon Columbiformes Trogloxene AR, IL ○ ○ ○
Columbasellus acheron Lewis et al., 2003 Isopoda Stygobiont WA ○ 1 ○
Colusa illecta (Casey, 1911) Coleoptera Troglophile AL ○ ○ ○
Comalcandona gibsoni Kulkoyluoglu, 2019 Gibson's Groundwater Ostracod Podocopida Stygobiont TX ○ ○ ○
Comalcandona mixoni Kulkoyluoglu et al., 2019 Podocopida Stygobiont TX ○ ○ ○
Comalcandona tressleri Kulkoyluoglu & Gibson, 2018 Podocopida Stygobiont TX ○ ○ ○
Comaldessus stygius Spangler & Barr, 1995 Comal Springs Diving Beetle Coleoptera Stygobiont G1 TX ○ 3 3
Conasellus burkensis (Lewis & Lewis, 2021) Isopoda Stygobiont GNR VA ○ ○ ○
Conasellus communis (Say, 1818) Eastern Waterslater Isopoda Stygoxene NY ○ ○ ○
Conasellus forbesi (Williams, 1970) Forbes' Waterslater Isopoda Stygoxene TN ○ ○ ○
Conasellus jeffersoni (Lewis, 2009) Isopoda Stygobiont VA ○ ○ ○
Conasellus kenki (Bowman, 1967) Isopoda Stygobiont G3 DC, MD, PA, VA ○ ○ ○
Conasellus phreaticus (Lewis & Holsinger, 1985) Isopoda Stygobiont G2G3 VA ○ ○ ○
Conasellus pricei (Levi, 1949) Price's Cave Isopod Isopoda Stygobiont G5 MD, PA, VA, WV ○ ○ ○
Conasellus washingtoni Lewis & Lewis, 2023 Isopoda Stygobiont DC ○ ○ ○
Condeicampa langei Ferguson, 1996 Rhabdura Troglobiont GNR TN ○ ○ ○
Conicera similis Haliday, 1833 Diptera Trogloxene AL ○ ○ ○
Conosoma velocipes Casey, 1895 Coleoptera Accidental ○ ○ ○
Conotyla blakei (Verhoeff, 1932) Chordeumatida Troglophile G4 MD, PA ○ ○ ○
Conotyla bollmani (McNeill, 1887) Bollman's Cave Millipede Chordeumatida Troglobiont G3 IN ○ 1 ○
Conotyla vista Shear, 1971 Chordeumatida Troglobiont GH WV ○ 1 ○
Copestylum vesicularium (Curran, 1947) Iridescent Bromeliad Fly Diptera Accidental G5 GA ○ ○ ○
Coquillettidia perturbans (Walker, 1856) Cattail Mosquito Diptera Trogloxene G5 ○ ○ ○
Coragyps atratus (Bechstein, 1793) Black Vulture Accipitriformes Trogloxene G5 LC AL, FL ○ ○ ○
Coras juvenilis (Keyserling, 1881) Araneae Troglophile GNR GA, KY ○ ○ ○
Coras lamellosus (Keyserling, 1887) Araneae Troglophile GNR AL, IL ○ ○ ○
Coras taugynus Chamberlin, 1925 Araneae Troglophile TN ○ ○ ○
Corbicula fluminea (O. F. Müller, 1774) Asian Basket Clam Venerida Stygoxene G5 LC AL, FL ○ ○ ○
Cordylophora caspia (Pallas, 1771) Ponto-Caspian Freshwater Hydroid Anthoathecata Stygoxene GNR FL ○ ○ ○
Corydalus cornutus (Linnaeus, 1758) Dobsonfly Megaloptera Accidental GNR GA ○ ○ ○
Corynorhinus rafinesquii (Lesson, 1827) Eastern Big-Eared Bat Chiroptera Trogloxene G3 LC AL, AR, GA, IN, KY, NC, TN ○ ○ ○
Corynorhinus townsendii (Cooper, 1837) Pacific Townsend's Big-Eared Bat Chiroptera Trogloxene G4 LC AR, KY, OK, PA ○ ○ ○
Cottus bairdii Girard, 1850 Mottled Sculpin Scorpaeniformes Stygophile G5 LC AL, GA, KY, TN ○ ○ ○
Cottus carolinae (Gill, 1861) Banded Sculpin Scorpaeniformes Stygophile G5 LC AL, GA, IL, KY, OK, TN ○ ○ ○
Cottus specus Adams et al. 2013 Grotto sculpin Scorpaeniformes Stygobiont G2 ESA MO ○ 10 21
Crangonyx anomalus Hubricht, 1943 Large Male Crangonyctid Amphipoda Stygophile G4 IL, IN, OH ○ ○ ○
Crangonyx antennatus Packard, 1881 Appalachian Valley crangonyctid Amphipoda Stygobiont G5 AL, GA, TN, VA ○ 222 1
Crangonyx baculispina Zhang & Holsinger, 2003 Lancaster County Amphipod Amphipoda Stygoxene G1 VA ○ ○ ○
Crangonyx barri Zhang & Holsinger, 2003 Barr's Cave Amphipod Amphipoda Stygobiont G5 IN, KY ○ 141 ○
Crangonyx bousfieldi Zhang & Holsinger, 2003 Amphipoda Stygophile G2 OH ○ ○ ○
Crangonyx caecus Zhang & Holsinger, 2003 Amphipoda Stygobiont G1 KY ○ 19 ○
Crangonyx castellanum Zhang & Holsinger, 2003 Amphipoda Stygobiont G1 KY ○ 13 ○
Crangonyx consimilis Zhang & Holsinger, 2003 Amphipoda Stygoxene G4 AL, GA, TN ○ ○ ○
Crangonyx cooperi Zhang & Holsinger, 2003 Amphipoda Stygophile G2 AL ○ ○ ○
Crangonyx dearolfi Shoemaker, 1942 Pennsylvania Cave Amphipod Amphipoda Stygobiont G2 EN MD, PA ○ 14 ○
Crangonyx floridanus Bousfield, 1963 Florida Crangonyctid Amphipoda Stygophile G5 AL, FL, KY, MO, NC, PA, SC, TN ○ ○ ○
Crangonyx forbesi (Hubricht & Mackin, 1940) Forbes Crangonyctid Amphipoda Stygophile G5 IL, IN, KY, MO, OH, OK ○ ○ ○
Crangonyx gracilis S.I. Smith, 1871 Northern Lake Crangonyctid Amphipoda Stygophile G4 IN, KY, WV ○ ○ ○
Crangonyx grandimanus Bousfield, 1963 Florida Cave Amphipod Amphipoda Stygobiont G2G3 VU FL ○ 67 13
Crangonyx hobbsi Shoemaker, 1941 Hobb's Cave Amphipod Amphipoda Stygobiont G2G3 VU FL ○ 80 146
Crangonyx indianensis Zhang & Holsinger, 2003 Indiana Cave Amphipod Amphipoda Stygobiont G3 IN ○ 33 ○
Crangonyx insolitus Zhang & Holsinger, 2003 Amphipoda Stygophile G1 MO ○ ○ ○
Crangonyx lewisi Zhang & Holsinger, 2003 Lewis' Cave Amphipod Amphipoda Stygobiont G2 IN, KY ○ 15 ○
Crangonyx longidactylus Zhang & Holsinger, 2003 Amphipoda Stygoxene G2 KY ○ ○ ○
Crangonyx manubrium Cannizzaro & Sawicki in Cannizzaro, Balding, Lazo-Wasem & Sawicki, 2019 Jackson County Cave Amphipod Amphipoda Stygobiont G1G2 FL ○ 32 123
Crangonyx minor Bousfield, 1958 Small Stream Crangonyctid Amphipoda Stygoxene G5 IL ○ ○ ○
Crangonyx montanus Zhang & Holsinger, 2003 Amphipoda Stygoxene G2 WV ○ ○ ○
Crangonyx obliquus (Hubricht & Mackin, 1940) Oblique-Hand Crangonyctid Amphipoda Stygophile G5 AR ○ ○ ○
Crangonyx ohioensis Zhang & Holsinger, 2003 Ohio Cave Amphipod Amphipoda Stygobiont G1G2 IN, KY, OH ○ 32 ○
Crangonyx packardi Smith, 1888 Packard's Cave Amphipod Amphipoda Stygobiont G4 IL, IN, KS, KY, MO ○ 103 ○
Crangonyx palustris Zhang & Holsinger, 2003 Amphipoda Stygoxene G4 NC ○ ○ ○
Crangonyx parhobbsi Cannizzaro & Sawicki in Cannizzaro, Balding, Lazo-Wasem & Sawicki, 2020 Florida Big Bend Cave Amphipod Amphipoda Stygobiont G1G2 FL, GA ○ 13 89
Crangonyx pseudogracilis Bousfield, 1958 Northern River Crangonyctid Amphipoda Stygoxene G5 KY ○ ○ ○
Crangonyx richmondensis Ellis, 1941 Alpine Lake Crangonyctid Amphipoda Stygoxene G5 AK, FL ○ ○ ○
Crangonyx rivularis Bousfield, 1958 Ontario Stream Crangonyctid Amphipoda Stygoxene G5 KY ○ ○ ○
Crangonyx setodactylus Bousfield, 1958 Hairy-Fingered Crangonyctid Amphipoda Stygoxene G5 OH ○ ○ ○
Crangonyx shoemakeri (Hubricht & Mackin, 1940) Shoemaker Crangonyctid Amphipoda Stygophile G5 IA, IL, IN, MI, OH, VA ○ ○ ○
Crangonyx specus Zhang & Holsinger, 2003 Amphipoda Stygobiont G1 KY ○ 23 ○
Crangonyx sulphurium Sawicki & Holsinger, 2017 Sulphurous Cave Amphipod Amphipoda Stygobiont G1 FL ○ 6 ○
Creaserinus fodiens (Cottle, 1863) Digger Crayfish Decapoda Accidental G5 IL ○ ○ ○
Creophilus maxillosus (Linnaeus, 1758) Hairy Rove Beetle Coleoptera Troglophile GA ○ ○ ○
Crosbycus goodnighti Roewer, 1951 Opiliones Troglophile VA ○ ○ ○
Crosbyella distincta Goodnight & Goodnight, 1942 Opiliones Troglobiont G1G2 AR ○ ○ ○
Crosbyella roeweri Goodnight & Goodnight, 1967 Opiliones Troglobiont G1G2 AR ○ ○ ○
Crosbyella spinturnix (Crosby & Bishop, 1924) Opiliones Troglophile GNR AL, AR, FL, GA ○ ○ ○
Crotalus adamanteus Palisot de Beauvois, 1799 Eastern Diamond-Backed Rattlesnake Squamata Accidental G3 LC FL ○ ○ ○
Crotalus horridus Linnaeus, 1758 Timber Rattlesnake Squamata Accidental G4 LC TN ○ ○ ○
Cryptachaea porteri (Banks, 1896) Cobweb Spiders Araneae Troglophile GNR AL, GA, IN, NM, TN ○ ○ ○
Cryptachaea rupicola (Emerton, 1882) Araneae Troglophile GNR AL, IL ○ ○ ○
Cryptobranchus alleganiensis (Sonnini de Manoncourt & Latreille, 1801) Hellbender Caudata Accidental G3 VU AL ○ ○ ○
Cryptocreagris destica Harvey & Muchmore, 2010 Pseudoscorpiones Troglobiont CO ○ 3 ○
Cryptocreagris steinmanni Harvey & Muchmore, 2010 Pseudoscorpiones Troglobiont CO ○ 3 ○
Cryptophagus valens Casey, 1900 Coleoptera Trogloxene GNR IL ○ ○ ○
Ctenophthalmus pseudagyrtes Baker, 1904 Siphonaptera Trogloxene GNR AL, GA, TN ○ ○ ○
Ctenus captiosus Gertsch, 1935 Florida False Wolf Spider Araneae Trogloxene FL ○ ○ ○
Culex erraticus (Dyar & Knab, 1906) Diptera Trogloxene GNR TN ○ ○ ○
Culex pipiens Linnaeus, 1758 Northern House Mosquito Diptera Trogloxene G5 IL, IN, TN ○ ○ ○
Culex restuans Theobald, 1901 White-Dotted Mosquito Diptera Trogloxene G5 TN ○ ○ ○
Culex territans Walker, 1856 Northern Frog-Biting Mosquito Diptera Trogloxene G5 GA ○ ○ ○
Culicoides paraensis (Goeldi, 1905) Diptera Trogloxene TN ○ ○ ○
Culiseta impatiens (Walker, 1848) Impatient Mosquito Diptera Trogloxene G5 ○ ○ ○
Cybaeopsis pantopla (Bishop & Crosby, 1935) Araneae Trogloxene G3 TN ○ ○ ○
Cybaeopsis tibialis (Emerton, 1888) Araneae Trogloxene GNR ○ ○ ○
Cybaeus giganteus Banks, 1892 Araneae Accidental GNR KY, TN ○ ○ ○
Cybaeus patritus Bishop & Crosby, 1926 Araneae Trogloxene TN ○ ○ ○
Cyclidium glaucoma Schewiakoff, 1889 Pleuronematida Stygoxene AL ○ ○ ○
Cyclocosmia truncata (Hentz, 1841) Ravine Trapdoor Spider Araneae Trogloxene GNR GA ○ ○ ○
Cyclocypris sharpei Furtos, 1933 Podocopida Stygoxene GNR TN ○ ○ ○
Cyclops cavernarum Ulrich, 1902 Cyclopoida Stygobiont GNR TX ○ ○ ○
Cyclops learii Ulrich, 1902 Cyclopoida Stygobiont TX ○ ○ ○
Cyclosa conica (Pallas, 1772) Cone-Shaped Orbweaver Araneae Accidental G5 ○ ○ ○
Cyclotrachelus fucatus (Freitag, 1969) Coleoptera Accidental GNR IL ○ ○ ○
Cyclotrachelus sodalis (LeConte, 1846) Blue-Tinged Harp Ground Beetle Coleoptera Accidental GNR IL ○ ○ ○
Cylindrodesmus hirsutus Pocock, 1889 Polydesmida Trogloxene ○ ○ ○
Cylindroiulus arborum Verhoeff, 1928 Julida Trogloxene IN ○ ○ ○
Cylisticus convexus (De Geer, 1778) Curly Woodlouse Isopoda Troglophile G5 AL, GA, IL, IN, KY, TN, VA ○ ○ ○
Cyllopsis gemma (Hübner, 1809) Gemmed Satyr Lepidoptera Accidental G5 TN ○ ○ ○
Cyphoderus inaequalis Folsom, JW, 1927 Entomobryomorpha Troglophile ○ ○ ○
Cyphoderus similis Denis, 1931 Entomobryomorpha Troglophile ○ ○ ○
Cypria lacrima Kulkoyluoglu et al., 2017 Podocopida Stygobiont GNR TX ○ ○ ○
Cypridopsis vidua (Müller, 1776) Cypris Veuve (La) Podocopida Stygophile GNR TX ○ ○ ○
Cyprinella spiloptera (Cope, 1867) Spotfin Shiner Cypriniformes Accidental G5 LC AL, KY ○ ○ ○
Cyprinodon diabolis Wales, 1930 Devil's Hole Pupfish Cyprinodontiformes Stygoxene G1 CR NV ○ ○ ○
Cyprinus carpio Linnaeus, 1758 Eurasian Carp Cypriniformes Accidental G5 LC AL ○ ○ ○
Cyrtodesmus archimedes (Chamberlin, 1940) Polydesmida Troglophile ○ ○ ○
Dactylocythere arcuata (C. W. Hart & Hobbs, 1961) Podocopida Stygophile G1 AL ○ ○ ○
Dactylocythere prionata (C. W. Hart & Hobbs, 1961) Podocopida Stygophile G2 KY ○ ○ ○
Dactylocythere steevesi (C. W. Hart & Hobbs, 1961) Podocopida Stygophile G4 AL, KY, TN ○ ○ ○
Dactylocythere ungulata (C. W. Hart & Hobbs, 1961) Podocopida Stygophile KY, TN ○ ○ ○
Dactylolabis hudsonica Alexander, 1931 Diptera Trogloxene GNR ○ ○ ○
Dactylolabis montana (Osten Sacken, 1860) Diptera Trogloxene GNR MO ○ ○ ○
Darlingtonea kentuckensis Valentine, 1952 Coleoptera Troglobiont G3G4 KY, TN ○ 15 88
Darwinula stevensoni (Brady & Robertson, 1870) Podocopida Stygophile TX ○ ○ ○
Dasycerus nikwasi Caterino & Harden, 2024 Coleoptera Trogloxene GNR AL ○ ○ ○
Dasypus novemcinctus Linnaeus, 1758 Nine-Banded Armadillo Cingulata Accidental G5 LC FL ○ ○ ○
Dasyscias franzi F. G. Thompson & Hershler, 1991 Shaggy ghostsnail Littorinimorpha Stygobiont G1 FL ○ 8 ○
Demogorgonichthys arcanus Hart, Niemiller, Sandel & Armbruster, 2026 Demon Cavefish Percopsiformes Stygobiont AL ○ ○ ○
Dendrobaena octaedra (Savigny in Cuvier, 1826) Octagonal-Tail Worm Crassiclitellata Troglophile GNR GA, NC, VA ○ ○ ○
Dendrocoelopsis americana (Hyman, 1939) Tricladida Stygobiont G3 AR, MO, OK, TX, MD ○ 30 ○
Dendrocoelopsis hymanae Kawakatsu, 1968 Lake Tahoe planarian Tricladida Stygobiont GNR CA, NV ○ 3 ○
Dendronucleata americana Moravec & Huffman, 2000 Neoechinorhynchida Stygobiont TX ○ 2 ○
Dermacentor variabilis Say, 1821 American Dog Tick Ixodida Accidental GA, IL ○ ○ ○
Deroceras laeve (O. F. Müller, 1774) Meadow Slug Stylommatophora Trogloxene IL ○ ○ ○
Deroceras reticulatum (O. F. Müller, 1774) Gray Fieldslug Stylommatophora Trogloxene ○ ○ ○
Derops divalis (Sanderson, 1947) Coleoptera Troglophile GNR IL ○ ○ ○
Desmognathus abditus Anderson & Tilley, 2003 Cumberland Dusky Salamander Caudata Accidental G2 VU ○ ○ ○
Desmognathus brimleyorum Stejneger, 1895 Ouachita Dusky Salamander Caudata Trogloxene G3 LC ○ ○ ○
Desmognathus conanti Rossman, 1958 Spotted Dusky Salamander Caudata Trogloxene G4 LC AL, GA, TN ○ ○ ○
Desmognathus fuscus (Green, 1818) Northern Dusky Salamander Caudata Trogloxene KY, OH ○ ○ ○
Desmognathus gvnigeusgwotli Pyron & Beamer, 2022 Cherokee Blackbelly Salamander Caudata Accidental G3 TN ○ ○ ○
Desmognathus monticola Dunn, 1916 Seal Salamander Caudata Trogloxene G5 LC AL, KY ○ ○ ○
Desmognathus ocoee Nicholls, 1949 Ocoee Salamander Caudata Accidental G4 LC AL, GA ○ ○ ○
Desoria trispinata (Mac Gillivray, 1896) Entomobryomorpha Troglophile GNR IL, TN ○ ○ ○
Desoria truncata Waltz & Hart, 1995 Entomobryomorpha Troglophile GNR TN ○ ○ ○
Desoria violacea (Tullberg, T, 1877) Entomobryomorpha Trogloxene GNR ○ ○ ○
Deuteraphorura jana (Christiansen, K & Bellinger, P, 1980) Poduromorpha Troglophile AL ○ ○ ○
Diacyclops alabamensis Reid, 1992 Cyclopoida Stygobiont AL ○ 2 ○
Diacyclops bicuspidatus (Claus, 1857) Damhops Cyclopoida Stygophile GNR IN, KY ○ ○ ○
Diacyclops chrisae Reid, 1992 Cyclopoida Stygobiont MD, GA, IN ○ 5 ○
Diacyclops clandestinus (Kiefer, 1926) Cyclopoida Stygophile IL, MO, TN ○ ○ ○
Diacyclops conversus Reid, 2004 Blue River Groundwater Copepod Cyclopoida Stygobiont G2 IN ○ 3 ○
Diacyclops crassicaudis (Sars G.O., 1863) Kompakthops Cyclopoida Stygoxene GNR IN ○ ○ ○
Diacyclops indianensis Reid, 2004 Indiana Groundwater Copepod Cyclopoida Stygobiont G2 IN, TN ○ ○ ○
Diacyclops jeanneli (Chappuis, 1929) Cyclopoida Stygobiont G1G2 IN, NC ○ 6 ○
Diacyclops lewisi Reid, 2004 Lewis' Groundwater Copepod Cyclopoida Stygobiont G1 IN ○ ○ ○
Diacyclops navus (Herrick, 1882) Cyclopoida Stygoxene GNR IN ○ ○ ○
Diacyclops nearcticus (Kiefer, 1934) Cyclopoida Stygophile G3 IN, KY, TN ○ ○ ○
Diacyclops nikolasarburni Suarez-Morales et al., 2013 Cyclopoida Stygobiont NM ○ 4 ○
Diacyclops salisae Reid, 2004 Salisa's Groundwater Copepod Cyclopoida Stygobiont G1 IN ○ ○ ○
Diacyclops sororum Reid, 1992 Cyclopoida Stygophile GNR IN, KY ○ ○ ○
Diacyclops yeatmani Reid, 1988 Yeatman's Groundwater Copepod Cyclopoida Stygobiont G2G3 IL, IN, MO, OK, TN ○ 5 ○
Diadophis punctatus (Linnaeus, 1766) Ring-Necked Snake Squamata Accidental G5 LC AL, AR, GA, KY ○ ○ ○
Dichogaster affinis (Michaelsen, 1890) Earthworm Crassiclitellata Troglophile ○ ○ ○
Dichogaster bolaui (Michaelsen, 1891) Earthworm Crassiclitellata Troglophile ○ ○ ○
Dicranomyia humidicola Osten Sacken, 1860 Diptera Trogloxene GNR KY ○ ○ ○
Dicranomyia stulta Osten Sacken, 1860 Diptera Trogloxene GNR ○ ○ ○
Dicranota flaveola (Osten Sacken, 1869) Diptera Trogloxene GNR ○ ○ ○
Dicrotendipes neomodestus (Malloch, 1915) Diptera Trogloxene GNR ○ ○ ○
Dictyostelium giganteum B.N.Singh, 1947 Dictyosteliales Stygophile GA ○ ○ ○
Dictyostelium mucoroides Bref., 1870 Dictyosteliales Stygophile GA ○ ○ ○
Dictyostelium purpureum Olive, 1901 Dictyosteliales Stygophile GA ○ ○ ○
Dictyostelium sphaerocephalum (Oudem.) Sacc. & Marchal, 1885 Dictyosteliales Stygophile GA ○ ○ ○
Didelphis virginiana Kerr, 1792 Virginia Opossum Didelphimorphia Trogloxene G5 LC AL, AR, GA, OK ○ ○ ○
Difflugia globulosa Dujardin Arcellinida Stygoxene KY ○ ○ ○
Dineutus assimilis (W. Kirby, 1837) Coleoptera Stygoxene G5 MO ○ ○ ○
Dinocheirus astutus Hoff, 1956 Pseudoscorpiones Troglophile NM ○ ○ ○
Dinocheirus cavicola Muchmore, 1992 Pseudoscorpiones Troglobiont G2G3 TX ○ ○ ○
Diplectrona marianae Reeves in Reeves & Paysen, 1999 Trichoptera Stygoxene G1 GA ○ ○ ○
Diplocardia caroliniana Eisen, 1899 Crassiclitellata Troglophile AL, NC ○ ○ ○
Diplocardia singularis (Ude, 1893) Crassiclitellata Troglophile GNR IL ○ ○ ○
Diplocardia verrucosa Ude, 1895 Crassiclitellata Troglophile MO ○ ○ ○
Discourella dubiosa (Schweizer, 1961) Mesostigmata Trogloxene IL ○ ○ ○
Discus catskillensis (Pilsbry, 1898) Angular Disc Stylommatophora Trogloxene G5 ○ ○ ○
Discus macclintocki (F. C. Baker, 1928) Iowa Pleistocene Snail Stylommatophora Trogloxene G1 LC IA ○ ○ ○
Discus nigrimontanus (Pilsbry, 1924) Black Mountain Disc Stylommatophora Trogloxene G4 TN ○ ○ ○
Discus patulus (Deshayes, 1832) Domed Disc Stylommatophora Trogloxene G5 AL, KY, TN ○ ○ ○
Discus whitneyi (Newcomb, 1864) Forest Disc Stylommatophora Trogloxene ○ ○ ○
Dissochaetus oblitus (LeConte, J. L., 1853) Coleoptera Troglophile GNR TN ○ ○ ○
Docosia paradichroa Fisher, 1937 Diptera Troglophile ○ ○ ○
Dohrniphora perplexa (Brues, 1904) Diptera Trogloxene GNR AL ○ ○ ○
Dolichopeza dorsalis (Johnson, 1909) Diptera Trogloxene GNR ○ ○ ○
Dolichopeza tridenticulata Alexander, 1931 Diptera Trogloxene GNR GA ○ ○ ○
Dolichopeza venosa (Johnson, 1909) Diptera Trogloxene GNR ○ ○ ○
Dolichopeza walleyi (Alexander, 1931) Diptera Trogloxene GNR GA ○ ○ ○
Dolichopus fulvipes Loew, 1862 Diptera Trogloxene GNR ○ ○ ○
Dolichopus penicillatus Van Duzee, 1921 Diptera Trogloxene GNR ○ ○ ○
Dolomedes scriptus Hentz, 1845 Dock Spider Araneae Trogloxene G5 IL, IN, KY, TN ○ ○ ○
Dolomedes tenebrosus Hentz, 1844 Dark Fishing Spider Araneae Trogloxene G5 IL, KY, OK ○ ○ ○
Dolomedes vittatus Walckenaer, 1837 Banded Fishing Spider Araneae Trogloxene GNR AL, IL, TN ○ ○ ○
Donnaldsoncythere hiwasseensis (Hobbs & Walton, 1961) Podocopida Stygophile KY ○ ○ ○
Donnaldsoncythere tuberosa (C. W. Hart & Hobbs, 1961) Podocopida Stygophile G2 TN ○ ○ ○
Dorosoma cepedianum (Lesueur, 1818) American Gizzard Shad Clupeiformes Stygoxene G5 LC AL, KY ○ ○ ○
Dorypteryx domestica (Smithers, 1958) Cave Barklouse Psocodea Trogloxene KY ○ ○ ○
Drosophila funebris (Fabricius, 1787) Azijnvlieg Diptera Trogloxene GNR ○ ○ ○
Dryobius sexnotatus Linsley, 1957 Maple Dryobius Coleoptera Accidental GNR NE KY ○ ○ ○
Dryomyza anilis Fallén, 1820 Hooded Dryomyza Diptera Trogloxene GNR ○ ○ ○
Dryophytes chrysoscelis (Cope, 1880) Cope's Gray Treefrog Anura Accidental G5 LC AL, GA, TN ○ ○ ○
Dryophytes cinereus (Schneider, 1799) Green Treefrog Anura Accidental G5 LC FL, KY ○ ○ ○
Dryophytes gratiosus (LeConte, 1856) Barking Treefrog Anura Accidental G5 LC FL, GA ○ ○ ○
Dryophytes squirellus (Daudin, 1800) Squirrel Treefrog Anura Accidental G5 LC FL ○ ○ ○
Dryophytes versicolor (LeConte, 1825) Gray Treefrog Anura Accidental G5 LC AR ○ ○ ○
Dugesia tigrina (Girard, 1850) Tricladida Stygoxene ○ ○ ○
Duvalius jureceki (Dodero, 1917) Coleoptera Troglophile ○ ○ ○
Duvalius lombardii Straneo, 1939 Coleoptera Troglophile ○ ○ ○
Eclipidrilus palustris (Smith, 1900) Aquatic Oligochaete Worm Lumbriculida Stygoxene TX ○ ○ ○
Eidmannella bullata Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ 1 ○
Eidmannella delicata Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ ○ ○
Eidmannella nasuta Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ ○ ○
Eidmannella pallida (Emerton, 1875) Pallid Cave Spider Araneae Troglophile GNR AL, AR, AZ, FL, GA, HI, IA, IL, IN, KY, MD, NM, OH, OK, PA, SC, TN, TX, VA, WV ○ ○ ○
Eidmannella reclusa Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ 13 ○
Eidmannella rostrata Gertsch, 1984 Araneae Troglobiont G4 TX ○ 262 ○
Eidmannella tuckeri Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 1 ○
Eisenia fetida (Savigny in Cuvier, 1826) Brandling Crassiclitellata Troglophile GNR KY ○ ○ ○
Eisenia rosea (Savigny in Cuvier, 1826) Crassiclitellata Troglophile IL ○ ○ ○
Eiseniella tetraedra (Savigny in Cuvier, 1826) Square-Tailed Worm Crassiclitellata Troglophile IA, VA, WV ○ ○ ○
Eisenoides carolinensis (Michaelsen, 1910) Carolina Worm Crassiclitellata Troglophile GNR IL ○ ○ ○
Eisenoides lonnbergi (Michaelsen, 1894) Crassiclitellata Troglophile NC ○ ○ ○
Elaphoidella bidens (Schmeil, 1893) Harpacticoida Stygoxene GA ○ ○ ○
Elaphropus ferrugineus (Dejean, 1831) Coleoptera Troglophile AL, GA ○ ○ ○
Elaphropus incurvus (Say, 1830) Coleoptera Trogloxene G5 IL ○ ○ ○
Elaver excepta (L. Koch, 1866) White Sac Spider Araneae Trogloxene G5 GA ○ ○ ○
Eleodes eschscholtzii Solier, 1848 Pinacate De Eschscholtz Coleoptera Trogloxene NM ○ ○ ○
Eleodes microps Aalbu et al., 2012 Coleoptera Troglobiont CA ○ 12 5
Eleutherodactylus planirostris (Cope, 1862) Greenhouse Frog Anura Trogloxene G5 LC FL ○ ○ ○
Elimia athearni (Clench & R. D. Turner, 1956) Knobby Elimia Cerithiida Accidental G3 LC FL ○ ○ ○
Elimia clenchi (Goodrich, 1924) Slackwater Elimia Cerithiida Stygoxene G3 LC FL ○ ○ ○
Elimia comalensis (Pilsbry, 1890) Balcones Elimia Cerithiida Stygophile G2 TX ○ ○ ○
Elimia curvicostata (Reeve, 1861) Cerithiida Stygoxene FL ○ ○ ○
Elimia proxima (Say, 1825) Sprite Elimia Cerithiida Stygoxene G5 LC GA ○ ○ ○
Elliotta howarthi (Elliott, 1976) Actinedida Troglobiont G1G2 WA ○ ○ ○
Emblyna florens (Ivie & Barrows, 1935) Araneae Accidental AL ○ ○ ○
Empherozoster antaeus Crabill, 1959 Geophilomorpha Troglophile NM ○ ○ ○
Enoplognatha ovata (Clerck, 1757) Common Candy-Striped Spider Araneae Accidental GNR ○ ○ ○
Enterobryus oxidi Lichtw., 1961 Eccrinales Stygoxene GA ○ ○ ○
Entomobrya bicolor Guthrie, JE, 1903 Entomobryomorpha Accidental GNR IA ○ ○ ○
Entomobrya confusa Christiansen, K, 1958 Entomobryomorpha Accidental GNR IA ○ ○ ○
Entomobrya intermedia Brook, G, 1884 Entomobryomorpha Trogloxene GNR IL ○ ○ ○
Entomobrya nivalis Christiansen, K, 1958 Entomobryomorpha Trogloxene ○ ○ ○
Entomobrya troglodytes Christiansen, 1958 Entomobryomorpha Troglobiont G2G3 CO, SD ○ ○ ○
Entomobryoides purpurascens (Packard, AS, 1872) Springtail Entomobryomorpha Troglophile GNR NM ○ ○ ○
Ephemera simulans Walker, 1853 Brown Drake Ephemeroptera Stygoxene G4 ○ ○ ○
Ephippiochthonius tetrachelatus (Preyssler, 1790) Dimple-Clawed Chthonid Pseudoscorpiones Troglophile AL, IL ○ ○ ○
Ephippiochthonius virginicus (J. C. Chamberlin, 1929) Pseudoscorpiones Troglophile GA ○ ○ ○
Epicharis rustica (Olivier, 1789) Hymenoptera Accidental ○ ○ ○
Epitedia cavernicola Traub, 1957 Siphonaptera Trogloxene VA ○ ○ ○
Epitedia wenmanni (Rothschild, 1904) Deermice Flea Siphonaptera Trogloxene GNR KY, TN ○ ○ ○
Eptesicus fuscus (Palisot de Beauvois, 1796) Big Brown Bat Chiroptera Trogloxene G5 LC AL, AR, GA, IL, IN, KY, OK, PA, TN, VA ○ ○ ○
Equus caballus Linnaeus, 1758 Domestic Horse Perissodactyla Accidental GNA FL ○ ○ ○
Erebomaster flavescens Cope, 1872 Golden Cave Harvestman Opiliones Troglophile G3 IN, KY ○ ○ ○
Ereboporus naturaconservatus Miller et al., 2019 Nature Conservancy Diving Beetle Coleoptera Stygobiont GNR TX ○ 1 8
Eremidrilus allegheniensis (Cook, 1971) Lumbriculida Stygobiont G1G2 TN ○ 3 ○
Eremidrilus coyote Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 19 8
Eremidrilus elegans Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 20 ○
Eremidrilus felini Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 28 ○
Eremidrilus parvitheca Fend, 2023 Lumbriculida Stygobiont TX ○ 6 ○
Eremidrilus ritocsi Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 10 ○
Eremidrilus samacos Fend, 2023 Lumbriculida Stygobiont TX ○ 8 ○
Erethizon dorsatum (Linnaeus, 1758) Rodentia Trogloxene KY ○ ○ ○
Ergodesmus remingtoni (Hoffman, 1962) Polydesmida Troglobiont G3 IL ○ 1 ○
Erichsonius nanus (Horn, 1884) Coleoptera Trogloxene G5 IL ○ ○ ○
Erichsonius patella (Horn, 1884) Kneecap Rove Beetle Coleoptera Trogloxene GNR AL ○ ○ ○
Erigone autumnalis Emerton, 1882 Autumn Money Spider Araneae Troglophile GNR IA, IL ○ ○ ○
Erigone infernalis Keyserling, 1886 Araneae Troglophile GNR IN, KY ○ ○ ○
Erigone stygius Gertsch, 1973 Araneae Troglobiont G1G2 HI ○ ○ ○
Erpobdella dubia (Moore & Meyer, 1951) Arhynchobdellida Stygoxene ○ ○ ○
Etheostoma artesiae (Hay, 1881) Redspot Darter Perciformes Accidental G5 LC AL ○ ○ ○
Etheostoma caeruleum Storer, 1845 Rainbow Darter Perciformes Accidental G5 LC AL, KY ○ ○ ○
Etheostoma duryi Henshall, 1889 Black Darter Perciformes Accidental G4 LC AL, TN ○ ○ ○
Etheostoma flabellare Rafinesque, 1819 Fantail Darter Perciformes Accidental G5 LC AL ○ ○ ○
Etheostoma swaini (Jordan, 1884) Gulf Darter Perciformes Accidental G5 LC FL ○ ○ ○
Ethopolys bipunctatus Wood (1862) Lithobiomorpha Trogloxene UT ○ ○ ○
Euborellia annulipes (Lucas, 1847) Ring-Legged Earwig Dermaptera Trogloxene GNR AL ○ ○ ○
Euborellia antoni Hebard, 1917 Dermaptera Trogloxene ○ ○ ○
Eucyclops agilis (Koch, 1838) Cyclopoida Stygophile GNR IL, IN, KY, NM, TN ○ ○ ○
Eucyclops bondi Kiefer, 1934 Cyclopoida Stygophile GA, IN ○ ○ ○
Eucyclops elegans (Herrick, 1884) Cyclopoida Stygophile GA, IN, KY, TN ○ ○ ○
Eucyclops speratus (Lilljeborg, 1901) Fintagghalehops Cyclopoida Stygoxene KY ○ ○ ○
Euglandina rosea (A. Férussac, 1821) Rosy Predator Snail Stylommatophora Trogloxene G5 FL ○ ○ ○
Euglesa casertana (Poli, 1791) Caserta Pea Mussel Sphaeriida Accidental LC ○ ○ ○
Euhadenoecus fragilis Hubbell, 1978 Tawneys Cave Cricket Orthoptera Troglophile KY, TN, VA ○ ○ ○
Euhadenoecus insolitus Hubbell, 1978 Highland Rim Crevice Cricket Orthoptera Troglophile AL, TN ○ ○ ○
Euhadenoecus puteanus (Scudder, 1877) Allegheny Crevice Cricket Orthoptera Troglophile GNR AL, GA, KY, TN ○ ○ ○
Eupeodes volucris Osten Sacken, 1877 Large-Tailed Aphideater Diptera Accidental G4 ○ ○ ○
Eurycea bislineata (Green, 1818) Caudata Trogloxene G5 LC OH ○ ○ ○
Eurycea braggi (Smith, 1968) Southern Grotto Salamander Caudata Troglobiont GNR LC AR, MO ○ 15 137
Eurycea chisholmensis Chippindale, Price, Wiens & Hillis, 2000 Salado Salamander Caudata Stygophile G1 EN TX ○ ○ ○
Eurycea cirrigera (Green, 1831) Southern Two-Lined Salamander Caudata Trogloxene G5 LC AL, FL, GA, KY, TN ○ ○ ○
Eurycea guttolineata (Holbrook, 1838) Three-Lined Salamander Caudata Trogloxene G5 LC AL, FL, GA ○ ○ ○
Eurycea longicauda (Green, 1818) Long-Tailed Salamander Caudata Troglophile G5 LC AL, GA, IL, KY, MO, TN ○ ○ ○
Eurycea lucifuga Rafinesque, 1822 Cave Salamander Caudata Troglophile G5 LC AL, AR, GA, IL, IN, KY, MO, OK, TN, VA, WV ○ ○ ○
Eurycea melanopleura (Cope, 1894) Dark-Sided Salamander Caudata Troglophile AR, OK ○ ○ ○
Eurycea multiplicata (Cope, 1869) Many-Ribbed Salamander Caudata Trogloxene G4 LC AR ○ ○ ○
Eurycea nerea (Bishop, 1944) Northern Grotto Salamander Caudata Troglobiont GNR LC AR, MO, KS ○ 67 144
Eurycea pterophila Burger, Smith & Potter, 1950 Fern Bank Salamander Caudata Stygophile G3 EN TX ○ ○ ○
Eurycea rathbuni (Stejneger, 1896) Texas Blind Salamander Caudata Stygobiont G1 CR ESA TX ○ 380 9
Eurycea robusta (Longley, 1978) Blanco Blind Salamander Caudata Stygobiont G1 DD TX ○ 1 ○
Eurycea spelaea (Stejneger, 1892) Western Grotto Salamander Caudata Troglobiont G4 LC AR, KS, MO, OK, MT ○ 2.1k 236
Eurycea tonkawae Chippindale et al., 2000 Jollyville Plateau Salamander Caudata Stygobiont G2 EN ESA TX ○ 232 22
Eurycea tridentifera Mitchell & Reddell, 1965 Comal Blind Salamander Caudata Stygobiont G3 VU TX ○ ○ ○
Eurycea wallacei (Carr, 1939) Georgia Blind Salamander Caudata Stygobiont G1G2 VU FL, GA ○ 175 22
Eurycea waterlooensis Hillis et al., 2001 Austin Blind Salamander Caudata Stygobiont G1 CR ESA TX ○ 14 5
Eurycea wilderae Dunn, 1920 Blue Ridge Two-Lined Salamander Caudata Trogloxene G5 LC TN ○ ○ ○
Euryurus erythropygos (Brandt, 1839) Polydesmida Trogloxene G3 IN ○ ○ ○
Euryurus evides (Bollman, 1887) Polydesmida Trogloxene AR ○ ○ ○
Euryurus leachii (Gray, 1832) Log Lurker Polydesmida Trogloxene G4 AL, KY ○ ○ ○
Euschoengastia pipistrelli Brennan, 1947 Trombidiformes Trogloxene AL, GA, TN ○ ○ ○
Euschoengastia staffordi Brennan & White, 1960 Trombidiformes Trogloxene AL ○ ○ ○
Eustala anastera (Walckenaer, 1841) Orb Weaver Araneae Accidental G5 ○ ○ ○
Eustilicus condei (Jarrige, 1960) Coleoptera Troglophile NM, TX ○ ○ ○
Eutrichota affinis (Stein, 1898) Diptera Trogloxene GNR IN ○ ○ ○
Eutrichota lipsia (Walker, 1849) Diptera Trogloxene GNR IN ○ ○ ○
Eutrombicula alfreddugesi (Oudemans, 1910) Trombidiformes Trogloxene AL ○ ○ ○
Exechia attrita Johannsen, 1912 Diptera Troglophile ○ ○ ○
Exechia auxiliaria Johannsen, 1912 Diptera Troglophile ○ ○ ○
Exechia bifurcata Fisher, 1934 Diptera Troglophile ○ ○ ○
Exechia obediens Johannsen, 1912 Diptera Troglophile GNR ○ ○ ○
Exechia shawi Fisher, 1934 Diptera Troglophile TN ○ ○ ○
Exechia umbratica (Aldrich, 1897) Diptera Troglophile GNR IN ○ ○ ○
Exechiopsis nugax (Johannsen, 1912) Diptera Troglophile ○ ○ ○
Faiditus cancellatus (Hentz, 1850) Araneae Accidental GNR AL ○ ○ ○
Faxonius neglectus (Faxon, 1885) Ringed Crayfish Decapoda Stygophile G5 MO, OK ○ ○ ○
Faxonius rusticus (Girard, 1852) Rusty Crayfish Decapoda Stygoxene G5 MO ○ ○ ○
Faxonius stygocaneyi (Hobbs, 2001) Caney Mountain Cave Crayfish Decapoda Stygobiont G1 LC MO ○ 2 5
Faxonius validus (Faxon, 1914) Powerful Crayfish Decapoda Stygoxene G5 AL ○ ○ ○
Faxonius virilis (Hagen, 1870) Virile Crayfish Decapoda Stygoxene G5 MO ○ ○ ○
Felis rufus Schreber, 1777 Carnivora Trogloxene AR ○ ○ ○
Feralia jocosa Guenée, 1852 Jocose Sallow Lepidoptera Trogloxene G5 ○ ○ ○
Fissilicreagris imperialis (Muchmore, 1969) Empire Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VU CA ○ ○ ○
Flabellorhagidia pecki Elliott, 1976 Actinedida Troglobiont G1G2 ID ○ ○ ○
Folsomia bisetosa Gisin, 1953 Entomobryomorpha Trogloxene GNR IL ○ ○ ○
Folsomia candida Willem, V, 1902 Springtail Entomobryomorpha Troglophile GNR AL, AR, GA, IA, IL, KY, MN, MO, NM, TN ○ ○ ○
Folsomia elongata (MacGillivray, 1896) Entomobryomorpha Trogloxene GNR IL ○ ○ ○
Folsomia fimetaria (Linnæus, C, 1758) Springtail Entomobryomorpha Troglophile GNR KY ○ ○ ○
Folsomia nivalis (Packard, AS, 1873) Entomobryomorpha Trogloxene GNR IA, IL, WI ○ ○ ○
Folsomia penicula Bagnall, 1939 Entomobryomorpha Troglophile GNR AL ○ ○ ○
Folsomia prima Mills, HB, 1931 Entomobryomorpha Trogloxene GNR IA, IL ○ ○ ○
Folsomia stella Christiansen, KA & Tucker, BE, 1977 Springtail Entomobryomorpha Troglophile GNR IA, IL, MN, TN, WI ○ ○ ○
Folsomia variabilis Fjellberg, A, 1984 Entomobryomorpha Trogloxene IL ○ ○ ○
Folsomides americanus Denis, 1931 Entomobryomorpha Accidental IA ○ ○ ○
Fontigens aldrichi (Call & Beecher, 1886) Hoosier Springsnail Littorinimorpha Stygophile G4 MO ○ ○ ○
Fontigens antroecetes Hubricht, 1972 Enigmatic ghostsnail Littorinimorpha Stygobiont G2 VU IL, MO ○ 35 1
Fontigens bottimeri (Walker, 1925) Appalachian Springsnail Littorinimorpha Stygobiont G2G3 DC, MD, VA, MO ○ 68 2
Fontigens cryptica Hubricht, 1963 Hidden springsnail Littorinimorpha Stygobiont G1 IN, KY ○ 7 1
Fontigens morrisoni Hershler, Holsinger & Hubricht, 1990 Virginia springsnail Littorinimorpha Stygobiont G2 VA ○ 8 ○
Fontigens orolibas Hubricht, 1957 Blue Ridge Springsnail Littorinimorpha Stygophile G3 DD VA ○ ○ ○
Fontigens proserpina Hubricht, 1940 Proserpine Springsnail Littorinimorpha Stygobiont G1 MO ○ 20 ○
Fontigens tartarea Hubtricht 1963 Organ cavesnail Littorinimorpha Stygobiont G2 VA, WV, MD ○ 42 2
Fontigens turritella Hubricht, 1976 Greenbrier Cavesnail Littorinimorpha Stygobiont G1 NT WV ○ 8 ○
Forbesichthys agassizii (Putnam, 1872) Spring Cavefish Percopsiformes Stygophile GNR LC TN ○ ○ ○
Forbesichthys papilliferus (Forbes, 1882) Percopsiformes Stygophile G3 IL, KY, TN ○ ○ ○
Forcipomyia pholetor Wirth & Howarth, 1982 Diptera Troglobiont HI ○ ○ ○
Foveacheles auricularia Zacharda & Elliott, 1985 Actinedida Troglobiont CA ○ ○ ○
Foveacheles goffi Zacharda, 1982 Actinedida Troglobiont G1G2 HI ○ 1 ○
Foveacheles tenorioae Zacharda, 1982 Actinedida Troglobiont G1G2 HI ○ ○ ○
Foveacheles titanica Zacharda & Elliott, 1985 Actinedida Troglobiont G2G3 CA ○ ○ ○
Foveacheles tuktoyaktukensis Zacharda, 1980 Actinedida Troglophile WV ○ ○ ○
Friesea claviseta Axelson, 1900 Bark-Tretornspringhale Poduromorpha Trogloxene GNR IA ○ ○ ○
Fumonelix wetherbyi (Bland, 1874) Clifty Covert Stylommatophora Trogloxene G2 TN ○ ○ ○
Fundulus notatus (Rafinesque, 1820) Blackstripe Topminnow Cyprinodontiformes Accidental G5 LC AL ○ ○ ○
Gabrius microphthalmus (Horn, 1884) Coleoptera Troglophile G5 IL ○ ○ ○
Gaeolaelaps queenslandicus (Womersley) Mesostigmata Troglophile IL ○ ○ ○
Galba modicella (Say, 1825) Rock Fossaria Lymnaeida Stygoxene ○ ○ ○
Galerita bicolor (Drury, 1773) Coleoptera Accidental GNR MO ○ ○ ○
Galerita lecontei Dejean, 1831 Coleoptera Accidental AL ○ ○ ○
Gallus gallus (Linnaeus, 1758) Red Junglefowl Galliformes Accidental G5 LC KY ○ ○ ○
Galumna alata (Hermann, 1804) Mite Oribatida Troglobiont G1G2 KY ○ ○ ○
Gambusia affinis (Baird & Girard, 1853) Western Mosquitofish Cyprinodontiformes Accidental G5 LC AL, FL ○ ○ ○
Gambusia holbrooki Girard, 1859 Eastern Mosquitofish Cyprinodontiformes Accidental G5 LC FL ○ ○ ○
Gammarus acherondytes Hubricht & Mackin, 1940 Illinois Cave Amphipod Amphipoda Stygobiont G2G3 EN ESA IL ○ 37 80
Gammarus cohabitus Holsinger & Shafer, 2008 Amphipoda Stygobiont G1G2 PA ○ 10 ○
Gammarus limnaeus S.I. Smith, 1874 Amphipoda Stygophile GNR IL, MO ○ ○ ○
Gammarus minus Say, 1818 Amphipoda Stygophile GNR VA, WV, AL, AR, GA, IL, IN, KY, MD, MO, OH, OK, PA, TN ○ ○ ○
Gammarus minus tenuipes Shoemaker, 1940 Amphipoda Stygobiont VA, WV ○ ○ ○
Gammarus pseudolimnaeus Bousfield, 1958 Northern Spring Amphipod Amphipoda Stygophile G5 IA, IL ○ ○ ○
Gammarus troglophilus Hubricht & Mackins, 1940 Amphipoda Stygophile GNR IL, MO ○ ○ ○
Gastrellarius honestus (Say, 1823) Coleoptera Accidental G5 KY ○ ○ ○
Gastrocopta abbreviata (Sterki, 1909) Plains Snaggletooth Stylommatophora Trogloxene G4 IL ○ ○ ○
Gastrocopta armifera (Say, 1821) Armed Snaggletooth Stylommatophora Trogloxene G5 IL, KY ○ ○ ○
Gastrocopta contracta (Say, 1822) Bottleneck Snaggletooth Stylommatophora Trogloxene G5 IL, KY ○ ○ ○
Gastrocopta pentodon (Say, 1822) Comb Snaggletooth Stylommatophora Trogloxene IL ○ ○ ○
Gastrocopta similis (Sterki, 1909) Great Lakes Snaggletooth Stylommatophora Trogloxene G5 IL ○ ○ ○
Gastrodonta interna (Say, 1821) Brown Bellytooth Stylommatophora Trogloxene G5 AL, GA, TN ○ ○ ○
Gastrophryne carolinensis (Holbrook, 1835) Eastern Narrow-Mouthed Toad Anura Accidental G5 LC ○ ○ ○
Gastrophryne olivacea (Hallowell, 1856) Great Plains Narrow-Mouthed Toad Anura Accidental G5 LC TX ○ ○ ○
Gaucelmus augustinus Keyserling, 1884 Araneae Troglophile G3 AL, FL, GA, MS, TX ○ ○ ○
Gaucelmus cavernicola (Petrunkevitch, 1910) Araneae Troglophile ○ ○ ○
Gaucelmus strinatii Brignoli, 1979 Araneae Troglophile ○ ○ ○
Gaucelmus tropicus Gertsch, 1984 Araneae Troglophile ○ ○ ○
Gennadota puberula (Casey, 1893) Coleoptera Trogloxene IA ○ ○ ○
Geocentrophora cavernicola Carpenter, 1970 Alloecoela Stygobiont G2 KY, VA, WV ○ 2 ○
Geocolus caecus Wenzel, 1944 Coleoptera Trogloxene AL ○ ○ ○
Geodromicus brunneus (Say, 1823) Coleoptera Trogloxene GNR AL, GA, IL, TN ○ ○ ○
Geomys bursarius (Shaw, 1800) Plains Pocket Gopher Rodentia Accidental G5 LC KY ○ ○ ○
Girardia dorotocephala (Woodworth, 1897) Flatworm Tricladida Stygophile NV ○ ○ ○
Glacicavicola bathyscioides Westcott, 1968 Coleoptera Troglobiont G3 VU ID, WY ○ 4 ○
Globocreagris nigrescens (J. C. Chamberlin, 1952) Pseudoscorpiones Troglophile VA ○ ○ ○
Glyphyalinia carolinensis (T. D. A. Cockerell, 1890) Spiral Mountain Glyph Stylommatophora Trogloxene G4 TN ○ ○ ○
Glyphyalinia cryptomphala (G. H. Clapp, 1915) Thin Glyph Stylommatophora Troglophile G5 AL, GA, KY, TN ○ ○ ○
Glyphyalinia indentata (Say, 1822) Carved Glyph Stylommatophora Troglophile G5 AL, GA, KY, TN, VA ○ ○ ○
Glyphyalinia latebricola Hubricht, 1968 Stone Glyph Stylommatophora Troglophile G2 AL, IL ○ ○ ○
Glyphyalinia lewisiana (G. H. Clapp, 1908) Pale Glyph Stylommatophora Trogloxene G4 KY, TN ○ ○ ○
Glyphyalinia luticola Hubricht, 1966 Furrowed Glyph Stylommatophora Trogloxene G4 IL ○ ○ ○
Glyphyalinia pecki Hubricht, 1966 Blind Glyph Stylommatophora Troglobiont G1G2 DD AL ○ 5 ○
Glyphyalinia praecox (H. B. Baker, 1930) Brilliant Glyph Stylommatophora Trogloxene G4 GA, TN ○ ○ ○
Glyphyalinia rhoadsi (Pilsbry, 1899) Sculpted Glyph Stylommatophora Troglophile G5 GA, TN ○ ○ ○
Glyphyalinia sculptilis (Bland, 1858) Suborb Glyph Stylommatophora Troglophile G4 AL, GA, KY, TN ○ ○ ○
Glyphyalinia specus Hubricht, 1965 Hollow glyph Stylommatophora Troglobiont G4 AL, GA, KY, TN, WV ○ 28 ○
Glyphyalinia wheatleyi (Bland, 1883) Bright Glyph Stylommatophora Troglophile G5 GA, TN ○ ○ ○
Gnathonaroides pedalis (Emerton, 1923) Araneae Trogloxene GNR ○ ○ ○
Gnophomyia tristissima Osten Sacken, 1860 Diptera Trogloxene GNR IL ○ ○ ○
Gopherus polyphemus (Daudin, 1801) Gopher Tortoise Testudines Accidental G3 VU ○ ○ ○
Gordionus lineatus Leidy, 1851 Gordioidea Trogloxene TN ○ ○ ○
Grandidierella palama Barnard, 1977 Amphipoda Stygobiont HI ○ 4 ○
Gymnophora luteiventris Schmitz, 1952 Diptera Trogloxene GNR ○ ○ ○
Gyraulus parvus (Say, 1817) Ash Gyro Lymnaeida Stygoxene FL ○ ○ ○
Gyrinophilus gulolineatus Brandon, 1965 Berry cave salamander Caudata Stygobiont G1 EN TN ○ 31 34
Gyrinophilus palleucus McCrady, 1954 Tennessee Cave Salamander Caudata Stygobiont G2G3 NT AL, GA, TN ○ 206 67
Gyrinophilus porphyriticus (Green, 1827) Spring Salamander Caudata Troglophile G5 LC AL, GA, KY, OH, TN, VA, WV ○ ○ ○
Gyrinophilus subterraneus Besharse & Holsinger, 1977 West Virginia Spring Salamander Caudata Stygobiont G1 CR ESA? WV ○ 30 45
Hadenoecus barri Hubbell, 1978 Barr's Cave Cricket Orthoptera Troglophile TN ○ ○ ○
Hadenoecus cumberlandicus Hubbell & Norton, 1978 Cumberland Cave Cricket Orthoptera Troglophile GNR KY ○ ○ ○
Hadenoecus jonesi Hubbell, 1978 Limrock Blowing Cave Cricket Orthoptera Troglophile GNR AL, TN ○ ○ ○
Hadenoecus opilionoides Hubbell, 1978 Orthoptera Troglophile TN ○ ○ ○
Hadenoecus subterraneus (Scudder, 1861) Mammoth Cave Cricket Orthoptera Troglophile GNR KY, TN ○ ○ ○
Haideoporus texanus Young & Longley, 1976 Edwards Aquifer Diving Beetle Coleoptera Stygobiont G1G2 NE TX ○ 43 8
Halocaridina palahemo Kensley & Williams, 1986 Decapoda Stygobiont G1 HI ○ 3 ○
Halocaridina rubra Holthuis, 1963 Decapoda Stygobiont G3 HI ○ 25 337
Hamohalacarus subterraneus Walter, 1931 Donaldson's Cave Water Mite Actinedida Stygobiont G1 IN ○ ○ ○
Haplophthalmus danicus Budde-Lund in Meinert, 1880 Spurred Ridgeback Isopoda Troglophile IL, IN, KY, NM, TN, VA ○ ○ ○
Haplotaxis brinkhursti Cook, 1975 Haplotaxida Stygobiont G1 WV ○ ○ ○
Haplotaxis gordioides (Hartmann in Oken, 1819) Haplotaxid Worm Haplotaxida Stygoxene GNR WV ○ ○ ○
Haplotrema concavum (Say, 1821) Gray-Foot Lancetooth Stylommatophora Trogloxene G5 AL, FL, TN ○ ○ ○
Harpalus pennsylvanicus Say, 1823 Coleoptera Trogloxene GA ○ ○ ○
Hawaiia minuscula (A. Binney, 1841) Minute Gem Stylommatophora Troglophile G5 GA, IL, MO, TN ○ ○ ○
Hawaiioscia micropthalma Taiti & Howarth, 1997 Isopoda Troglobiont G1G2 HI ○ ○ ○
Hawaiioscia paeninsulae Taiti & Howarth, 1997 Isopoda Troglobiont G1G2 HI ○ ○ 1
Hawaiioscia parvituberculata Schultz, 1973 Isopoda Troglobiont G1G2 HI ○ 7 ○
Hawaiioscia rotundata Taiti & Howarth, 1997 Isopoda Troglobiont G1G2 HI ○ 1 3
Hawinella lava Bellinger & Christiansen, 1974 Entomobryomorpha Troglobiont G2G3 HI ○ 1 ○
Heleomyza brachypterna (Loew, 1873) Diptera Trogloxene AR ○ ○ ○
Heleomyza serrata (Linnaeus, 1758) Diptera Trogloxene GNR IN, WI ○ ○ ○
Helicina orbiculata (Say, 1818) Cycloneritida Trogloxene AL, TN ○ ○ ○
Helicodiscus barri Hubricht, 1962 Raccoon coil Stylommatophora Troglobiont G3G4 AL, GA, TN ○ 13 106
Helicodiscus fimbriatus Wetherby, 1881 Fringed Coil Stylommatophora Trogloxene G4 AL ○ ○ ○
Helicodiscus hadenoecus Hubricht, 1962 Cricket Coil Stylommatophora Troglophile AL, KY, TN ○ ○ ○
Helicodiscus inermis H. B. Baker, 1929 Stylommatophora Troglophile AL ○ ○ ○
Helicodiscus multidens Hubricht, 1962 Twilight Coil Stylommatophora Troglophile G3 TN ○ ○ ○
Helicodiscus notius Hubricht, 1962 Tight Coil Stylommatophora Troglophile G5 AL, GA, IL, KY, MO, TN ○ ○ ○
Helicodiscus notius specus Hubricht, 1962 Hollow Coil Stylommatophora Troglobiont G2 KY, TN, AL, GA, IL, MO ○ ○ ○
Helicodiscus parallelus (Say, 1821) Compound Coil Stylommatophora Troglophile G5 GA, TN ○ ○ ○
Helicodiscus punctatellus J. P. E. Morrison, 1942 Stylommatophora Troglophile KY ○ ○ ○
Helicodiscus shimeki Hubricht, 1962 Temperate Coil Stylommatophora Trogloxene G5 ○ ○ ○
Helius flavipes (Macquart, 1855) Diptera Trogloxene GNR ○ ○ ○
Helodrilus foetidus Michaelsen, 1913 Crassiclitellata Troglophile KY ○ ○ ○
Helodrilus tenuis (Eisen, 1874) Crassiclitellata Troglophile KY ○ ○ ○
Hemidactylium scutatum (Temminck in Temminck & Schlegel, 1838) Four-Toed Salamander Caudata Accidental G5 LC OH ○ ○ ○
Heminothrus targionii (Berlese, 1885) Sarcoptiformes Trogloxene GNR ○ ○ ○
Hemiscolopendra marginata Say (1821) Eastern Bark Centipede Scolopendromorpha Trogloxene GNR IN ○ ○ ○
Hemisotoma thermophila (Axelson, 1900) Entomobryomorpha Trogloxene GNR IL ○ ○ ○
Hemitremia flammea (Jordan & Gilbert, 1878) Flame Chub Cypriniformes Accidental G3 NT AL ○ ○ ○
Hentziectypus globosus (Hentz, 1850) Fingered Cobweaver Araneae Accidental GNR AL ○ ○ ○
Hesperochernes bradybaughi Harvey & Wynne, 2014 Pseudoscorpiones Troglobiont GNR AZ ○ ○ ○
Hesperochernes holsingeri Muchmore, 1994 Holsinger's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 IN ○ ○ ○
Hesperochernes mirabilis (Banks, 1895) Southeastern Cave Pseudoscorpion Pseudoscorpiones Troglobiont G5 AL, GA, IN, KY, OH, TN, VA, IL ○ 2 ○
Hesperochernes occidentalis (Hoff & Bolsterli, 1956) Guano Pseudoscorpion Pseudoscorpiones Troglobiont G5 AR, MO, OK, TX ○ 9 ○
Hesperonemastoma inops (Packard, 1884) Opiliones Troglobiont G1G2 KY ○ ○ ○
Hesperonemastoma pallidimaculosum (Goodnight & Goodnight, 1945) Opiliones Troglobiont G3 AL ○ ○ ○
Hesperonemastoma smilax Shear, 2010 Opiliones Troglobiont GNR WV ○ ○ ○
Heteraphorura subtenuis (Folsom, 1917) Poduromorpha Troglophile GNR IL ○ ○ ○
Heterelmis comalensis Bosse, Tuff & Brown, 1988 Comal Springs Riffle Beetle Coleoptera Stygophile G1 TX ○ ○ ○
Heteromurus nitidus (Templeton, R in Templeton, R & Westwood, JO, 1836) Entomobryomorpha Troglophile IA ○ ○ ○
Heterophrys glabrescens Penard, 1904 Piroplasmida Stygoxene AL, KY ○ ○ ○
Hexabathynella hessleri Cho, 2001 Bathynellacea Stygobiont CA ○ ○ ○
Hexabathynella muliebris Cho, 2001 Bathynellacea Stygobiont CA ○ ○ ○
Hexabathynella otayana Cho, 2001 Bathynellacea Stygobiont CA ○ 2 ○
Hexabathynella virginiae Cho & Schminke, 2006 Bathynellacea Stygobiont VA ○ ○ ○
Hibana gracilis (Hentz, 1847) Garden Ghost Spider Araneae Accidental GNR AL ○ ○ ○
Hickmania troglodytes (Higgins & Petterd, 1883) Tasmanian Cave Spider Araneae Troglophile ○ ○ ○
Hirundo rustica Linnaeus, 1758 Barn Swallow Passeriformes Trogloxene G5 LC AR, TN ○ ○ ○
Hobbsinella edwardensis Camacho et al., 2017 Bathynellacea Stygobiont GNR TX ○ 5 1
Hobbsinella gunnisonensis Camacho & Taylor, 2023 Bathynellacea Stygobiont CO ○ ○ ○
Hogna frondicola (Emerton, 1885) Forest Wolf Spider Araneae Trogloxene G5 ○ ○ ○
Holsingeria unthanksensis Hershler, 1989 Thankless ghostsnail Littorinimorpha Stygobiont G2 VA, WV ○ 20 1
Holsingerius samacos (Holsinger, 1980) Amphipoda Stygobiont G1G2 TX ○ 78 ○
Homaeotarsus bicolor (Gravenhorst, 1802) Coleoptera Trogloxene GNR IL ○ ○ ○
Homaeotarsus carolinus (Erichson, 1840) Coleoptera Trogloxene GNR AL ○ ○ ○
Homidia socia Denis, 1929 Three-Lined Springtail Entomobryomorpha Trogloxene IL ○ ○ ○
Horologion hubbardi Harden & Davidson, 2024 Coleoptera Troglobiont GNR VA ○ ○ 10
Horologion speokites Valentine, 1932 Arbuckle Cave Beetle Coleoptera Troglobiont GH WV ○ ○ ○
Horreolanus orphanus Mitchell, 1955 Trombidiformes Stygobiont IL, OH ○ 42 5
Howaia mogera (Yaginuma, 1972) Kasholenspin Araneae Troglophile HI ○ ○ ○
Hubbardia shoshonensis Briggs & Hom, 1972 Shoshone Cave Whipscorpion Schizomida Troglobiont G1 CA ○ 1 ○
Hyalella azteca (Saussure, 1858) Mexican Freshwater Shrimp Amphipoda Stygoxene G5 FL ○ ○ ○
Hyalella cretae Cannizzaro et al., 2023 Amphipoda Stygobiont GNR NV ○ ○ ○
Hygrotus sayi J. Balfour-Browne, 1944 Say's Diving Beetle Coleoptera Stygoxene G5 ○ ○ ○
Hylarana grandocula (Taylor, 1920) Big-Eyed Frog Anura Trogloxene ○ ○ ○
Hypena scabra (Fabricius, 1798) Green Cloverworm Lepidoptera Trogloxene G5 IL, IN, VA ○ ○ ○
Hypentelium nigricans (Lesueur, 1817) Northern Hog Sucker Cypriniformes Stygoxene G5 LC AL ○ ○ ○
Hypoaspis angusta Karg, 1962 Mesostigmata Troglophile IL ○ ○ ○
Hypochilus bonneti Gertsch, 1964 Araneae Trogloxene GNR CO, NM ○ ○ ○
Hypochilus coylei Platnick, 1987 Araneae Trogloxene G3 NC ○ ○ ○
Hypochilus gertschi Hoffman, 1963 Araneae Trogloxene G3 VA ○ ○ ○
Hypochilus pococki Platnick, 1987 Pocock's Lampshade-Web Spider Araneae Trogloxene G4 NC, TN ○ ○ ○
Hypochilus thorelli Marx, 1888 Thorell's Lampshade-Web Spider Araneae Trogloxene G4 AL, GA, TN, VA ○ ○ ○
Hypogastrura antra Christiansen, K & Bellinger, P, 1980 Poduromorpha Troglophile KY, TN ○ ○ ○
Hypogastrura assimilis (Krausbauer, 1898) Mark-Tornspringhale Poduromorpha Trogloxene GNR AL, IL, TN ○ ○ ○
Hypogastrura matura (Folsom, JW, 1916) Poduromorpha Trogloxene GNR IL, MO ○ ○ ○
Hypogastrura notha (Macnamara, 1922) Poduromorpha Trogloxene IL ○ ○ ○
Hypogastrura purpurescens (Lubbock, J, 1870) Purpur-Tornspringhale Poduromorpha Trogloxene GNR ○ ○ ○
Hypogastrura sparta Christiansen & Bellinger, 1980 Poduromorpha Trogloxene AL ○ ○ ○
Hypogastrura tullbergi (Schäffer, 1900) Poduromorpha Troglophile GNR AL ○ ○ ○
Ictalurus punctatus (Rafinesque, 1818) Channel Catfish Siluriformes Stygoxene G5 LC AL, FL ○ ○ ○
Ictidomys tridecemlineatus (Mitchill, 1821) Thirteen-Lined Ground Squirrel Rodentia Accidental G5 LC ○ ○ ○
Ictiobus bubalus (Rafinesque, 1818) Smallmouth Buffalo Cypriniformes Stygoxene G5 LC AL ○ ○ ○
Idagona jasperi Shear, 2007 Chordeumatida Troglobiont UT ○ ○ ○
Idagona lehmanensis Shear, 2007 Chordeumatida Troglobiont NV ○ 34 ○
Idagona westcotti Buckett & Gardner, 1967 Idaho Lave Tube Millipede Chordeumatida Troglobiont G1G2 ID ○ ○ ○
Illacme tobini Marek et al., 2016 Siphonophorida Troglobiont CA ○ ○ ○
Inflectarius inflectus (Say, 1821) Shagreen Stylommatophora Trogloxene G5 AL, FL, IL, KY, MO, TN ○ ○ ○
Inflectarius rugeli (Shuttleworth, 1852) Deep-Tooth Shagreen Stylommatophora Trogloxene G5 AL, GA, KY, TN ○ ○ ○
Iolania frankanstonei Hoch, 2024 Homoptera Troglobiont HI ○ ○ ○
Islandiana cavealis Ivie, 1965 Cave Sheet-web Spider Araneae Troglobiont GNR IN ○ ○ ○
Islandiana flaveola (Banks, 1892) Unequal Whiskered Money Spider Araneae Troglophile G5 IL ○ ○ ○
Islandiana lewisi Milne & Wells, 2018 cave sheet-web spider Araneae Troglobiont G1 IN ○ ○ ○
Islandiana muma Ivie, 1965 Araneae Troglobiont G1G2 AL, VA, IL ○ 1 ○
Islandiana princeps Brændegaard, 1932 Eastern Whiskered Money Spider Araneae Troglophile GNR IL ○ ○ ○
Islandiana speophila Ivie, 1965 Cavern Sheet-web Spider Araneae Troglobiont G1 WV ○ ○ ○
Islandiana unicornis Ivie, 1965 Araneae Troglobiont G2G3 OK, TX, IL ○ 4 ○
Isotoma andrei Mills, HB, 1934 Entomobryomorpha Trogloxene IL ○ ○ ○
Isotoma gelida Folsom, 1937 Entomobryomorpha Trogloxene IL ○ ○ ○
Isotoma tariva Wray, 1953 Entomobryomorpha Troglophile TN ○ ○ ○
Isotoma trispinata Tuxen, 1944 Entomobryomorpha Trogloxene IA, IL, MN ○ ○ ○
Isotoma viridis Bourlet, C, 1839 Engspringhale Entomobryomorpha Trogloxene GNR IA, MO ○ ○ ○
Isotomiella minor (Schäffer, 1896) Springtail Entomobryomorpha Trogloxene GNR IA, IL, TN ○ ○ ○
Ixodes conepati Cooley & Kohls, 1943 Ixodida Trogloxene NM, TX ○ ○ ○
Ixodes cookei Packard, 1869 American Castor Bean Tick Ixodida Trogloxene GA, KY ○ ○ ○
Ixodes scapularis Say, 1821 Black Legged Tick Ixodida Trogloxene GNR OK ○ ○ ○
Ixodes woodi Bishopp, 1911 Ixodida Trogloxene GNR TN ○ ○ ○
Jalysus wickhami Van Duzee, 1906 Spined Stilt Bug Hemiptera Accidental GNR KY ○ ○ ○
Kenkia glandulosa Hyman, 1956 Pink Cave Flatworm Tricladida Stygobiont G2G3 IA, MO ○ 11 ○
Kenkia lewisi Kenk, 1975 Lewis' cave planarian Tricladida Stygobiont G1 IL, MO ○ 15 ○
Kenkia rhynchida Hyman, 1937 Malheur Cave Flatworm Tricladida Stygobiont G1G2 NE OR ○ 2 ○
Kleptochthonius affinis Muchmore, 1976 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius anophthalmus Muchmore, 1970 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius attenuatus Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 KY ○ ○ ○
Kleptochthonius barri Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius binoculatus Muchmore, 1974 Pseudoscorpiones Troglobiont G1G2 VA ○ ○ ○
Kleptochthonius cerberus Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 KY ○ 1 ○
Kleptochthonius charon Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius daemonius Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius erebicus Muchmore, 1965 Pseudoscorpiones Troglobiont G1 KY ○ ○ ○
Kleptochthonius gertschi Malcolm & Chamberlin, 1961 Gertsch's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius griseomanus Muchmore, 2000 Gray-handed Cave Pseudoscorpion Pseudoscorpiones Troglobiont G2 IN ○ ○ ○
Kleptochthonius hageni Muchmore, 1963 Pseudoscorpiones Troglobiont G1G2 KY ○ 7 ○
Kleptochthonius henroti (Vachon, 1952) Greenbrier Valley Cave Pseudoscorpion Pseudoscorpiones Troglobiont G2 WV ○ 1 ○
Kleptochthonius hetricki Muchmore, 1974 Organ Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Kleptochthonius hubrichti Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 KY ○ ○ ○
Kleptochthonius infernalis Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius krekeleri Muchmore, 1965 Pseudoscorpiones Troglobiont G2G4 KY ○ ○ ○
Kleptochthonius lewisorum Muchmore, 2000 Lewis Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 IN ○ ○ ○
Kleptochthonius lutzi Malcolm & Chamberlin, 1961 Lutz's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius magnus Muchmore, 1966 Pseudoscorpiones Troglobiont G1 GA, TN ○ ○ ○
Kleptochthonius microphthalmus Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1G2 KY ○ ○ ○
Kleptochthonius multispinosus (Hoff, 1945) Pseudoscorpiones Troglophile AL ○ ○ ○
Kleptochthonius myopius Malcolm & J. C. Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius orpheus Muchmore, 1965 Orpheus Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Kleptochthonius packardi (Hagen, 1879) Packard's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G2 IN, KY ○ 6 ○
Kleptochthonius pluto Muchmore, 1965 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius proserpinae Muchmore, 1965 Proserpina Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Kleptochthonius proximosetus Muchmore, 1976 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius regulus Muchmore, 1970 Pseudoscorpiones Troglobiont G1G2 VA ○ ○ ○
Kleptochthonius rex Malcolm & J. C. Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius similis Muchmore, 1976 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius stygius Muchmore, 1965 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius tantalus Muchmore, 1966 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Komarekiona eatoni Gates, 1974 Kentucky Earthworm Crassiclitellata Troglophile GNR DD IL ○ ○ ○
Labidostomma neotropica Stoll, 1891 Trombidiformes Troglophile TN ○ ○ ○
Laccophilus maculosus (Germar, 1823) Dingy Diver Coleoptera Stygoxene G5 ○ ○ ○
Lacrimacandona wisei Kulkoyluoglu et al., 2017 Podocopida Stygobiont GNR TX ○ ○ ○
Lacunicambarus diogenes (Girard, 1852) Devil Crawfish Decapoda Stygoxene G5 IL, KY ○ ○ ○
Ladislavella elodes (Say, 1821) Marsh Pondsnail Lymnaeida Stygoxene ○ ○ ○
Laelaps cavernicola Packard, 1888 Mesostigmata Troglobiont KY ○ ○ ○
Lampetra aepyptera (Abbott, 1860) Least Brook Lamprey Petromyzontiformes Stygoxene G5 LC AL, KY ○ ○ ○
Lampropeltis holbrooki Stejneger, 1902 Speckled Kingsnake Squamata Accidental G5 LC AR ○ ○ ○
Larca aalbui (Muchmore, 1984) Aalbu's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Larca cavicola (Muchmore, 1981) Grand Canyon Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Larca guadalupensis (Muchmore, 1981) Guadalupe Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Larca laceyi Muchmore, 1981 Lacey's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Larca welbourni (Muchmore, 1981) Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Lascoria ambigualis Walker, 1866 Ambiguous Moth Lepidoptera Trogloxene G5 AL ○ ○ ○
Lasionycteris noctivagans (Le Conte, 1831) Black Bat Chiroptera Trogloxene G4 LC AR ○ ○ ○
Lasiurus borealis (Müller, 1776) Eastern Red Bat Chiroptera Trogloxene G3 LC AR, GA, KY, TN ○ ○ ○
Lasiurus cinereus (Palisot de Beauvois, 1796) Chiroptera Trogloxene AR, KY, TN ○ ○ ○
Lasiurus seminolus (Rhoads, 1895) Seminole Bat Chiroptera Trogloxene G5 LC ○ ○ ○
Latrodectus variolus Walckenaer, 1837 Northern Black Widow Araneae Accidental GNR TN ○ ○ ○
Lebia ornata Say, 1823 Ornate Harp Ground Beetle Coleoptera Accidental G5 AL ○ ○ ○
Leia cuneola (Adams, 1903) Diptera Troglophile AL, GA ○ ○ ○
Leiobunum aldrichi Weed, 1893 Aldrich's Harvestman Opiliones Trogloxene GNR AL ○ ○ ○
Leiobunum calcar (Wood, 1868) Opiliones Trogloxene GNR TN ○ ○ ○
Leiobunum longipes Menge, 1854 Opiliones Trogloxene KY ○ ○ ○
Leiobunum politum Weed, 1889 Opiliones Trogloxene GNR ○ ○ ○
Leiobunum vittatum (Say, 1821) Eastern Harvestman Opiliones Trogloxene ○ ○ ○
Lepidocyrtus atropurpureus Packard, 1888 Entomobryomorpha Troglophile KY ○ ○ ○
Lepidocyrtus helenae Snider, RJ, 1967 Entomobryomorpha Trogloxene G3 TN ○ ○ ○
Lepidocyrtus paradoxus Uzel, 1890 Entomobryomorpha Trogloxene GNR ○ ○ ○
Lepidocyrtus usitatus Folsom, JW, 1927 Entomobryomorpha Trogloxene ○ ○ ○
Lepomis auritus (Linnaeus, 1758) Redbreast Sunfish Centrarchiformes Stygoxene G5 LC AL ○ ○ ○
Lepomis cyanellus Rafinesque, 1819 Green Sunfish Centrarchiformes Stygoxene G5 LC AL, AR, IL, KY ○ ○ ○
Lepomis gulosus (Cuvier, 1829) Warmouth Centrarchiformes Stygoxene G5 LC FL ○ ○ ○
Lepomis humilis (Girard, 1858) Orangespotted Sunfish Centrarchiformes Stygoxene G5 LC AL ○ ○ ○
Lepomis macrochirus Rafinesque, 1819 Bluegill Centrarchiformes Stygoxene G5 LC AL, FL, KY ○ ○ ○
Lepomis megalotis (Rafinesque, 1820) Mojarra De Cuatro Cienegas Centrarchiformes Stygoxene GNR LC KY ○ ○ ○
Lepomis microlophus (Günther, 1859) Redear Sunfish Centrarchiformes Stygoxene G5 LC AL ○ ○ ○
Lepthyphantes leprosus (Ohlert, 1865) Household Fine Sheetweaver Araneae Troglophile GNR IL ○ ○ ○
Leptocera atra (Adams, 1903) Diptera Troglophile NM ○ ○ ○
Leptocera caenosa (Rondani, 1880) Diptera Troglophile GNR GA ○ ○ ○
Leptopeza compta Coquillett, 1895 Diptera Trogloxene GNR ○ ○ ○
Leptotrombidium myotis (Ewing, 1929) Trombidiformes Trogloxene GA ○ ○ ○
Lepus americanus Erxleben, 1777 Snowshoe Hare Lagomorpha Accidental G5 LC ○ ○ ○
Lesteva cribratula (Casey, 1893) Coleoptera Trogloxene AL ○ ○ ○
Lesteva pallipes LeConte, J. L., 1863 Coleoptera Troglophile G5 AL, GA, IA, IL, IN, KY, TN ○ ○ ○
Lestodiplosis grassator (Fyles, 1883) Diptera Trogloxene GNR ○ ○ ○
Lestodiplosis satiata Felt, 1919 Diptera Trogloxene ○ ○ ○
Lethemurus missus Mills, 1948 Missus Cave Springtail Entomobryomorpha Troglobiont G4 CO, IL, IN, KY, MO, TN, Mexico ○ 5 ○
Leucauge venusta (Walckenaer, 1841) Orchard Orbweaver Araneae Trogloxene GNR AL, IL, KY ○ ○ ○
Leucohya heteropoda Chamberlin, 1946 Pseudoscorpiones Troglophile ○ ○ ○
Leucohya texana Muchmore, 1986 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Leydigia leydigi (Schödler, 1863) Gewone Breedstaartwatervlo Diplostraca Stygoxene FL ○ ○ ○
Liagoceradocus lonomaka (Barnard, 1977) Amphipoda Stygobiont G1G2 HI ○ ○ ○
Liancalus genualis Loew, 1861 Diptera Trogloxene GNR GA, IN ○ ○ ○
Ligidium elrodii (Packard, 1873) Isopoda Troglophile G4 AL, AR, GA, IL, KY, TN, VA, WV ○ ○ ○
Ligidium kofoidi Maloney, 1930 Isopoda Troglobiont G1G2 CA ○ 5 ○
Limomyza cavernicola Marshall, 1997 Diptera Trogloxene CO ○ ○ ○
Limonia annulata Lackschewitz, 1940 Syyssienikiekko Diptera Trogloxene ○ ○ ○
Limonia badia (Walker, 1848) Diptera Trogloxene GNR ○ ○ ○
Limonia indigena (Osten Sacken, 1860) Diptera Trogloxene GNR ○ ○ ○
Limonia maculicosta (Coquillett, 1905) Diptera Trogloxene GNR ○ ○ ○
Limonia stulta (Osten Sacken, 1859) Diptera Trogloxene AR ○ ○ ○
Linopodes mammouthia Banks, 1897 Trombidiformes Troglobiont KS, KY ○ ○ ○
Liocranoides archeri Platnick, 1999 Archer's Two-clawed Cave Spider Araneae Troglobiont G2 AL, TN ○ ○ 6
Liocranoides coylei Platnick, 1999 Araneae Troglophile TN, VA ○ ○ ○
Liocranoides tennesseensis Platnick, 1999 Araneae Troglophile G1 TN ○ ○ ○
Liocranoides unicolor Keyserling, 1881 Araneae Troglobiont GU KY, TN, AL, GA ○ 5 ○
Liposcelis decolor (Pearman, 1925) Book Louse Psocodea Troglophile GA ○ ○ ○
Liposcelis divinatoria (Müller, 1776) Psocodea Trogloxene KY ○ ○ ○
Lirceolus bisetus (Steeves, 1968) Isopoda Stygobiont G1G2 TX ○ 14 5
Lirceolus cocytus Lewis, 2001 Isopoda Stygobiont GNR TX ○ 47 10
Lirceolus hardeni Lewis & Bowman, 1996 Isopoda Stygobiont G2G3 TX ○ 227 21
Lirceolus nidulus Lewis, 2001 Isopoda Stygobiont GNR TX, Mexico ○ 5 ○
Lirceolus pilus (Steeves, 1968) Isopoda Stygobiont G2G3 TX ○ 117 8
Lirceolus smithii (Ulrich, 1902) Texas Troglobitic Water Slater Isopoda Stygobiont G1G2 TX ○ 432 ○
Lirceus bicuspidatus Hubricht & Mackin, 1949 Isopoda Stygophile G3 AR ○ ○ ○
Lirceus bidentatus Hubricht & Mackin, 1949 Isopoda Stygophile G1 AR ○ ○ ○
Lirceus bisetus Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ 3
Lirceus culveri Estes & Holsinger, 1976 Rye Cove Cave Isopod Isopoda Stygobiont G1 VU VA ○ 9 9
Lirceus fontinalis Rafinesque, 1820 Lower Ohio Waterslater Isopoda Stygophile GNR GA, IL, KY, TN ○ ○ ○
Lirceus garmani Hubricht & Mackin, 1949 Isopoda Stygophile GNR KS, OK ○ ○ ○
Lirceus holsingeri Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ ○
Lirceus hoppinae (Faxon, 1889) Blunt-Toothed Springslater Isopoda Stygophile GNR MO, OK ○ ○ ○
Lirceus littonensis Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ 3
Lirceus orndorffi Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ 9
Lirceus slayorum Lewis & Lewis, 2023 Flat Waterslaters Isopoda Stygoxene MO ○ ○ ○
Lirceus usdagalun Holsinger & Bowman, 1973 Lee County Cave Isopod Isopoda Stygobiont G1 EN ESA VA ○ 14 11
Lirceus zigleri Lewis & Lewis, 2023 Flat Waterslaters Isopoda Stygobiont GNR VA ○ ○ 3
Lissocreagris eurydice (Muchmore, 1969) Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Lissocreagris nickajackensis (Muchmore, 1966) Pseudoscorpiones Troglobiont G1G2 AL, TN ○ ○ ○
Lissocreagris persephone (Chamberlin, 1962) Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Lissocreagris pluto (Chamberlin, 1962) Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Lissocreagris subatlantica (J. C. Chamberlin, 1962) Pseudoscorpiones Troglophile G3 AL, GA ○ ○ ○
Lissocreagris valentinei (Chamberlin, 1962) Valentine's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Litargus balteatus LeConte, 1856 Stored Grain Hairy Fungus Beetle Coleoptera Troglophile GNR IA ○ ○ ○
Lithasia obovata (Say, 1829) Shawnee Rocksnail Cerithiida Stygoxene G4 KY ○ ○ ○
Lithobates catesbeianus (Shaw, 1802) American Bullfrog Anura Trogloxene AL, AR, FL, GA, IL, IN, KY, TN ○ ○ ○
Lithobates clamitans (Latreille in Sonnini de Manoncourt & Latreille, 1801) Green Frog Anura Trogloxene AL, AR, FL, GA, IN, KY, OK, TN ○ ○ ○
Lithobates grylio (Stejneger, 1901) Anura Trogloxene FL ○ ○ ○
Lithobates heckscheri (Wright, 1924) Anura Accidental ○ ○ ○
Lithobates palustris (LeConte, 1825) Pickerel Frog Anura Trogloxene G5 LC AL, AR, CT, GA, IL, KY, OK, TN ○ ○ ○
Lithobates sphenocephalus (Cope, 1886) Southern Leopard Frog Anura Trogloxene G5 LC AR, FL, GA, KY, TN ○ ○ ○
Lithobates sylvaticus (LeConte, 1825) Anura Trogloxene AR, NY, TN, VA ○ ○ ○
Lithobius atkinsoni Bollman, 1887 Lithobiomorpha Troglophile AL, FL, GA, TN ○ ○ ○
Litocampa cookei (Packard, 1871) Cooke's Cave Dipluran Rhabdura Troglobiont G5 AL, KY, TN, VA, GA ○ 2 ○
Litocampa fieldingae (Conde, 1949) Fielding's Cave Dipluran Rhabdura Troglobiont G2G3 VA, WV ○ ○ ○
Litocampa henroti (Conde, 1949) Rhabdura Troglobiont G1G2 AL ○ ○ ○
Litocampa hoffmani Ferguson, 2009 Rhabdura Troglobiont G3 VA ○ ○ ○
Litocampa jonesi (Conde, 1949) Rhabdura Troglobiont G2G3 KY, TN ○ ○ ○
Litocampa pucketti Conde & Bareth, 1996 Puckett's Cave Dipluran Rhabdura Troglobiont G1G2 VA ○ ○ ○
Litocampa valentinei (Conde, 1949) Rhabdura Troglobiont G5 AL, TN ○ ○ ○
Lobosculum pustuloides (Bland, 1858) Tiny Liptooth Stylommatophora Trogloxene G4 AL ○ ○ ○
Lontra canadensis (Schreber, 1777) Common Otter Carnivora Stygoxene G5 LC OK ○ ○ ○
Lophomus skamania (Causey, 1972) Chordeumatida Troglobiont WA ○ 1 ○
Loxosceles panama Gertsch, 1958 Violin Spiders Araneae Trogloxene ○ ○ ○
Lucilla inermis (H. B. Baker, 1929) Oldfield Coil Stylommatophora Troglophile AL, GA, TN ○ ○ ○
Lucilla singleyana (Pilsbry, 1889) Singley's Subterranean Discus Snail Stylommatophora Trogloxene G5 IL, TN ○ ○ ○
Lumbricus rubellus Hoffmeister, 1843 Blood Worm Crassiclitellata Troglophile GNR GA, IL, NC, VA ○ ○ ○
Lumbricus terrestris Linnaeus, 1758 Nightcrawler Crassiclitellata Troglophile GNR IL ○ ○ ○
Lustrochernes minor Chamberlin, 1938 Pseudoscorpiones Troglophile ○ ○ ○
Luxilus chrysocephalus Rafinesque, 1820 Striped Shiner Cypriniformes Stygoxene G5 LC AL ○ ○ ○
Lycosa howarthi Gertsch, 1973 Araneae Troglobiont G1G2 HI ○ 2 ○
Lymantes nadineae Paquin & Anderson, 2009 Coleoptera Troglobiont GNR TX ○ 5 ○
Lynx rufus (Schreber, 1777) Bobcat Carnivora Trogloxene G5 LC AR, KY, TN ○ ○ ○
Lyogyrus retromargo (F. G. Thompson, 1968) Indented Duskysnail Littorinimorpha Stygophile GU LC FL ○ ○ ○
Lythrurus fasciolaris (Gilbert, 1891) Scarlet Shiner Cypriniformes Accidental G5 LC KY ○ ○ ○
Macrobrachium carcinus (Linnaeus, 1758) Bigclaw River Shrimp Decapoda Stygoxene G5 LC FL ○ ○ ○
Macrocera hirsuta Loew, 1870 Diptera Troglophile IN ○ ○ ○
Macrocera nobilis Johnson, 1922 Diptera Troglophile GNR AR, IL, KY, OK, TN ○ ○ ○
Macrocheles penicilliger (Berlese, 1904) Mesostigmata Troglophile TN ○ ○ ○
Macrocheles robustulus (Berlese, 1904) Mite Mesostigmata Troglophile TN ○ ○ ○
Macrocheles stygius (Packard, 1888) Mesostigmata Troglobiont G1G2 KY ○ ○ ○
Macrocheles troglodytes (Packard, 1888) Mesostigmata Troglobiont G1G2 KY ○ ○ ○
Macrocyclops albidus (Jurine, 1820) White Copepod Cyclopoida Stygophile FL, GA, IL, IN, KY, TN ○ ○ ○
Macrocyclops fuscus (Jurine, 1820) Giant Copepod Cyclopoida Stygophile IN, KY ○ ○ ○
Macromastus umpqua Shear, 1982 Chordeumatida Troglobiont G1G2 OR ○ 1 ○
Maevia inclemens (Walckenaer, 1837) Dimorphic Jumper Araneae Accidental GNR GA ○ ○ ○
Malaconothrus monodactylus (Michael, 1888) Sarcoptiformes Accidental IA ○ ○ ○
Mangora placida (Hentz, 1847) Tuftlegged Orbweaver Araneae Accidental GNR AL ○ ○ ○
Manota obscura (Walker, 1948) Diptera Troglophile IN ○ ○ ○
Margarinotus egregius (Casey, 1916) Coleoptera Accidental G5 GA ○ ○ ○
Marmota monax (Linnaeus, 1758) Groundhog Rodentia Accidental G5 LC IN ○ ○ ○
Marpissa lineata (C. L. Koch, 1846) Four-Lined Slender Jumping Spider Araneae Accidental GNR KY ○ ○ ○
Marstonia comalensis (Pilsbry & Ferriss, 1906) Comal Marstonia Littorinimorpha Stygophile G2 TX ○ ○ ○
Marstonia ogmorhaphe F. G. Thompson, 1977 Royal Marstonia Littorinimorpha Stygoxene G1 TN ○ ○ ○
Martes americana (Turton, 1806) American Marten Carnivora Trogloxene G5 LC KY ○ ○ ○
Mastigoproctus giganteus (Lucas, 1835) Giant Vinegaroon Uropygi Trogloxene GNR FL ○ ○ ○
Maymena ambita (Barrows, 1940) Araneae Troglophile GNR AL, GA, IL, TN ○ ○ ○
Mecyclothorax aa Liebherr, 2008 Coleoptera Troglobiont HI ○ ○ ○
Mecynogea lemniscata (Walckenaer, 1841) Basilica Orbweaver Araneae Accidental GNR AL ○ ○ ○
Megacyclops donnaldsoni (Chappuis, 1929) Donaldson's Cave Copepod Cyclopoida Stygobiont G3G4 GA, IN, KY, TN ○ 1 ○
Megacyclops latipes (Lowndes, 1927) Cyclopoida Stygoxene IN, TN ○ ○ ○
Megacyclops viridis (Jurine, 1820) Cyclopoid Copepod Cyclopoida Stygophile IN ○ ○ ○
Megalothorax minimus Womersley, 1939 Neelipleona Troglophile IA, IL ○ ○ ○
Megalothorax tristani Denis, J-R, 1933 Neelipleona Trogloxene IL ○ ○ ○
Megapallifera mutabilis (Hubricht, 1951) Changeable Mantleslug Stylommatophora Trogloxene G5 KY ○ ○ ○
Megascops asio (Linnaeus, 1758) Eastern Screech Owl Strigiformes Trogloxene G5 LC AL, OK ○ ○ ○
Megaselia breviterga (Lundbeck, 1919) Diptera Troglophile GA ○ ○ ○
Megaselia cavernicola (Brues, 1906) Diptera Troglophile GNR AL, AR, GA, IA, IL, KY, TN ○ ○ ○
Megaselia nigriceps (Loew, 1866) Diptera Troglophile GNR IN ○ ○ ○
Megaselia rufipes (Meigen, 1804) Hump-Backed Fly Diptera Troglophile GNR IN, KY, NM ○ ○ ○
Megaselia spelunciphila Reeves & Disney, 1999 Diptera Troglophile GA ○ ○ ○
Melanoides tuberculata (O. F. Müller, 1774) Cerithiida Stygoxene FL ○ ○ ○
Melanolestes picipes (Herrich-Schaeffer, 1848) Black Corsair Heteroptera Trogloxene GNR IL ○ ○ ○
Mephitis mephitis (Schreber, 1776) Striped Skunk Carnivora Trogloxene G5 LC AR, KY, OK ○ ○ ○
Mermessus antraeus (Crosby, 1926) Araneae Trogloxene GNR IL, NM, TX ○ ○ ○
Mermessus denticulatus (Banks, 1898) Amerikaanse Kasdwergspin Araneae Trogloxene GNR AL, IL, NM ○ ○ ○
Mermessus indicabilis (Crosby & Bishop, 1928) Araneae Trogloxene GNR IL ○ ○ ○
Mermessus maculatus (Banks, 1892) Sheet-Web Spiders Araneae Trogloxene GNR AL, GA, IL, MO, TN ○ ○ ○
Mermessus tridentatus (Emerton, 1882) Spider Araneae Trogloxene GNR AL, IL, TN ○ ○ ○
Meropelopia flavifrons (Johannsen, 1905) Diptera Trogloxene ○ ○ ○
Mesaphorura iowensis (Mills, HB, 1932) Poduromorpha Troglophile GNR GA, IA ○ ○ ○
Mesaphorura krausbaueri Jordana et al., 1987 Poduromorpha Troglophile GA ○ ○ ○
Mesaphorura pacifica Rusek, 1976 Poduromorpha Accidental GNR IA ○ ○ ○
Mesaphorura silvicola (Folsom, 1932) Poduromorpha Trogloxene IL ○ ○ ○
Mesocyclops edax (Forbes, 1891) Cyclopoida Stygoxene IL, IN, KY ○ ○ ○
Mesodon clausus (Say, 1821) Yellow Globelet Stylommatophora Trogloxene G5 IL ○ ○ ○
Mesodon thyroidus (Say, 1817) White-Lip Globe Stylommatophora Trogloxene G5 FL, TN ○ ○ ○
Mesodon zaletus (A. Binney, 1837) Toothed Globe Stylommatophora Trogloxene G5 AL, FL, KY ○ ○ ○
Mesomphix andrewsae (Pilsbry, 1895) Mountain Button Stylommatophora Trogloxene G3 TN ○ ○ ○
Mesomphix anurus Hubricht, 1962 Frog Button Stylommatophora Trogloxene G4 AL ○ ○ ○
Mesomphix capnodes (W. G. Binney, 1857) Dusky Button Stylommatophora Trogloxene G5 AL ○ ○ ○
Mesomphix cupreus (Rafinesque, 1831) Copper Button Stylommatophora Trogloxene KY, TN ○ ○ ○
Mesomphix perlaevis (Pilsbry, 1900) Smooth Button Stylommatophora Trogloxene G4 KY ○ ○ ○
Mesomphix pilsbryi (G. H. Clapp, 1904) Striate Button Stylommatophora Trogloxene G4 AL ○ ○ ○
Meta ovalis (Gertsch, 1933) Cave Orb Weaver Araneae Troglophile G5 AL, AR, GA, IA, IL, IN, KY, MO, OK, TN, VA ○ ○ ○
Metajapyx subterraneus (Packard, 1874) Dicellurata Troglophile KY ○ ○ ○
Metalimnobia cinctipes (Say, 1823) Diptera Trogloxene GNR ○ ○ ○
Metaltella simoni (Keyserling, 1878) Hacklemesh Weaver Araneae Accidental GNR GA ○ ○ ○
Metisotoma grandiceps Stebaeva, 2015 Entomobryomorpha Accidental IA ○ ○ ○
Mexichthonius exoticus Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Mexistenasellus coahuila Cole & Minckley, 1972 Coahuila Groundwater Isopod Isopoda Stygobiont G2G3 EN TX, Mexico ○ 44 2
Mexistenasellus floridensis Lewis & Sawicki, 2016 Florida Cave Isopod Isopoda Stygobiont G1 FL ○ 3 ○
Mexiweckelia hardeni Holsinger, 1992 Amphipoda Stygobiont G2G3 TX ○ 21 ○
Microbisium parvulum (Banks, 1895) Pseudoscorpiones Trogloxene GNR IL, KY, TN ○ ○ ○
Microcreagris grandis Muchmore, 1962 Great Basin Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 NV ○ 12 2
Microcyclops rubellus (Lilljeborg, 1901) Cyclopoida Stygoxene IL, IN, KY, TN ○ ○ ○
Microcylloepus pusillus (LeConte, 1852) Coleoptera Stygoxene GNR TX ○ ○ ○
Micropogonias undulatus (Linnaeus, 1766) Croaker Perciformes Accidental G5 LC ○ ○ ○
Micropterus dolomieu Lacepède, 1802 Smallmouth Bass Centrarchiformes Stygoxene GNR LC TN ○ ○ ○
Micropterus punctulatus (Rafinesque, 1819) Spotted Bass Centrarchiformes Stygoxene G5 LC KY ○ ○ ○
Micropterus salmoides (Lacepède, 1802) American Black Bass Centrarchiformes Stygoxene GNR LC AL, FL ○ ○ ○
Microtus ochrogaster (Wagner, 1842) Prairie Vole Rodentia Accidental G5 LC KY ○ ○ ○
Microtus pennsylvanicus (Ord, 1815) Meadow Vole Rodentia Accidental G5 LC KY ○ ○ ○
Microtus pinetorum (Le Conte, 1830) Pine Vole Rodentia Accidental G5 LC AR, KY ○ ○ ○
Microtus xanthognathus (Leach, 1815) Taiga Vole Rodentia Accidental G5 LC ○ ○ ○
Microvelia americana (Uhler, 1884) Hemiptera Stygoxene GNR GA, IL ○ ○ ○
Miktoniscus barri Vandel, 1965 Barr's Terrestrial Cave Isopod Isopoda Troglobiont G2G4 IN, KY, TN ○ 4 ○
Miktoniscus linearis (Patience, 1908) Isopoda Troglophile AL ○ ○ ○
Miktoniscus medcofi Van Name, 1940 Small Alabama Sowbug Isopoda Troglophile GNR AL, FL, IL, NC, TN, VA, WV ○ ○ ○
Miktoniscus morganensis Schultz, 1976 Isopoda Troglophile AL ○ ○ ○
Miktoniscus oklahomensis Vandel, 1965 Racovitza's Cave Isopod Isopoda Troglobiont G1G2 OK ○ ○ ○
Miktoniscus racovitzai Vandel, 1950 Racovitza's Terrestrial Cave Isopod Isopoda Troglobiont G3G4 VA, KY, OK ○ 20 ○
Mimetus notius Chamberlin, 1923 Reticulated Pirate Spider Araneae Accidental GNR ○ ○ ○
Mimetus puritanus Chamberlin, 1923 Common Pirate Spider Araneae Accidental GNR AL ○ ○ ○
Minicreagris pumila (Muchmore, 1969) Pseudoscorpiones Troglophile GNR AL, GA ○ ○ ○
Mixojapyx reddelli Muegge, 1992 Dicellurata Troglobiont G3G4 TX ○ 9 ○
Mochlonyx cinctipes (Coquillett, 1903) Diptera Trogloxene GNR ○ ○ ○
Molothrus ater (Boddaert, 1783) Brown-Headed Cowbird Passeriformes Accidental G5 LC IL ○ ○ ○
Monopis crocicapitella (Clemens, 1859) Bird Nest Moth Lepidoptera Trogloxene GNR AL, TN ○ ○ ○
Monopsyllus wagneri Baker, 1904 Siphonaptera Accidental IA ○ ○ ○
Montanabathynella pecosensis Camacho & Hutchins, 2021 Bathynellacea Stygobiont TX ○ ○ ○
Montanabathynella salish Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Moraria cristata Chappuis, 1929 Harpacticoida Stygoxene IN ○ ○ ○
Mormoops megalophylla (Peters, 1864) Ghost-Faced Bat Chiroptera Trogloxene G5 LC TX ○ ○ ○
Morone saxatilis (Walbaum, 1792) Maryland Bass Hybrid Acanthuriformes Stygoxene G3 LC FL ○ ○ ○
Moxostoma erythrurum (Rafinesque, 1818) Golden Redhorse Cypriniformes Stygoxene G5 LC KY ○ ○ ○
Mundochthonius cavernicola Muchmore, 1968 Pseudoscorpiones Troglobiont G1G2 IL, MO ○ ○ ○
Mundochthonius holsingeri Benedict & Malcolm, 1974 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Mus musculus Linnaeus, 1758 House Mouse Rodentia Trogloxene G5 LC AL, KY ○ ○ ○
Musca domestica Linnaeus, 1758 Common House Fly Diptera Accidental GNR IN ○ ○ ○
Muscina prolapsa (Harris, 1780) Muscoid Fly Diptera Trogloxene GNR GA ○ ○ ○
Musculium securis (Prime, 1852) Sphaeriida Accidental ○ ○ ○
Mustela nivalis Linnaeus, 1766 Least Weasel Carnivora Trogloxene G5 LC ○ ○ ○
Mycetophila analis (Coquillett, 1901) Diptera Troglophile IN ○ ○ ○
Mycetophila byersi (Laffoon, 1957) Diptera Troglophile GNR ○ ○ ○
Mycetophila fisherae (Laffoon, 1957) Diptera Troglophile GNR ○ ○ ○
Mycetophila ichneumonea Say, 1823 Diptera Troglophile GNR IN, TN ○ ○ ○
Mycetophila ruficollis Meigen, 1818 Diptera Troglophile ○ ○ ○
Mycetophila sordida Wulp, 1875 Diptera Troglophile GNR ○ ○ ○
Mycomya nitida (Zetterstedt, 1852) Diptera Troglophile GNR ○ ○ ○
Myodopsylla insignis (Rothschild, 1903) Bat Flea Siphonaptera Trogloxene GNR KY ○ ○ ○
Myotis austroriparius (Rhoads, 1897) Southeastern Myotis Chiroptera Trogloxene G4 LC AL, AR, FL, GA, IL, KY ○ ○ ○
Myotis grisescens A. H. Howell, 1909 Gray Bat Chiroptera Trogloxene G3 VU ESA AL, AR, FL, GA, IL, KS, KY, MO, MS, OK, TN, VA ○ ○ ○
Myotis leibii (Audubon & Bachman, 1842) Eastern Small-Footed Bat Chiroptera Trogloxene G4 EN AR, GA, KY, PA, TN, VA ○ ○ ○
Myotis lucifugus (Le Conte, 1831) Little Brown Bat Chiroptera Trogloxene G3 EN AL, AR, GA, IL, IN, KY, PA, TN, VA ○ ○ ○
Myotis septentrionalis (Trouessart, 1897) Northern Long-eared Bat Chiroptera Trogloxene G2 NT ESA AL, AR, FL, GA, IL, KY, OK, PA, TN, VA ○ ○ ○
Myotis sodalis Miller & G. M. Allen, 1928 Cluster Bat Chiroptera Trogloxene G2 NT ESA AL, AR, FL, GA, IL, KY, PA, TN, VA ○ ○ ○
Myotis velifer (J. A. Allen, 1890) Cave Myotis Chiroptera Trogloxene G4 LC ○ ○ ○
Myrmecina americana Emery, 1895 American Mite-Eating Ant Hymenoptera Accidental G5 GA ○ ○ ○
Mystacides sepulchralis (Walker, 1852) Black Dancer Caddisfly Amblypygi Troglophile G5 ○ ○ ○
Nadabius ameles Chamberlin, 1944 Lithobiomorpha Trogloxene MO ○ ○ ○
Nais variabilis Piguet, 1906 Aquatic Oligochaete Worm Tubificida Stygophile GNR WV ○ ○ ○
Namiotkocypria haysensis Kulkoyluoglu, 2018 Podocopida Stygobiont TX ○ ○ ○
Nampabius mycophor Chamberlin, 1940 Lithobiomorpha Trogloxene AL ○ ○ ○
Nampabius turbator Crabill, 1952 Lithobiomorpha Troglobiont G1 VA ○ ○ ○
Nannopetalum fontis Shear, 2003 Chordeumatida Troglobiont AL ○ ○ ○
Nannopetalum vespertilo Shear, 2003 Chordeumatida Troglobiont NC ○ ○ ○
Narceus americanus (Palisot de Beauvouis, 1817) American Giant Millipede Spirobolida Trogloxene G5 AL, AR, TN ○ ○ ○
Narceus annularis (Rafinesque, 1820) Spirobolida Trogloxene G5 TN ○ ○ ○
Neanura hawaiiensis (Bellinger & Christiansen, 1974) Poduromorpha Troglobiont G1G2 HI ○ ○ ○
Neanura muscorum (Templeton, R in Templeton, R & Westwood, JO, 1836) Moss Springtail Poduromorpha Trogloxene GNR IA ○ ○ ○
Neanura persimilis Mills, HB, 1934 Poduromorpha Trogloxene GNR IL ○ ○ ○
Neanura pseudoquadrioculata Stach, 1951 Poduromorpha Trogloxene IL ○ ○ ○
Neaphaenops tellkampfi (Erichson, 1844) Coleoptera Troglobiont G3G4 KY, IN ○ ○ ○
Necrophilus pettitii Horn, 1880 Coleoptera Trogloxene GNR AL, TN ○ ○ ○
Neelides minutus (Folsom, JW, 1901) Neelipleona Troglophile GNR GA, IA, IL ○ ○ ○
Nelima elegans (Weed, 1889) Opiliones Trogloxene KY, TN ○ ○ ○
Nelima paessleri (Roewer, 1910) Aggregating Harvestman Opiliones Trogloxene GNR ○ ○ ○
Nelsonites jonesi Valentine, 1952 Coleoptera Troglobiont G2G3 KY ○ ○ ○
Nelsonites walteri Valentine, 1952 Coleoptera Troglobiont G3 TN ○ 3 ○
Nemadus hornii Hatch, 1933 Coleoptera Troglophile AL, GA, TN ○ ○ ○
Nemoura trispinosa Claassen, 1923 Plecoptera Stygoxene IL ○ ○ ○
Neoallochernes incertus Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 NM ○ ○ ○
Neoallochernes stercoreus (Turk, 1949) Pseudoscorpiones Troglobiont G4 TX ○ 10 ○
Neobisnius paederoides (LeConte, J. L., 1863) Coleoptera Trogloxene GNR IL ○ ○ ○
Neochthonius imperialis Muchmore, 1996 Empire Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ 1 ○
Neochthonius troglodytes Muchmore, 1969 Pseudoscorpiones Troglobiont G2G3 CA ○ ○ ○
Neoechinorhynchus cylindratus (Van Cleave, 1913) Neoechinorhynchida Stygoxene KY ○ ○ ○
Neogale vison (Schreber, 1777) American Mink Carnivora Trogloxene G5 LC GA, IN ○ ○ ○
Neohelix albolabris (Say, 1817) Eastern Whitelip Stylommatophora Trogloxene G5 ○ ○ ○
Neolithobius voracior Chamberlin (1912) Lithobiomorpha Trogloxene GA ○ ○ ○
Neoporus clypealis (Sharp, 1882) Coleoptera Stygoxene GNR FL ○ ○ ○
Neoscona domiciliorum (Hentz, 1847) Red-Femured Spotted Orbweaver Araneae Accidental GNR ○ ○ ○
Neospintharus trigonum (Hentz, 1850) Horned Parasitic Cobweaver Araneae Trogloxene GNR AL, IN ○ ○ ○
Neotoma floridana (Ord, 1818) Eastern Woodrat Rodentia Trogloxene G5 LC ESA AL, AR, FL, GA, OK ○ ○ ○
Neotoma magister Baird, 1857 Allegheny Woodrat Rodentia Trogloxene G3 NT AL, IN, KY, TN, VA ○ ○ ○
Nepalomyia sombrea (Harmston & Knowlton, 1945) Diptera Trogloxene GA ○ ○ ○
Neriene radiata (Walckenaer, 1841) Filmy Dome Spider Araneae Trogloxene G5 GA, IL, IN, KY ○ ○ ○
Nerodia erythrogaster (Forster, 1771) Plain-Bellied Water Snake Squamata Accidental G5 LC ○ ○ ○
Nerodia fasciata (Linnaeus, 1766) Southern Water Snake Squamata Accidental G5 LC FL ○ ○ ○
Nerodia sipedon (Linnaeus, 1758) Common Watersnake Squamata Accidental G5 LC GA, TN ○ ○ ○
Nesidiolestes ana Gagne & Howarth, 1974 Ana Wingless Thread Bug Heteroptera Troglobiont G1G2 NE HI ○ 1 ○
Nesticus barri Gertsch, 1984 Barr's Cave Spider Araneae Troglobiont G3 AL, TN ○ 4 57
Nesticus barrowsi Gertsch, 1984 Araneae Troglobiont G1G2 TN ○ 4 9
Nesticus brimleyi Gertsch, 1984 Araneae Troglobiont G1G2 NC ○ 7 9
Nesticus carolinensis (Bishop, 1950) Linville Caverns Spider Araneae Troglophile G2 NC ○ ○ ○
Nesticus carteri Emerton, 1875 Carter's Cave Spider Araneae Troglophile GNR IN, KY, TN, VA, WV ○ ○ ○
Nesticus cressleri Zigler & Milne, 2022 Araneae Troglobiont GNR GA ○ 2 8
Nesticus dilutus Gertsch, 1984 Grassy Creek Cave Spider Araneae Troglobiont G1 TN ○ 2 4
Nesticus furtivus Gertsch, 1984 Racoon Mountain Caverns Cave Spider Araneae Troglobiont G1 TN ○ 2 7
Nesticus georgia Gertsch, 1984 Georgia Cave Spider Araneae Troglobiont G1G2 GA ○ 2 4
Nesticus gertschi Coyle & McGarity, 1992 Araneae Troglophile G1 TN ○ ○ ○
Nesticus holsingeri Gertsch, 1984 Holsinger's Cave Spider Araneae Troglobiont G3G4 VA ○ 4 16
Nesticus jonesi Gertsch, 1984 Cave Spring Cave Spider Araneae Troglobiont G1 AL ○ 1 4
Nesticus lula Zigler & Milne, 2022 Araneae Troglobiont GNR GA ○ 2 6
Nesticus mimus Gertsch, 1984 Araneae Troglophile G3 VA ○ ○ ○
Nesticus paynei Gertsch, 1984 Araneae Troglobiont G3G4 TN, VA ○ 18 27
Nesticus pecki Hedin & Dellinger, 2005 Monteagle Cave Spider Araneae Troglobiont GNR TN ○ 3 4
Nesticus potterius (Chamberlin, 1933) Araneae Troglobiont G1G2 CA ○ ○ ○
Nesticus reclusus Gertsch, 1984 Lost Nantahala Cave Spider Araneae Troglophile G3 NC ○ ○ ○
Nesticus sodanus Gertsch, 1984 Araneae Troglophile CA ○ ○ ○
Nesticus stupkai Gertsch, 1984 Araneae Troglophile TN ○ ○ ○
Nesticus stygius Gertsch, 1984 Araneae Troglobiont G1G2 TN ○ 3 7
Nesticus tennesseensis (Petrunkevitch, 1925) Tennessee Cave Spider Araneae Troglobiont G3G4 TN, VA, WV ○ 10 25
Neuratelia sayi (Aldrich, 1897) Diptera Troglophile IN ○ ○ ○
Nevadesmus hubbsi (Chamberlin, 1943) Polydesmida Troglobiont GNR NV ○ ○ ○
Nevadesmus ophimontis Shear, 2009 Polydesmida Troglobiont NV ○ 1 ○
Nevadesmus superstitiona Shear, Pape & Marek, 2025 Polydesmida Troglobiont AZ ○ ○ ○
Niptus arcanus Aalbu & Andrews, 1992 Coleoptera Troglobiont CA ○ 18 ○
Nitidotachinus scrutator (Gemminger & Harold, 1868) Coleoptera Trogloxene GNR IN ○ ○ ○
Nitocrellopsis texana Fiers & Iliffe, 2000 Harpacticoida Stygobiont GNR TX ○ 1 ○
Nopoiulus kochii (Gervais, 1847) Haakwang Julida Trogloxene G5 KY, TN ○ ○ ○
Notemigonus crysoleucas (Mitchill, 1814) Golden Shiner Cypriniformes Accidental G5 LC AL ○ ○ ○
Notiobia terminata (Say, 1823) Coleoptera Accidental G5 AL ○ ○ ○
Notophthalmus viridescens (Rafinesque, 1820) Eastern Newt Caudata Accidental G5 LC AL, GA, KY, TN ○ ○ ○
Notropis atherinoides Rafinesque, 1818 Emerald Shiner Cypriniformes Stygoxene G5 LC KY ○ ○ ○
Novobisium carolinense (Banks, 1895) Pseudoscorpiones Trogloxene GA ○ ○ ○
Novobisium ingratum (J. C. Chamberlin, 1962) Pseudoscorpiones Troglophile GNR AL ○ ○ ○
Nycteridopsylla chapini Jordan, 1929 Siphonaptera Trogloxene KY ○ ○ ○
Nycticeius humeralis (Rafinesque, 1818) Evening Bat Chiroptera Trogloxene G5 LC AL, AR, KY ○ ○ ○
Nyctinomops femorosaccus (Merriam, 1889) Pocketed Free-Tailed Bat Chiroptera Trogloxene G5 LC ○ ○ ○
Oaphantes prometheus Paquin et al., 2020 Araneae Troglobiont CA ○ 2 ○
Occasjapyx kofoidi (Silvestri, 1928) Dicellurata Troglobiont CA ○ ○ ○
Ochrotrichia spinosa (Ross, 1938) Trichoptera Trogloxene G5 AR ○ ○ ○
Ocrepeira ectypa (Walckenaer, 1841) Araneae Accidental GNR AL ○ ○ ○
Octolasion lacteum (Örley, 1881) White Worm Crassiclitellata Troglophile TN, WV ○ ○ ○
Octolasion tyrtaeum (Savigny in Cuvier, 1826) Woodland White Worm Crassiclitellata Troglophile GNR AL, GA, IL, NC, TN, VA ○ ○ ○
Odiellus pictus (Wood, 1868) Painted Harvestman Opiliones Trogloxene GNR ○ ○ ○
Odocoileus virginianus (Zimmermann, 1780) White-Tailed Deer Artiodactyla Accidental G5 LC ESA AL, FL, KY ○ ○ ○
Odontotaenius disjunctus (Illiger, 1800) Patent Leather Beetle Coleoptera Accidental GNR IL ○ ○ ○
Oecothea fenestralis (Fallén, 1820) Diptera Trogloxene GNR IN, KY, TN ○ ○ ○
Oecothea specus (Aldrich, 1897) Small Cave Sun Fly Diptera Troglophile AL, AR, GA, IA, IL, IN, KY, TN, VA, WI ○ ○ ○
Oedipodrilus macbaini (Holt, 1955) Branchiobdellida Stygophile TN ○ ○ ○
Oliarus gagneri Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus kalaupapae Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus lorettae Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus makaiki Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus polyphemus Fennah, 1973 Homoptera Troglobiont G1G2 HI ○ 4 29
Oliarus priola Fennah, 1973 Homoptera Troglobiont G1G2 HI ○ 2 ○
Oliarus waikau Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Omalium rivulare (Paykull, 1789) Ocillate Rove Beetle Coleoptera Trogloxene GNR TN ○ ○ ○
Omorgus alatus (MacLeay, 1888) Coleoptera Trogloxene ○ ○ ○
Omorgus amictus (Haaf, 1954) Coleoptera Trogloxene ○ ○ ○
Omorgus batesi (Harold, 1872) Coleoptera Trogloxene ○ ○ ○
Omorgus brucki (Harold, 1872) Coleoptera Trogloxene ○ ○ ○
Omorgus carinatus Loomis, 1922 Coleoptera Trogloxene ○ ○ ○
Omorgus costatus (Wiedemann, 1823) Coleoptera Trogloxene ○ ○ ○
Omorgus fuliginosus (Robinson, 1941) Coleoptera Trogloxene ○ ○ ○
Omorgus lindemannae (Petrovitz, 1975) Coleoptera Trogloxene ○ ○ ○
Omorgus punctatus (Germar, 1823) Coleoptera Trogloxene ○ ○ ○
Omorgus rubricans (Robinson, 1946) Coleoptera Trogloxene ○ ○ ○
Omorgus suberosus (Fabricius, 1775) Hide Beetle Coleoptera Trogloxene GNR ○ ○ ○
Oncopodura cruciata Bonet, 1943 Entomobryomorpha Troglobiont G1G2 MT ○ 10 ○
Oncopodura fenestra Christiansen & Bonet, 1996 Entomobryomorpha Troglobiont G2G3 TX ○ 1 ○
Oncopodura hoffi Christiansen & Bonet, 1980 Hoff's Cave Springtail Entomobryomorpha Troglobiont G1G2 MO ○ 3 ○
Oncopodura hubbardi Christiansen & Bellinger 1996 Entomobryomorpha Troglobiont G2 VA ○ 1 ○
Oncopodura iowae Christiansen, 1980 Entomobryomorpha Troglobiont G3G4 IA, IL, MO ○ 3 ○
Oncopodura mala Christiansen, K & Bellinger, P, 1996 Malheur Cave Springtail Entomobryomorpha Troglophile G2 CA, OR ○ ○ ○
Oncopodura prietoi Bonet, 1943 Entomobryomorpha Troglobiont G1G2 NM, Mexico ○ 2 ○
Oncopodura subhoffi Christiansen & Bellinger, 1998 Entomobryomorpha Troglobiont G3 CO ○ 2 ○
Oniscus asellus Linnaeus, 1758 Common Shiny Woodlouse Isopoda Trogloxene GNR ○ ○ ○
Onychiurus eous Christiansen & Bellinger, 1980 Poduromorpha Troglophile IA ○ ○ ○
Onychiurus gelus Christiansen & Bonet, 1980 Poduromorpha Troglobiont G2G3 IA, WA, WI ○ ○ ○
Onychiurus janus Christiansen & Bellinger, 1980 Poduromorpha Troglobiont G2G3 AL, VA, WV ○ ○ ○
Onychiurus obesus Mills, 1934 Poduromorpha Troglobiont G3G4 IA, MN, MO, WI ○ ○ ○
Onychiurus opus Christiansen & Bellinger, 1980 Poduromorpha Troglophile IA ○ ○ ○
Onychiurus oregonensis Bonet, 1931 Poduromorpha Troglobiont G1G2 OR ○ ○ ○
Onychiurus paro Christiansen & Bellinger, 1980 Poduromorpha Troglobiont G1 AL ○ ○ ○
Onychiurus parvicornis Mills, 1934 Poduromorpha Troglophile IA ○ ○ ○
Onychiurus pipistrellae Soto-Adames & Taylor, 2013 Poduromorpha Troglobiont IL ○ ○ ○
Onychiurus pseudoarmatus Aitchison, CW, 1984 Poduromorpha Trogloxene TN ○ ○ ○
Onychiurus pseudofimetarius Folsom, 1917 Poduromorpha Troglophile TN ○ ○ ○
Onychiurus reluctus Christiansen, 1961 Poduromorpha Troglobiont GNR WV, Mexico, IA, IL, TN ○ 9 ○
Onychiurus similis Folsom, 1917 Poduromorpha Troglophile IA ○ ○ ○
Opheodrys aestivus (Linnaeus, 1766) Greensnake Squamata Accidental G5 LC AR ○ ○ ○
Ophisaurus ventralis (Linnaeus, 1766) Eastern Glass Lizard Squamata Accidental G5 LC FL ○ ○ ○
Ophyiulus pilosus (Newport, 1843) Furry Snake Millipede Julida Trogloxene G5 AL, KY, TN ○ ○ ○
Opiona catorycha Shear & Crawford, 2019 Chordeumatida Troglobiont OR ○ ○ ○
Opiona graeningi Shear, 2011 Chordeumatida Troglobiont CA ○ 1 ○
Oppia nitens Koch, 1835 Sarcoptiformes Accidental GNR IA ○ ○ ○
Orchesella hexfasciata (Harvey, FL, 1895) Entomobryomorpha Accidental GNR IA ○ ○ ○
Orchopeas leucopus (Baker, 1904) Pulga Siphonaptera Trogloxene GNR ○ ○ ○
Orchopeas sexdentatus (Baker, 1904) Siphonaptera Trogloxene KY, VA ○ ○ ○
Orconectes australis (Rhoades, 1941) Southern Cave Crayfish Decapoda Stygobiont G5 LC AL, TN, KY ○ 158 152
Orconectes barri Buhay & Crandall, 2008 Cumberland Plateau Cave Crayfish Decapoda Stygobiont G2 DD KY, TN ○ 14 66
Orconectes incomptus Hobbs & Barr, 1972 Tennessee Cave Crayfish Decapoda Stygobiont G2 VU TN ○ 8 17
Orconectes inermis Cope, 1872 Ghost Crayfish Decapoda Stygobiont G5 LC IN, KY, TN ○ 114 3
Orconectes packardi Rhoades, 1944 Appalachian Cave Crayfish Decapoda Stygobiont G2 EN KY, TN ○ 34 33
Orconectes pellucidus (Tellkampf, 1844) Mammoth Cave Crayfish Decapoda Stygobiont G4 LC KY, TN, AL, IN ○ 139 2
Oregonasellus elliotti Lewis, 2008 Isopoda Stygobiont OR ○ ○ ○
Oreonetides beattyi Paquin et al., 2009 Beatty's Sheetweb Cave Spider Araneae Troglobiont GNR IN, MD, PA, TN, VA ○ ○ ○
Oreonetides vaginatus (Thorell, 1872) Undergrowth Sheet-Web Weaver Araneae Troglophile G5 ○ ○ ○
Oreophantes recurvatus (Emerton, 1913) Araneae Troglophile G5 ○ ○ ○
Orfelia elegans (Coquillett, 1895) Diptera Troglophile ○ ○ ○
Origanates rostratus (Emerton, 1882) Araneae Trogloxene GNR AL, TN ○ ○ ○
Ormosia affinis (Lundbeck, 1898) Cranefly Diptera Trogloxene GNR ○ ○ ○
Ormosia meigenii (Osten Sacken, 1860) Diptera Trogloxene GNR IN ○ ○ ○
Oropsylla arctomys (Baker, 1904) Siphonaptera Accidental GNR IA ○ ○ ○
Orthocyclops modestus (Herrick, 1883) Cyclopoida Stygoxene IL, IN, TN ○ ○ ○
Orus rubens (Casey, 1905) Coleoptera Troglophile NM ○ ○ ○
Oryzaephilus surinamensis (Linnaeus, 1758) Sawtoothed Grain Beetle Coleoptera Accidental GNR IA ○ ○ ○
Osphranticum labronectum Forbes S.A., 1882 Calanoida Stygoxene KY, TN ○ ○ ○
Osteopilus septentrionalis (Duméril & Bibron, 1841) Cuban Treefrog Anura Accidental G5 LC FL ○ ○ ○
Otiorhynchus ovatus (Linnaeus, 1758) Strawberry Root Weevil Coleoptera Accidental GNR ○ ○ ○
Otostigmus cooperi Chamberlin, 1942 Scolopendromorpha Troglophile ○ ○ ○
Oxidus gracilis (Koch, 1847) Greenhouse Millipede Polydesmida Troglophile G5 AL, FL, GA, KY, OK, PA, TN, VA ○ ○ ○
Oxyopes salticus Hentz, 1845 Lynx Spider Araneae Accidental GNR AL ○ ○ ○
Oxytelus exiguus Erichson, 1840 Coleoptera Trogloxene IL ○ ○ ○
Ozarkia alabama (Gertsch, 1974) Araneae Troglobiont GNR AL ○ 27 ○
Ozarkia arkansa (Gertsch, 1974) Araneae Troglobiont AR ○ 8 ○
Ozarkia blanda (Gertsch, 1974) Araneae Troglobiont GNR AL ○ 1 ○
Ozarkia georgia (Gertsch, 1974) Araneae Troglobiont GNR GA ○ 25 ○
Ozarkia novaegalleciae (Brignoli, 1979) Araneae Troglobiont NM ○ ○ ○
Ozarkia serena (Gertsch, 1974) Araneae Troglobiont G1G2 AL ○ 15 ○
Pachydesmus crassicutis (Wood, 1864) Polydesmida Accidental G5 AL ○ ○ ○
Pacificabathynella kalispellensis Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Pacificabathynella ruthae Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Pacificabathynella sequoiae Schminke & Noodt, 1988 Bathynellacea Stygobiont GNR CA, WA ○ 5 ○
Pacificabathynella stanfordi Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Pacificabathynella yupik Camacho et al., 2016 Bathynellacea Stygobiont AK ○ 47 ○
Packardesmus cavicola (Packard, 1877) Polydesmida Troglophile UT ○ ○ ○
Paeromopus cavicolens Chamberlin, 1953 Julida Troglophile CA ○ ○ ○
Paitobius arienus Chamberlin (1911) Lithobiomorpha Trogloxene AL ○ ○ ○
Paitobius carolinae Chamberlin (1911) Lithobiomorpha Trogloxene AL ○ ○ ○
Paitobius juventus Bollman (1887) Lithobiomorpha Trogloxene AL ○ ○ ○
Palaemon kadiakensis (M. J. Rathbun, 1902) Mississippi Grass Shrimp Decapoda Accidental G5 FL ○ ○ ○
Palaemon paludosus (Gibbes, 1850) Eastern Grass Shrimp Decapoda Stygoxene G5 FL ○ ○ ○
Palaemonetes antrorum Benedict, 1896 Balcones Cave Shrimp Decapoda Stygobiont G2G3 EN TX ○ ○ ○
Palaemonetes cummingi Chace, 1954 Squirrel Chimney Cave Shrimp Decapoda Stygobiont GH CR ESA FL ○ ○ ○
Palaemonias alabamae Smalley, 1961 Alabama Cave Shrimp Decapoda Stygobiont G1 EN ESA AL ○ 5 14
Palaemonias ganteri Hay, 1901 Mammoth Cave Shrimp Decapoda Stygobiont G1 VU ESA KY ○ 11 ○
Paleonura petebellingeri Palacios-Vargas & Benito, 2007 Poduromorpha Troglobiont G1 VA ○ ○ ○
Pangaeus docilis (Walker, 1867) Hemiptera Troglophile ○ ○ ○
Panstrongylus geniculatus (Latreille, 1811) Barbeiro Heteroptera Trogloxene ○ ○ ○
Pantelozetes cavaticus (Kunst, 1962) Sarcoptiformes Trogloxene TN ○ ○ ○
Pantherophis guttatus (Linnaeus, 1766) Corn Snake Squamata Trogloxene G5 LC ○ ○ ○
Pantherophis obsoletus (Say, 1823) Pilot Black Snake Squamata Trogloxene G5 LC AR, KY ○ ○ ○
Pantherophis quadrivittatus (Holbrook, 1836) Yellow Rat Snake (Quadrivittata) Squamata Trogloxene FL ○ ○ ○
Pantherophis spiloides (Duméril, Bibron, & Duméril, 1854) Gray Rat Snake Squamata Trogloxene G4 LC AL, FL ○ ○ ○
Parabogidiella americana Holsinger, 1980 Amphipoda Stygobiont G2G3 TX ○ 65 4
Paracyclops canadensis (Willey, 1934) Cyclopoida Stygoxene IN ○ ○ ○
Paracyclops chiltoni (Thomson G.M., 1883) Cyclopoida Stygoxene IN, KY ○ ○ ○
Paracyclops poppei (Rehberg, 1880) Copepod Cyclopoida Stygoxene GNR KY ○ ○ ○
Paragnetina media (Walker, 1852) Embossed Stone Plecoptera Stygoxene G4 ○ ○ ○
Paraholsingerius smaragdinus (Holsinger, 1992) Amphipoda Stygobiont GNR TX ○ 17 ○
Parametriocnemus lundbeckii (Johannsen, 1905) Diptera Trogloxene GNR ○ ○ ○
Paramexiweckelia ruffoi Holsinger, 1993 Ruffo's Cave Amphipod Amphipoda Stygobiont G1G2 TX ○ 40 ○
Paramoera lokowai Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 6 ○
Paramoera paakai Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 17 ○
Paramoera rua Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 11 ○
Paraplanaria occulta (Kenk, 1969) Tricladida Stygophile VA ○ ○ ○
Pararhynchelmis texana Fend, 2023 Lumbriculida Stygobiont TX ○ 7 ○
Parascalops breweri (Bachman, 1842) Brewer's Mole Eulipotyphla Accidental G5 LC KY ○ ○ ○
Parasitus fimetorum (Berlese, 1904) Mesostigmata Trogloxene TN ○ ○ ○
Parasitus remberti (Oudemans, 1912) Mesostigmata Troglophile IA ○ ○ ○
Parasteatoda tepidariorum (Koch, 1841) Common House Spider Araneae Troglophile GNR AL, AR, GA, IL, KY, TN, VA ○ ○ ○
Paratachys aaa Liebherr, 2021 Coleoptera Troglobiont HI ○ ○ ○
Paratheridula perniciosa (Keyserling, 1886) Cobweb Spiders Araneae Troglophile ○ ○ ○
Paravachonium bolivari Beier, 1956 Pseudoscorpiones Troglophile ○ ○ ○
Paravitrea capsella (A. A. Gould, 1851) Stylommatophora Trogloxene KY, TN ○ ○ ○
Paravitrea multidentata (A. Binney, 1840) Dentate Supercoil Stylommatophora Trogloxene G5 TN ○ ○ ○
Parazaona cavicola Chamberlin, 1938 Pseudoscorpiones Troglophile ○ ○ ○
Paria sexnotata (Say, 1824) Juniper Paria Coleoptera Accidental GNR AR ○ ○ ○
Parisotoma notabilis (Schäffer, 1896) Entomobryomorpha Troglophile GNR FL, IA, IL, MN, MO, TN, WI ○ ○ ○
Parobisium charlotteae Chamberlin, 1962 Pseudoscorpiones Troglobiont G1G2 OR ○ 3 6
Parobisium petilum Harvey & Cullen, 2020 Pseudoscorpiones Troglobiont CA ○ 1 ○
Parobisium yosemite Cokendolpher & Krejca, 2010 Pseudoscorpiones Troglobiont CA ○ ○ ○
Parvidrilus strayeri Erseus, 1999 Lumbriculida Stygobiont G1G2 AL ○ 17 ○
Patera appressa (Say, 1821) Flat Bladetooth Stylommatophora Trogloxene G5 AL, GA, KY, TN, VA ○ ○ ○
Patera laevior (Pilsbry, 1940) Smooth Bladetooth Stylommatophora Trogloxene G4 KY, TN ○ ○ ○
Patera perigrapta (Pilsbry, 1894) Engraved Bladetooth Stylommatophora Trogloxene G5 AL, GA, TN ○ ○ ○
Patera sargentiana (C. W. Johnson & Pilsbry, 1892) Grand Bladetooth Stylommatophora Trogloxene G2 AL ○ ○ ○
Patrobus longicornis (Say, 1823) Long-Horned Patrobus Beetle Coleoptera Trogloxene G5 AL, IL, IN, KY, MO ○ ○ ○
Pegoplata infirma (Meigen, 1826) Diptera Trogloxene GNR ○ ○ ○
Pelegrina exigua (Banks, 1892) Araneae Accidental GNR IL ○ ○ ○
Pelegrina proterva (Walckenaer, 1837) Common White-Cheeked Jumping Spider Araneae Accidental G5 IL ○ ○ ○
Penaeus setiferus (Linnaeus, 1767) Northern White Shrimp Decapoda Stygoxene GNR FL ○ ○ ○
Percina caprodes (Rafinesque, 1818) Common Logperch Perciformes Accidental G5 LC AL, KY ○ ○ ○
Percina nigrofasciata (Agassiz, 1854) Blackbanded Darter Perciformes Accidental G5 LC ○ ○ ○
Percymoorensis lateralis (Say, 1824) Arhynchobdellida Stygoxene ○ ○ ○
Perichaeta minima Horst, 1894 Crassiclitellata Troglophile GA ○ ○ ○
Pericoma scotiae (Curran, 1924) Diptera Trogloxene TN ○ ○ ○
Perimyotis subflavus (F. Cuvier, 1832) Tricolored Bat Chiroptera Trogloxene G3 VU AL, AR, CT, FL, GA, IL, IN, KS, KY, MA, MO, MS, NC, NJ, NY, OH, OK, ON, PA, TN, TX, VA, WI, WV ○ ○ ○
Periplaneta americana (Linnaeus, 1758) American Cockroach Blattodea Trogloxene GNR ○ ○ ○
Perissopalla flagellisetula Brennan & White, 1960 Trombidiformes Trogloxene AL ○ ○ ○
Peromyscopsylla hesperomys (Baker, 1904) Siphonaptera Trogloxene GNR TN ○ ○ ○
Peromyscus attwateri J. A. Allen, 1895 Attwater's Mouse Rodentia Trogloxene G5 LC AR ○ ○ ○
Peromyscus boylii (Baird, 1855) Brush Deermouse Rodentia Trogloxene G5 LC NM ○ ○ ○
Peromyscus gossypinus (Le Conte, 1853) Cotton Deermouse Rodentia Trogloxene G5 LC FL ○ ○ ○
Peromyscus leucopus (Rafinesque, 1818) White-Footed Deermouse Rodentia Trogloxene G5 LC IN, KY, NM, TN ○ ○ ○
Peromyscus maniculatus (Wagner, 1845) Deer Mouse Rodentia Trogloxene G5 LC IL ○ ○ ○
Perpolita binneyana (E. S. Morse, 1864) Blue Glass Stylommatophora Trogloxene ○ ○ ○
Petrochelidon pyrrhonota (Vieillot, 1817) American Cliff Swallow Passeriformes Trogloxene G5 LC OK ○ ○ ○
Phagocata angusta Kenk, 1977 Tricladida Stygobiont G1 WV ○ 7 ○
Phagocata bursaperforata Darlington, 1959 Tricladida Stygophile AL ○ ○ ○
Phagocata carolinensis Kenk, 1979 Tricladida Stygobiont G1G2 NC ○ 4 ○
Phagocata gracilis (Haldeman, 1840) Slender Spring Flatworm Tricladida Stygophile G5 IL, IN, KY, MO, OH, PA, VA ○ ○ ○
Phagocata holleri Kenk, 1979 Holler's Cave Flatworm Tricladida Stygobiont G1G2 NC ○ 4 ○
Phagocata morgani (Stevens & Boring, 1906) Tricladida Stygophile G4 AL, KY, MD, NC, NY, VA, VT, WV ○ ○ ○
Phagocata notorchis Kenk, 1987 Tricladida Stygobiont TN ○ 4 ○
Phagocata oregonensis Hyman, 1963 Oregon Cave Flatworm Tricladida Stygobiont G1G2 OR ○ ○ ○
Phagocata procera Kenk, 1984 Tricladida Stygobiont G1G2 NC ○ 14 ○
Phagocata pygmaea Kenk, 1987 Tricladida Stygobiont NC ○ 2 ○
Phagocata spuria Kenk, 1987 Tricladida Stygobiont NC ○ 3 ○
Phalangodes armata Tellkampf, 1844 Opiliones Troglobiont G3G4 KY, IN ○ 15 4
Phanetta subterranea (Emerton, 1875) Subterranean Sheetweb Spider Araneae Troglobiont G5 AL, AR, GA, IL, IN, KY, MO, NC, OH, TN, VA, WV ○ 9 47
Phaonia soccata (Walker, 1849) Diptera Accidental GNR ○ ○ ○
Phelister fulvulus Marseul, 1870 Coleoptera Trogloxene ○ ○ ○
Phelister luculentus Bickhardt, 1917 Escarabajo Payaso Coleoptera Trogloxene ○ ○ ○
Phenacomys intermedius Merriam, 1889 Heather Vole Rodentia Trogloxene G5 LC ○ ○ ○
Pheretima rodericensis (Grube, 1879) Crassiclitellata Troglophile ○ ○ ○
Phidippus purpuratus Keyserling, 1885 Marbled Purple Jumper Araneae Accidental G5 ○ ○ ○
Philomycus carolinianus (Bosc, 1802) Carolina Mantleslug Stylommatophora Trogloxene G5 FL ○ ○ ○
Philomycus togatus (A. A. Gould, 1841) Toga Mantleslug Stylommatophora Trogloxene G5 IL, KY ○ ○ ○
Philonthus cyanipennis (Fabricius, 1792) Staph Beetle Coleoptera Trogloxene GA ○ ○ ○
Philonthus lomatus Erichson, 1840 Coleoptera Trogloxene G5 IN ○ ○ ○
Phoberus capensis (Scholtz, 1979) Coleoptera Trogloxene ○ ○ ○
Phoberus pusillus (Péringuey, 1908) Coleoptera Trogloxene ○ ○ ○
Phoberus sulcatus (Thunberg, 1787) Grooveneck Keratin Beetle Coleoptera Trogloxene ○ ○ ○
Pholcus dade Huber, 2011 Araneae Troglophile GA ○ ○ ○
Pholcus jusahi Huber, 2011 Araneae Troglophile VA ○ ○ ○
Pholcus lanieri Huber, 2011 Araneae Troglophile GA ○ ○ ○
Pholcus phalangioides (Fuesslin, 1775) Cosmopolitan Cellar Spider Araneae Troglophile GNR IL, KY, NM, TN ○ ○ ○
Phoxinus erythrogaster (Rafinesque, 1820) Cypriniformes Stygoxene KY ○ ○ ○
Phreatoceras taylori (Hershler & Longley, 1986) Nymph trumpet Littorinimorpha Stygobiont G1G2 DD TX ○ 20 5
Phreatodrobia bulla Perez & Castañeda, 2023 Littorinimorpha Stygobiont TX ○ ○ 4
Phreatodrobia conica Hershler & Longley, 1986 Hueco Cavesnail Littorinimorpha Stygobiont G1 TX ○ 27 7
Phreatodrobia coronae Hershler, 1987 Littorinimorpha Stygobiont G1G2 DD TX ○ 9 ○
Phreatodrobia embossa Perez et al., 2025 Littorinimorpha Stygobiont TX ○ ○ ○
Phreatodrobia imitata Hershler & Longley, 1986 Mimic cavesnail Littorinimorpha Stygobiont G1 VU TX ○ 42 2
Phreatodrobia micra (Pilsbry & Ferriss, 1906) Flattened Cavesnail Littorinimorpha Stygobiont G2G3 TX, AL ○ 573 10
Phreatodrobia nugax (Pilsbry & Ferriss, 1906) Domed Cavesnail Littorinimorpha Stygobiont G1G2 DD TX ○ 103 13
Phreatodrobia plana Hershler & Longley, 1986 Disc cavesnail Littorinimorpha Stygobiont G2 TX ○ 7 3
Phreatodrobia punctata Hershler & Longley, 1986 High-hat cavesnail Littorinimorpha Stygobiont G2 TX ○ 12 3
Phreatodrobia rotunda Hershler & Longley, 1986 Beaked cavesnail Littorinimorpha Stygobiont G1G2 DD TX ○ 9 8
Phreatodrobia spica Perez & Alvear, 2020 Spiky Cavesnail Littorinimorpha Stygobiont GNR TX ○ 7 1
Phronia exigua (Zetterstedt, 1852) Diptera Troglophile GNR ○ ○ ○
Phronia flavipes Winnertz, 1864 Diptera Troglophile GNR ○ ○ ○
Phronia incerta (Adams, 1907) Diptera Troglophile IN ○ ○ ○
Physella acuta (Draparnaud, 1805) European Physa Basommatophora Stygophile IL, MO ○ ○ ○
Physella gyrina (Say, 1821) Tadpole Physa Basommatophora Stygophile G5 LC GA ○ ○ ○
Physella spelunca Turner & Clench, 1974 Cave Physa Basommatophora Stygobiont G1 VU WY ○ 11 4
Physocyclus enaulus Crosby, 1926 Araneae Troglophile NM ○ ○ ○
Physocypria denticulata (Daday, 1905) Podocopida Stygophile TX ○ ○ ○
Physocypria globula Furtos, 1933 Podocopida Stygophile TX ○ ○ ○
Pimephales promelas Rafinesque, 1820 Fathead Minnow (Rosy Reds) Cypriniformes Stygoxene G5 LC KY ○ ○ ○
Pimoa altioculata (Keyserling, 1886) Araneae Accidental GNR ○ ○ ○
Pinodytes pusio Horn, 1892 Coleoptera Troglophile CA ○ ○ ○
Pinodytes rothi (Hatch, 1957) Coleoptera Troglophile OR ○ ○ ○
Pirata alachuus Gertsch & Wallace, 1935 Araneae Accidental GNR GA ○ ○ ○
Pirata sedentarius Montgomery, 1904 Araneae Trogloxene GNR IL, TN ○ ○ ○
Piratula insularis (Emerton, 1885) Lonely Wolf Spider Araneae Accidental G5 GA ○ ○ ○
Pisaurina mira (Walckenaer, 1837) Nursery Web Spider Araneae Accidental G5 AL, IL ○ ○ ○
Plathemis lydia (Drury, 1773) Common Whitetail or Long-Tailed Skimmer Odonata Accidental G5 LC MO ○ ○ ○
Platynus angustatus Dejean, 1828 Coleoptera Trogloxene GNR KY ○ ○ ○
Platynus cincticollis (Say, 1823) Coleoptera Troglophile GNR IN ○ ○ ○
Platynus parmarginatus Hamilton, 1893 Coleoptera Trogloxene GNR GA ○ ○ ○
Platynus tenuicollis (LeConte, 1846) Slender Ground Beetle Coleoptera Troglophile G5 AL, AR, IL, IN, KY, MO, TN, VA ○ ○ ○
Plestiodon fasciatus (Linnaeus, 1758) Common Five-Lined Skink Squamata Accidental G5 LC AR, KY ○ ○ ○
Plethodon albagula Grobman, 1944 Western Slimy Salamander Caudata Troglophile G5 LC AR, OK ○ ○ ○
Plethodon angusticlavius Grobman, 1944 Ozark Salamander Caudata Troglophile G4 LC AR, OK ○ ○ ○
Plethodon caddoensis Pope & Pope, 1951 Caddo Mountain Salamander Caudata Trogloxene G2 NT ○ ○ ○
Plethodon cinereus (Green, 1818) Eastern Red-Backed Salamander Caudata Trogloxene G5 LC IN, OH ○ ○ ○
Plethodon dorsalis Cope, 1889 Northern Zigzag Salamander Caudata Troglophile G5 LC IL, IN, KY, TN ○ ○ ○
Plethodon glutinosus (Green, 1818) Northern Slimy Salamander Caudata Troglophile G5 LC AL, GA, IL, IN, KY, OH, TN, VA ○ ○ ○
Plethodon grobmani Allen & Neill, 1949 Southeastern Slimy Salamander Caudata Troglophile G5 FL ○ ○ ○
Plethodon petraeus Wynn, Highton & Jacobs, 1988 Pigeon Mountain Salamander Caudata Troglophile G2 NT GA ○ ○ ○
Plethodon richmondi Netting & Mittleman, 1938 Ravine Salamander Caudata Trogloxene G5 LC KY, WV ○ ○ ○
Plethodon serratus Grobman, 1944 Southern Red-Backed Salamander Caudata Accidental G5 LC AR, GA ○ ○ ○
Plethodon ventralis Highton, 1997 Southern Zigzag Salamander Caudata Troglophile G4 LC AL, GA, TN ○ ○ ○
Plethodon websteri Highton, 1979 Webster's Salamander Caudata Trogloxene G3 LC AL ○ ○ ○
Plumatyla humerosa (Loomis, 1943) Chordeumatida Troglobiont G2G3 CA ○ 2 ○
Plutomurus californicus (Folson, 1913) Entomobryomorpha Troglobiont G2G3 CA, Mexico ○ 13 ○
Plutomurus grahami (Christiansen, 1980) Entomobryomorpha Troglobiont G1 NM ○ 2 ○
Pnyxia scabiei (Hopkins, 1895) Potato Scab Gnat Diptera Trogloxene KY ○ ○ ○
Poecilochirus carabi Canestrini & Canestrini, 1882 Mesostigmata Trogloxene TN ○ ○ ○
Poecilophysis extraneostella Zacharda 1985 Actinedida Troglobiont G2 VA, WV ○ ○ ○
Poecilophysis pratensis (C. L. Koch, 1835) Mite Actinedida Troglophile WV ○ ○ ○
Poecilophysis weyerensis (Packard, 1888) Actinedida Troglobiont G3 KY, TN, VA, WV, IN ○ ○ ○
Pogonognathellus bidentatus (Folsom, JW, 1913) Entomobryomorpha Troglophile GNR AL, GA, IL, KY, TN, VA ○ ○ ○
Pogonognathellus danieli Felderhoff, KL, Bernard, EC & Moulton, JK, 2010 Entomobryomorpha Troglophile VA ○ ○ ○
Pogonognathellus dubius (Christiansen, 1964) Ghost Silver Bullet Springtail Entomobryomorpha Troglophile GNR AL, GA, KY, TN ○ ○ ○
Pogonognathellus flavescens Dethier & Hubart, 2000 Entomobryomorpha Troglophile AL, GA, IA, IL, IN, KY, MO, WI ○ ○ ○
Pogonognathellus nigritus (Maynard, EA, 1951) Entomobryomorpha Troglophile IL, TN ○ ○ ○
Polycentropus barri Ross & Yamamoto Trichoptera Stygoxene G3 KY ○ ○ ○
Polycentropus elarus Ross, 1944 Trichoptera Stygoxene G3 KY ○ ○ ○
Polydesmus inconstans Latzel, 1884 European Sculpted Millipede Polydesmida Accidental G4 IL, PA ○ ○ ○
Polydrusus impressifrons Gyllenhal, 1834 Pale Green Weevil Coleoptera Accidental GNR ○ ○ ○
Polygonia comma (Harris, 1852) Eastern Comma Lepidoptera Accidental G5 KY ○ ○ ○
Polynoncus gemmingeri (Harold, 1872) Coleoptera Trogloxene ○ ○ ○
Polysphondylium violaceum Bref., 1884 Dictyosteliales Stygophile GA ○ ○ ○
Pomatiopsis lapidaria (Say, 1817) Slender Walker Littorinimorpha Stygoxene G5 LC IL, TN ○ ○ ○
Pomoxis nigromaculatus (Lesueur, 1829) Black Crappie Centrarchiformes Stygoxene G5 LC FL ○ ○ ○
Ponera pennsylvanica Buckley, 1866 Common Porthole Ant Hymenoptera Accidental G5 KY ○ ○ ○
Porcellio laevis Latreille, 1804 Smooth Slater Isopoda Trogloxene TN ○ ○ ○
Porcellio scaber Latreille, 1804 Common Rough Woodlouse Isopoda Trogloxene GNR IN ○ ○ ○
Porcellio spinicornis Say, 1818 Painted Woodlouse Isopoda Trogloxene GNR ○ ○ ○
Porrhomma cavernicola (Keyserling, 1886) Appalachian Cave Spider Araneae Troglobiont G5 AL, AR, GA, IA, IL, IN, KY, MD, MO, NC, OK, ON, PA, TN, VA, WV ○ 14 ○
Porrhomma egeria Simon, 1884 Kelderkleinoogje Araneae Troglophile ○ ○ ○
Porrhomma terrestre (Emerton, 1882) Terrestrial Wide-Eyed Sheetweaver Araneae Troglophile GNR IL ○ ○ ○
Potamocypris fulva (Brady, 1868) Podocopida Stygophile GA ○ ○ ○
Potamothrix bavaricus (Oschmann, 1913) Aquatic Oligochaete Worm Tubificida Stygophile TX ○ ○ ○
Pratherodesmus despaini Shear, 2009 Polydesmida Troglobiont CA ○ 1 ○
Pratherodesmus ecclesia Shear, 2009 Polydesmida Troglobiont AZ ○ 1 ○
Pratherodesmus voylesi Shear, 2009 Polydesmida Troglobiont AZ ○ 1 ○
Prenolepis imparis (Say, 1836) American Winter Ant Hymenoptera Accidental G5 AL ○ ○ ○
Prietella phreatophila Carranza, 1954 Mexican Blindcat Siluriformes Stygobiont GNR VU ESA TX, Mexico ○ 142 8
Primicimex cavernis Barber, 1941 Hemiptera Trogloxene TX ○ ○ ○
Prionochaeta opaca (Say, 1825) Small Carrion Beetle Coleoptera Troglophile G5 AL, FL, GA, IL, IN, KY, MO, NC, TN ○ ○ ○
Pristiloma subrupicola (Dall, 1877) Southern Tightcoil Stylommatophora Trogloxene ○ ○ ○
Pristina leidyi Smith, 1896 Aquatic Oligochaete Worm Tubificida Stygophile GNR SC ○ ○ ○
Procambarus acherontis (Lonnberg, 1894) Orlando Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 18 5
Procambarus alleni (Faxon, 1884) Everglades Crayfish Decapoda Stygoxene G4 LC FL ○ ○ ○
Procambarus attiguus Hobbs & Franz, 1992 Silver Glen Springs Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 8 5
Procambarus blandingii (Harlan, 1830) Santee Crayfish Decapoda Stygoxene G4 LC ○ ○ ○
Procambarus clarkii (Girard, 1852) Red Swamp Crayfish Decapoda Stygoxene G5 LC TX ○ ○ ○
Procambarus cuevachicae (Hobbs, 1941) Decapoda Stygoxene LC ○ ○ ○
Procambarus delicatus Hobbs & Franz, 1986 Bigcheek Cave Crayfish Decapoda Stygobiont G1 CR FL ○ 3 2
Procambarus erythrops Relyea & Sutton, 1975 Santa Fe Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 27 6
Procambarus fallax (Hagen, 1870) Slough Crayfish Decapoda Stygoxene G5 LC FL ○ ○ ○
Procambarus franzi Hobbs & Lee, 1976 Orange Lake Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 13 5
Procambarus horsti Hobbs & Means, 1972 Big Blue Spring Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 10 5
Procambarus leitheuseri Franz & Hobbs, 1983 Coastal Lowland Cave Crayfish Decapoda Stygobiont G1G2 EN FL ○ 12 5
Procambarus lucifugus (Hobbs, 1940) Light-fleeing Cave Crayfish Decapoda Stygobiont G1G2 LC FL ○ 89 10
Procambarus milleri Hobbs, 1971 Miami Cave Crayfish Decapoda Stygobiont G1 EN ESA? FL ○ 14 6
Procambarus morrisi Hobbs & Franz, 1991 Putnum County Cave Crayfish Decapoda Stygobiont G1 CR FL ○ 13 5
Procambarus orcinus Hobbs & Means, 1972 Woodville Karst Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 25 6
Procambarus paeninsulanus (Faxon, 1914) Peninsula Crayfish Decapoda Stygoxene G5 LC FL ○ ○ ○
Procambarus pallidus (Hobbs, 1940) Pallid Cave Crayfish Decapoda Stygobiont G1G2 NT FL, GA ○ 84 5
Procambarus pilosimanus (Ortmann, 1906) Decapoda Stygoxene LC ○ ○ ○
Procambarus spiculifer (LeConte, 1856) White Tubercled Crayfish Decapoda Stygoxene G5 LC FL ○ ○ ○
Procambarus toltecae Hobbs, 1943 Toltecae Crayfish Decapoda Stygoxene LC ○ ○ ○
Procambarus versutus (Hagen, 1870) Sly Crayfish Decapoda Stygoxene G5 LC FL ○ ○ ○
Procambarus villalobosi Hobbs, 1969 Decapoda Stygoxene DD ○ ○ ○
Procaris hawaiana Holthuis, 1973 Decapoda Stygobiont G1 ESA HI ○ 14 4
Proceratium micrommatum (Roger, 1863) Hymenoptera Accidental ○ ○ ○
Procladius bellus (Loew, 1866) Diptera Trogloxene GNR GA ○ ○ ○
Procyon lotor (Linnaeus, 1758) Common Raccoon Carnivora Trogloxene G5 LC AL, AR, FL, GA, IL, IN, KY, OK, TN, VA ○ ○ ○
Progomphus obscurus (Rambur, 1842) Common Sanddragon Odonata Accidental G5 LC GA ○ ○ ○
Progradungula carraiensis Forster & Gray, 1979 Carrai Cave Spider Araneae Troglophile ○ ○ ○
Proisotoma frisoni Folsom, JW, 1937 Entomobryomorpha Accidental IA ○ ○ ○
Proisotoma minuta (Tullberg, 1871) Springtail Entomobryomorpha Troglophile GNR MN ○ ○ ○
Proisotoma sepulcralis (Folsom, JW, 1902) Entomobryomorpha Trogloxene IL ○ ○ ○
Proisotoma veca (Wray, 1952) Entomobryomorpha Trogloxene IA ○ ○ ○
Proisotoma vesiculata Folsom, 1937 Entomobryomorpha Trogloxene IA ○ ○ ○
Prosimulium mixtum Syme & Davies, 1958 Mixed-Up Black Fly Diptera Trogloxene G5 ○ ○ ○
Prosimulium saltus Stone & Jamnback, 1955 Diptera Trogloxene GA ○ ○ ○
Prostoma graecense (Böhmig, 1892) Monostilifera Stygoxene GA ○ ○ ○
Proteocephalus poulsoni Whitaker & Zober, 1978 Proteocephalidea Stygobiont KY ○ ○ ○
Protolimnesia ventriplacophora Smith & Cook, 1996 Trombidiformes Stygobiont TX ○ ○ ○
Psephenus herricki (DeKay, 1844) Herrick's Water Penny Beetle Coleoptera Stygoxene G5 KY ○ ○ ○
Pseudachorutes aureofasciatus (Mac Gillivray, 1893) Poduromorpha Trogloxene GNR IA, IL ○ ○ ○
Pseudachorutes saxatilis Macnamara, 1920 Poduromorpha Trogloxene GNR IL ○ ○ ○
Pseudachorutes subcrassoides Mills, HB, 1934 Poduromorpha Trogloxene IA ○ ○ ○
Pseudacris crucifer (Wied-Neuwied, 1838) Spring Peeper Anura Accidental G5 LC AL, AR, FL, GA, OK ○ ○ ○
Pseudacris feriarum (Baird, 1854) Southeastern Chorus Frog Anura Accidental G5 LC AL, GA ○ ○ ○
Pseudacris triseriata (Wied-Neuwied, 1838) Striped Chorus Frog Anura Accidental G5 LC IL ○ ○ ○
Pseudanophthalmus abditus Krekeler, 1973 Concealed Cave Beetle Coleoptera Troglobiont G3 KY ○ ○ ○
Pseudanophthalmus acherontis Barr, 1959 Echo Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus alabamae Valentine, 1932 Alabama Cave Beetle Coleoptera Troglobiont G2 AL ○ 8 ○
Pseudanophthalmus alladini Valentine, 1945 Alladin's Cave Beetle Coleoptera Troglobiont G3G4 AL ○ ○ ○
Pseudanophthalmus assimilis Barr, 1981 West Wills Valley Cave Beetle Coleoptera Troglobiont G1G2 AL ○ ○ ○
Pseudanophthalmus audax (Horn, 1883) Bold Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 1 ○
Pseudanophthalmus avernus (Valentine, 1945) Avernus Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus barberi Jeannel, 1931 Coleoptera Troglobiont G2G3 KY ○ 2 3
Pseudanophthalmus barri Krekeler, 1973 Coleoptera Troglobiont G1G2 IN ○ 2 ○
Pseudanophthalmus beakleyi Valentine, 1937 Beakley's Cave Beetle Coleoptera Troglobiont G3G4 KY, TN ○ ○ ○
Pseudanophthalmus bendermani Barr, 1959 Benderman's Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Pseudanophthalmus caecus Krekeler, 1973 Clifton Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus calcareus Barr, 1981 Limestone Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus catherinae Barr, 1959 Catherine's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus catoryctos Krekeler, 1973 Lesser Adams Cave Beetle Coleoptera Troglobiont G1 KY ○ 3 ○
Pseudanophthalmus cerberus Barr, 1985 Coleoptera Troglobiont G3G4 KY, TN ○ 27 ○
Pseudanophthalmus chthonius Krekeler, 1973 Coleoptera Troglobiont G3 IN, KY ○ 2 ○
Pseudanophthalmus ciliaris Valentine, 1937 Coleoptera Troglobiont G3G4 KY, TN ○ 5 15
Pseudanophthalmus cnephosus Krekeler, 1973 Larue County Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus colemanensis Barr, 1959 Coleman Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ 7
Pseudanophthalmus conditus Krekeler, 1973 Hidden Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus cordicollis Barr, 1981 Little Kennedy Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus cumberlandus Valentine, 1937 Coleoptera Troglobiont G1G2 TN ○ 1 ○
Pseudanophthalmus darlingtoni Barr, 1985 Coleoptera Troglobiont G3 KY ○ ○ ○
Pseudanophthalmus deceptivus Barr, 1981 Deceptive Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus delicatus Valentine, 1932 Delicate Cave Beetle Coleoptera Troglobiont G3G4 VA, TN ○ ○ ○
Pseudanophthalmus desertus Krekeler, 1973 Coleoptera Troglobiont G3 KY ○ 4 ○
Pseudanophthalmus digitus Valentine, 1932 Coleoptera Troglobiont G1G2 GA, TN ○ 1 ○
Pseudanophthalmus distinguens Valentine, 1948 Coleoptera Troglobiont G1G2 AL, VA ○ ○ ○
Pseudanophthalmus egberti Barr, 1965 New River Valley Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudanophthalmus elongatus Krekeler, 1973 Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus emersoni Krekeler, 1958 Coleoptera Troglobiont G1 IN ○ 2 ○
Pseudanophthalmus englehardti (Barber, 1928) Englehardt's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus eremita (Horn, 1871) Coleoptera Troglobiont G1G2 IN, KY, OH ○ 2 ○
Pseudanophthalmus exiguus Krekeler, 1973 Coleoptera Troglobiont G3G4 KY ○ 4 ○
Pseudanophthalmus exoticus Krekeler, 1973 Exotic Cave Beetle Coleoptera Troglobiont G1 KY ○ 1 ○
Pseudanophthalmus farrelli Barr, 1959 Coleoptera Troglobiont G1G2 TN ○ 2 ○
Pseudanophthalmus fastigatus Barr, 1981 Tapered Cave Beetle Coleoptera Troglobiont G1 GA ○ ○ ○
Pseudanophthalmus fluviatilis Valentine, 1948 Coleoptera Troglobiont G3 AL ○ ○ ○
Pseudanophthalmus fowlerae Barr, 1980 Fowler's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus frigidus Barr, 1981 Icebox Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus fulleri Valentine, 1932 Coleoptera Troglobiont G2G3 GA, TN, AL ○ 1 ○
Pseudanophthalmus fuscus Valentine, 1931 Coleoptera Troglobiont G4 WV ○ 5 ○
Pseudanophthalmus georgiae Barr, 1981 Georgia Cave Beetle Coleoptera Troglobiont G1G2 GA ○ ○ ○
Pseudanophthalmus globiceps Barr, 1985 Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus gracilis Valentine, 1931 Clover Hollow Cave Beetle Coleoptera Troglobiont G1G2 VA ○ 1 ○
Pseudanophthalmus grandis Valentine, 1931 Coleoptera Troglobiont G4 WV ○ 9 ○
Pseudanophthalmus hadenoecus Barr, 1965 Timber Ridge Cave Beetle Coleoptera Troglobiont G1 WV ○ 1 ○
Pseudanophthalmus henroti Jeannel, 1949 Coleoptera Troglobiont GNR WV ○ ○ ○
Pseudanophthalmus hesperus Barr, 1959 Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus higginbothami Valentine, 1931 Coleoptera Troglobiont G2 WV ○ 2 ○
Pseudanophthalmus hirsutus Valentine, 1931 Cumberland Gap Cave Beetle Coleoptera Troglobiont G1G2 TN, VA ○ 3 ○
Pseudanophthalmus hoffmani Barr, 1965 Hoffman's Cave Beetle Coleoptera Troglobiont G2G3 VA ○ 1 ○
Pseudanophthalmus holsingeri Barr, 1965 Holsinger's Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudanophthalmus horni (Garman, 1892) Garman's Cave Beetle Coleoptera Troglobiont G3 KY ○ 8 ○
Pseudanophthalmus hortulanus Barr, 1965 Garden Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudanophthalmus hubbardi Barber, 1928 Hubbard's Cave Beetle Coleoptera Troglobiont G1 VA ○ 5 ○
Pseudanophthalmus hubrichti Valentine, 1948 Hubricht's Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus humeralis Valentine, 1931 Coleoptera Troglobiont G2 TN ○ 2 ○
Pseudanophthalmus hypertrichosis Valentine, 1932 Coleoptera Troglobiont G3 WV ○ 2 ○
Pseudanophthalmus hypolithos Barr, 1981 Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus illinoisensis Barr & Peck, 1966 Illinois Cave Beetle Coleoptera Troglobiont G1 IL ○ 1 ○
Pseudanophthalmus inexpectatus Barr, 1959 Surprising Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 5 ○
Pseudanophthalmus inquisitor Barr, 1980 Inquirer Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus insularis Barr, 1959 Baker Station Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus intermedius (Valentine, 1931) Intermediate Cave Beetle Coleoptera Troglobiont G2 TN ○ 2 ○
Pseudanophthalmus intersectus Barr, 1965 Crossroads Cave Beetle Coleoptera Troglobiont G1G2 VA ○ ○ ○
Pseudanophthalmus jonesi Valentine, 1945 Grassy Cove Cave Beetle Coleoptera Troglobiont G1G2 TN, AL ○ 3 ○
Pseudanophthalmus krameri Krekeler, 1973 Kramer's Cave Beetle Coleoptera Troglobiont G1 OH ○ 2 ○
Pseudanophthalmus krekeleri Barr, 1965 Rich Mountain Cave Beetle Coleoptera Troglobiont GX WV ○ 1 ○
Pseudanophthalmus lallemanti Jeannel, 1949 Coleoptera Troglobiont G1 WV ○ ○ ○
Pseudanophthalmus leonae Barr, 1960 Coleoptera Troglobiont G1 IN ○ ○ ○
Pseudanophthalmus limicola Jeannel, 1931 Maddens Cave Beetle Coleoptera Troglobiont G1G2 VA ○ 1 ○
Pseudanophthalmus lodingi Valentine, 1931 Coleoptera Troglobiont G1G2 AL ○ 6 ○
Pseudanophthalmus loganensis Barr, 1959 Coleoptera Troglobiont G2G3 KY, TN ○ ○ 10
Pseudanophthalmus longiceps Barr, 1981 Long-headed Cave Beetle Coleoptera Troglobiont G1G2 TN, VA ○ ○ ○
Pseudanophthalmus macradyi Valentine, 1948 Macrady's Cave Beetle Coleoptera Troglobiont G1G2 TN ○ 1 ○
Pseudanophthalmus major Krekeler, 1973 Beaver Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus menetriesi (Motschulsky, 1862) Coleoptera Troglobiont G3 KY ○ ○ ○
Pseudanophthalmus meridionalis Valentine, 1945 Coleoptera Troglobiont G2 AL ○ ○ ○
Pseudanophthalmus montanus Barr, 1965 Dry Fork Valley Cave Beetle Coleoptera Troglobiont G1 WV ○ 1 ○
Pseudanophthalmus nelsoni Barr, 1965 Nelson's Cave Beetle Coleoptera Troglobiont G2 VA ○ 1 ○
Pseudanophthalmus nickajackensis Barr, 1981 Nickajack Cave Beetle Coleoptera Troglobiont G1 AL, TN ○ ○ ○
Pseudanophthalmus nortoni Barr, 1981 Norton's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus occidentalis Barr, 1959 Western Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Pseudanophthalmus ohioensis Krekeler, 1973 Ohio Cave Beetle Coleoptera Troglobiont G1 OH ○ 2 ○
Pseudanophthalmus orientalis Krekeler, 1973 Surprising Cave Beetle Coleoptera Troglobiont G2G3 KY ○ ○ ○
Pseudanophthalmus orthosulcatus Valentine, 1932 Coleoptera Troglobiont G1 WV ○ 1 ○
Pseudanophthalmus packardi Barr, 1959 Packard's Cave Beetle Coleoptera Troglobiont G1G2 KY ○ ○ ○
Pseudanophthalmus pallidus Barr, 1981 Pale Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Pseudanophthalmus paradoxus Barr, 1981 Ridetop Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus parvicollis Jeannel, 1931 Hupps Hill Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus parvus Krekeler, 1973 Tatum Cave Beetle Coleoptera Troglobiont GH KY ○ 2 ○
Pseudanophthalmus paulus Barr, 1981 Nobletts Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus paynei Barr, 1981 Payne's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus petrunkevitchi Valentine, 1945 Petrunkevitch's Cave Beetle Coleoptera Troglobiont G1G2 VA ○ 1 ○
Pseudanophthalmus pholeter Krekeler, 1973 Greater Adams Cave Beetle Coleoptera Troglobiont G1 KY ○ 2 ○
Pseudanophthalmus pilosus Barr, 1985 Coleoptera Troglobiont G2G3 KY ○ ○ ○
Pseudanophthalmus pontis Barr, 1965 Natural Bridge Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudanophthalmus potamaca Valentine, 1932 South Branch Valley Cave Beetle Coleoptera Troglobiont G3G4 VA, WV ○ ○ ○
Pseudanophthalmus praetermissus Barr, 1981 Overlooked Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus princeps Barr, 1979 Coleoptera Troglobiont G1G2 KY, TN ○ 3 4
Pseudanophthalmus productus Barr, 1980 Coleoptera Troglobiont G3G4 TN ○ ○ ○
Pseudanophthalmus profundus Valentine, 1945 Coleoptera Troglobiont G2 AL ○ ○ ○
Pseudanophthalmus pubescens (Horn, 1868) Coleoptera Troglobiont G3 KY ○ 24 6
Pseudanophthalmus punctatus Valentine, 1931 Spotted Cave Beetle Coleoptera Troglobiont G2 VA ○ ○ ○
Pseudanophthalmus pusillus Barr, 1981 Tiny Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus pusio (Horn, 1868) Ellet Valley Cave Beetle Coleoptera Troglobiont G2G3 VA ○ 5 ○
Pseudanophthalmus puteanus Krekeler, 1973 Old Well Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 1 ○
Pseudanophthalmus quadratus Barr, 1965 Straley's Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudanophthalmus rittmani Krekeler, 1973 Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus robustus Valentine, 1931 Robust Cave Beetle Coleoptera Troglobiont G3G4 TN ○ 7 4
Pseudanophthalmus rogersae Barr, 1981 Roger's Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus rotundatus Valentine, 1932 Rotund Cave Beetle Coleoptera Troglobiont G2 TN, VA ○ ○ ○
Pseudanophthalmus sanctipauli Barr, 1981 Saint Paul Cave Beetle Coleoptera Troglobiont G1G2 VA ○ ○ ○
Pseudanophthalmus scholasticus Barr, 1981 Schoolhouse Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus scutilis Barr, 1981 Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus seclusus Barr, 1981 Rye Cove Cave Beetle Coleoptera Troglobiont G2G3 VA ○ ○ ○
Pseudanophthalmus senecae Valentine, 1932 Seneca Cave Beetle Coleoptera Troglobiont G1 WV ○ 1 ○
Pseudanophthalmus sequoyah Barr, 1981 Sequoyah Cave Beetle Coleoptera Troglobiont G1 AL ○ ○ ○
Pseudanophthalmus sericus Barr, 1981 Silken Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus shilohensis Krekeler, 1958 Shiloh Cave Beetle Coleoptera Troglobiont G2 IN ○ 2 ○
Pseudanophthalmus sidus Barr, 1965 Meredith Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus simplex Barr, 1980 Simple Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Pseudanophthalmus simulans Barr, 1985 Cub Run Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus solivagus Krekeler, 1973 Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus steevesi Barr, 1981 Steeves' Cave Beetle Coleoptera Troglobiont G1G2 AL ○ ○ ○
Pseudanophthalmus striatus (Motschulsky, 1862) Coleoptera Troglobiont G2G3 KY ○ 5 ○
Pseudanophthalmus stricticollis Jeannel, 1931 Marengo Cave Beetle Coleoptera Troglobiont G4 IN, IL ○ 5 ○
Pseudanophthalmus sylvaticus Barr, 1967 Coleoptera Troglophile GNR WV ○ ○ ○
Pseudanophthalmus templetoni Valentine, 1948 Templeton's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus tenebrosus Krekeler, 1973 Stevens Creek Cave Beetle Coleoptera Troglobiont G1 KY ○ 2 ○
Pseudanophthalmus tennesseensis Valentine, 1937 Coleoptera Troglobiont G3 TN ○ 1 ○
Pseudanophthalmus tenuis (Horn, 1871) Coleoptera Troglobiont G4 IN ○ 12 ○
Pseudanophthalmus thomasi Barr, 1981 Thomas' Cave Beetle Coleoptera Troglobiont G1G2 VA ○ ○ ○
Pseudanophthalmus tiresias Barr, 1959 Indian Grave Point Cave Beetle Coleoptera Troglobiont G1 TN ○ 1 ○
Pseudanophthalmus transfluvialis Barr, 1985 Coleoptera Troglobiont G1G2 KY ○ ○ ○
Pseudanophthalmus troglodytes Krekeler, 1973 Louisville Cave Beetle Coleoptera Troglobiont G1 KY ○ 2 ○
Pseudanophthalmus tullahoma Barr, 1959 Duck River Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Pseudanophthalmus umbratilis Krekeler, 1973 Coleoptera Troglobiont G2G3 KY ○ 2 ○
Pseudanophthalmus unionis Barr, 1981 Union County Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Pseudanophthalmus valentinei Jeannel, 1949 Valentine's Cave Beetle Coleoptera Troglobiont G3G4 TN ○ ○ ○
Pseudanophthalmus vanburenensis Barr, 1962 Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus ventus Barr, 1981 Blowing Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus vicarius Barr, 1965 Vicariant Cave Beetle Coleoptera Troglobiont G2G3 VA ○ 1 ○
Pseudanophthalmus virginicus (Barr, 1960) Maiden Spring Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus wallacei Barr, 1981 Wallace's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ 2
Pseudanophthalmus youngi Krekeler, 1958 Young's Cave Beetle Coleoptera Troglobiont G3G4 IN ○ 3 ○
Pseudemys peninsularis Carr, 1938 Peninsular Cooter Testudines Accidental G5 LC FL ○ ○ ○
Pseudobaicalasellus bowmani (Lewis, 1980) Natural Bridge Cave Isopod Isopoda Stygobiont G1 VA ○ ○ ○
Pseudobaicalasellus henroti (Bresson, 1955) Henrot's Cave Isopod Isopoda Stygobiont G2 VA ○ ○ ○
Pseudobaicalasellus holsingeri (Steeves, 1963) Holsinger's Cave Isopod Isopoda Stygobiont G5 MD, VA, WV ○ ○ ○
Pseudobaicalasellus incurvus (Steeves & Holsinger, 1968) Incurved Cave Isopod Isopoda Stygobiont G2G4 TN, VA ○ ○ ○
Pseudobaicalasellus magniezi Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ ○
Pseudobaicalasellus mausi (Lewis, 2009) Isopoda Stygobiont G2 MD, VA ○ ○ ○
Pseudobaicalasellus vandeli (Bresson, 1955) Vandel's Cave Isopod Isopoda Stygobiont G3G4 MD, VA ○ ○ ○
Pseudocandona albicans (Brady, 1864) Podocopida Stygophile GNR TX ○ ○ ○
Pseudocandona jeanneli (Klie, 1931) Jeannel's Cave Ostracod Podocopida Stygobiont G3 IN, KY, TN ○ 3 ○
Pseudocandona marengoensis (Klie, 1931) Marengo Cave Ostracod Podocopida Stygobiont G1G2 IN ○ ○ ○
Pseudodifflugia gracilis Schlumb. Gromiida Stygoxene AL ○ ○ ○
Pseudogarypus hypogeus Muchmore, 1981 Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Pseudogarypus orpheus Muchmore, 1981 Music Hall Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Pseudogarypus speleus Benedict & Malcolm, 1978 Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Pseudoleria parvitarsus Garrett, 1925 Diptera Troglophile GNR NM ○ ○ ○
Pseudopolydesmus erasus (Loomis, 1943) Polydesmida Trogloxene TN ○ ○ ○
Pseudopolydesmus pinetorum (Bollman, 1888) Polydesmida Trogloxene GNR AL, AR, GA, IL ○ ○ ○
Pseudopolydesmus serratus (Say, 1821) Common Pink Flat-Back Polydesmida Trogloxene G5 AL, IN, PA, TN ○ ○ ○
Pseudosinella aera Christiansen & Bellinger, 1980 Entomobryomorpha Troglophile G4 IL, TN ○ ○ ○
Pseudosinella alba (Packard, AS, 1873) Entomobryomorpha Troglophile GNR MN, TN ○ ○ ○
Pseudosinella argentea (Loksa, 1960) Entomobryomorpha Troglophile GNR AL, AR, IL, KY, MO, TN ○ ○ ○
Pseudosinella bona Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G2 VA ○ 1 ○
Pseudosinella boneti Bagnall, 1941 Entomobryomorpha Troglophile KY, TN ○ ○ ○
Pseudosinella certa Christiansen & Bellinger, 1980 Gandy Creek Cave Springtail Entomobryomorpha Troglobiont G1 WV ○ 1 ○
Pseudosinella christianseni Salmon, 1965 Christiansen's Cave Springtail Entomobryomorpha Troglobiont G5 AL, GA, KY, TN, VA ○ ○ ○
Pseudosinella collina Wray, 1952 Entomobryomorpha Troglophile GNR AL, TN ○ ○ ○
Pseudosinella dubia Christiansen, 1961 Entomobryomorpha Troglobiont G3 AR, OK, Mexico ○ 10 ○
Pseudosinella duodecimpunctata Denis, J-R, 1931 Entomobryomorpha Troglophile KY, MO, TN, VA ○ ○ ○
Pseudosinella erehwon Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G2 VA ○ 2 ○
Pseudosinella espana Christiansen, 1961 Espana Cave Springtail Entomobryomorpha Troglobiont G3 MO, Mexico, TN ○ 1 ○
Pseudosinella espanita Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G1 KY, TN ○ 2 ○
Pseudosinella extra Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G1G2 VA ○ 1 ○
Pseudosinella flatua Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G1G2 NC, VA ○ 2 ○
Pseudosinella folsomi (Mills, HB, 1931) Entomobryomorpha Troglophile GNR AL, MO ○ ○ ○
Pseudosinella fonsa Christiansen & Bellinger, 1996 Fountain Cave Springtail Entomobryomorpha Troglobiont G3G4 IN, OH, TN ○ 1 ○
Pseudosinella georgia Christiansen, K & Bellinger, P, 1998 Entomobryomorpha Troglophile GA ○ ○ ○
Pseudosinella gisini Charistiansen, 1960 Entomobryomorpha Troglobiont G3G4 NC, VA, WV, Mexico, TN ○ 15 ○
Pseudosinella gisini carolina Christiansen, K & Bellinger, P, 1996 Entomobryomorpha Troglophile NC ○ ○ ○
Pseudosinella gisini virginia Christiansen, K & Bellinger, P, 1996 Entomobryomorpha Troglophile VA ○ ○ ○
Pseudosinella granda Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G3 VA ○ 1 ○
Pseudosinella hirsuta (Delamare Deboutteville, 1949) Hirsute Cave Springtail Entomobryomorpha Troglobiont G5 AL, GA, KY, TN, VA, Mexico ○ 8 ○
Pseudosinella josemarii Soto-Adames 2010 Entomobryomorpha Troglobiont GNR WV ○ ○ ○
Pseudosinella meganporteri Soto-Adames 2010 Entomobryomorpha Troglobiont G2 WV ○ ○ ○
Pseudosinella nata Christiansen & Bellinger, 1980 Entomobryomorpha Troglobiont G1 AL, Mexico, TN ○ 2 ○
Pseudosinella ops Christiansen & Bellinger, 1998 Entomobryomorpha Troglobiont VA ○ 1 ○
Pseudosinella orba Christiansen, 1960 Entomobryomorpha Troglobiont G3G4 KY, NC, TN, VA, WV, Mexico ○ 65 ○
Pseudosinella pecki Christiansen & Bellinger, 1980 Peck's Cave Springtail Entomobryomorpha Troglobiont G2G3 GA, NC, TN, AL, FL ○ 1 ○
Pseudosinella rolfsi Mills, HB, 1932 Entomobryomorpha Troglophile GNR AL ○ ○ ○
Pseudosinella sexoculata Schött, 1902 Entomobryomorpha Troglophile KY, NM ○ ○ ○
Pseudosinella spinosa (Delamare Deboutteville, 1949) Spiny Cave Springtail Entomobryomorpha Troglobiont G5 AL, GA, TN, Mexico ○ 5 ○
Pseudosinella testa Christiansen & Bellinger, 1980 Shelled Cave Springtail Entomobryomorpha Troglobiont G2G3 AR, VA, WV, Mexico ○ 4 ○
Pseudosinella vespera Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G2 NC, VA ○ 1 ○
Pseudosinella violenta (Folsom, JW, 1924) Entomobryomorpha Troglophile GNR AL, IA, TX, WI ○ ○ ○
Pseudosinella vita Christiansen & Bonet, 1980 Entomobryomorpha Troglobiont G1G2 NM, Mexico ○ 14 ○
Pseudotremia acheron Shear, 1972 Chordeumatida Troglobiont G1G2 TN ○ 2 ○
Pseudotremia aeacus Shear, 1972 Chordeumatida Troglobiont G1G2 GA ○ 3 ○
Pseudotremia amphiorax Shear, 1972 Chordeumatida Troglobiont G1G2 KY ○ 2 ○
Pseudotremia armesi Shear, 1972 Chordeumatida Troglophile G2 WV ○ ○ ○
Pseudotremia barri Lewis, 2005 Barr's Cave Millipede Chordeumatida Troglobiont G2 TN ○ ○ 9
Pseudotremia blacki Lewis, 2000 Black's Cave Milliped Chordeumatida Troglobiont G1 IN ○ ○ ○
Pseudotremia burnsorum Lewis, 2000 Burn's Cave Milliped Chordeumatida Troglobiont G1G2 IN ○ ○ ○
Pseudotremia callae Lewis, 2009 Call's Cave Milliped Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia carterensis (Packard, 1883) Chordeumatida Troglobiont G2G3 KY, IN ○ 2 ○
Pseudotremia cavernarum Cope, 1869 Ellett Valley Pseudotremia Millipede Chordeumatida Troglobiont G2G3 VA, IN, KY, TN ○ 8 ○
Pseudotremia cerberus Shear, 2011 Cerberus Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia cercops Shear, 1972 Chordeumatida Troglobiont G1 TN ○ 3 ○
Pseudotremia cocytus Shear, 1972 Chordeumatida Troglophile TN ○ ○ ○
Pseudotremia conservata Hoffman & Lewis, 1997 TNC Cave Milliped Chordeumatida Troglobiont G1G2 IN ○ ○ ○
Pseudotremia contorta Shear, 2011 Twisted-gonopod Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia cookorum Lewis, 2000 Cook's Cave Milliped Chordeumatida Troglobiont G1G2 IN ○ ○ ○
Pseudotremia culveri Shear, 2011 Culver's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia deprehendor Shear, 1972 Chordeumatida Troglobiont G1 TN, VA ○ 2 ○
Pseudotremia douglasi Lewis, 2009 Douglas' Cave Milliped Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia eburnea Loomis, 1939 Chordeumatida Troglobiont G2G4 AL, GA, TN, VA ○ 3 ○
Pseudotremia fergusoni Shear, 2011 Ferguson's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia fongi Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia fracta Chamberlin, 1951 Chordeumatida Troglophile GA, TN ○ ○ ○
Pseudotremia fulgida Loomis, 1943 Greenbrier Valley Cave Millipede Chordeumatida Troglobiont G3 WV ○ 2 ○
Pseudotremia garlandae Lewis, 2005 Garland's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia handi Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia hobbsi Hoffman, 1950 Chordeumatida Troglophile G4 VA, WV ○ ○ ○
Pseudotremia hoffmani Lewis, 2009 Hoffman's Cave Millipede Chordeumatida Troglobiont GNR TN ○ ○ ○
Pseudotremia hollidayi Lewis, 2005 Holliday's Cave Milliped Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia hubbardi Shear, 2011 Hubbard's Cave Milliped Chordeumatida Troglobiont G1G2 VA ○ ○ ○
Pseudotremia indianae Hoffman, 1958 Blue River Cave Millipede Chordeumatida Troglobiont G4 IN ○ 5 ○
Pseudotremia inexpectata Shear, 2011 Devault's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia jaculohamatum Shear, 2011 Harpoon Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia johnholsingeri Shear, 2011 Holsinger's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia kerboi Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia laudermilki Lewis, 2009 Laudermilk's Cave Millipede Chordeumatida Troglobiont GNR KY ○ ○ ○
Pseudotremia lethe Shear, 1972 Chordeumatida Troglobiont G1 TN ○ 1 ○
Pseudotremia lictor Shear, 1972 Chordeumatida Troglobiont G1 TN ○ 2 ○
Pseudotremia loomisi Shear, 2011 Loomis's Rough-backed Millipede Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia lusciosa (Loomis, 1939) Germany Valley Cave Millipede Chordeumatida Troglobiont G1 WV ○ 2 ○
Pseudotremia manni Lewis, 2005 Mann's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia merops Shear, 1972 Chordeumatida Troglobiont G1 KY ○ 2 ○
Pseudotremia minos Shear, 1972 Chordeumatida Troglobiont G1 AL, TN ○ 2 10
Pseudotremia momus Shear, 1972 Chordeumatida Troglophile G2 VA ○ ○ ○
Pseudotremia nefanda Shear, 1972 Clark Cave Millipede Chordeumatida Troglobiont G3G4 IN ○ 2 ○
Pseudotremia nodosa Loomis, 1939 Chordeumatida Troglobiont G4 TN, VA ○ 2 ○
Pseudotremia nyx Shear, 1972 Chordeumatida Troglobiont G1 AL ○ 3 ○
Pseudotremia orndorffi Shear, 2011 Orndorff's Cave Milliped Chordeumatida Troglobiont G2 VA ○ ○ ○
Pseudotremia peponocranium Shear, 2011 Pumpkin-headed Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia piscator Shear, 2011 Fisher Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia princeps Loomis, 1939 South Branch Valley Cave Millipede Chordeumatida Troglophile G1 WV ○ ○ ○
Pseudotremia purselli Lewis, 2000 Pursell's Cave Milliped Chordeumatida Troglobiont G1 IN ○ ○ ○
Pseudotremia reynoldsae Lewis, 2003 Reynolds Cave Millipede Chordeumatida Troglobiont G1 IN ○ ○ ○
Pseudotremia rhadamanthus Shear, 1972 Chordeumatida Troglobiont G1G2 AL, TN ○ 2 ○
Pseudotremia roebuckorum Lewis, 2005 Roebuck's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia ryensis Shear, 2011 Rye Cove Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia salifodina Shear, 2011 Jones' Saltpetre Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Pseudotremia salisae Lewis, 2000 Salisa's Cave Milliped Chordeumatida Troglobiont G2 IN, OH ○ 3 ○
Pseudotremia schneiderae Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia sheari Lewis, 2009 Shear's Cave Millipede Chordeumatida Troglobiont GNR TN ○ ○ ○
Pseudotremia shelleyi Lewis, 2009 Shelley's Cave Millipede Chordeumatida Troglobiont GNR NC ○ 3 ○
Pseudotremia spira Shear, 1972 Chordeumatida Troglobiont G1 KY ○ 2 ○
Pseudotremia styx Lewis, 2009 Chordeumatida Troglobiont GNR KY ○ ○ ○
Pseudotremia tuberculata Loomis, 1939 Chordeumatida Troglobiont G3 VA ○ 1 ○
Pseudotremia unca Shear, 1972 Chordeumatida Troglobiont G1 KY ○ 2 ○
Pseudotremia valga Loomis, 1943 Chordeumatida Troglobiont G1G2 TN, VA ○ 2 ○
Pseudotremia wallaceae Lewis, 2005 Wallace's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotremia withersi Lewis, 2009 Withers' Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudotriton montanus Baird, 1850 Mud Salamander Caudata Trogloxene G5 LC KY ○ ○ ○
Pseudotriton ruber (Latreille, 1801) Red Salamander Caudata Troglophile G5 LC AL, GA, KY, TN ○ ○ ○
Psilochorus cordatus (Bilimek, 1867) Araneae Troglophile ○ ○ ○
Psychoda cinerea Banks, 1894 Diptera Troglophile GNR KY ○ ○ ○
Psychoda minuta Banks, 1894 Diptera Troglophile GNR IN ○ ○ ○
Psychoda phalaenoides (Linnaeus, 1758) Diptera Troglophile GNR ○ ○ ○
Psychoda pusilla Tonnoir, 1922 Diptera Troglophile GA ○ ○ ○
Psychoda reevesi Quate, 2000 Diptera Troglophile GA, TN ○ ○ ○
Psychoda satchelli Quate, 1955 Diptera Troglophile GNR ○ ○ ○
Psychoda umbracola Quate, 1955 Diptera Troglophile IL, TN ○ ○ ○
Psychopomporus felipi Jean et al., 2012 Coleoptera Stygobiont GNR TX ○ 3 3
Psyllipsocus ramburii Selys-Longchamps, 1872 Cave Barklouse Psocodea Troglophile GNR AL, GA, IA, KY, NM, OK, TN ○ ○ ○
Ptenothrix atra (Linnæus, C, 1758) Symphypleona Trogloxene GNR IL, TN ○ ○ ○
Ptenothrix marmorata (Packard, AS, 1873) Symphypleona Trogloxene GNR TN ○ ○ ○
Ptenothrix quadrangularis (Mills, HB, 1934) Symphypleona Trogloxene TN ○ ○ ○
Pteronotropis harperi Fowler, 1941 Redeye Chub Cypriniformes Stygoxene FL ○ ○ ○
Pteronotus rubiginosus (Wagner, 1843) Wagner's Common Mustached Bat Chiroptera Trogloxene LC ○ ○ ○
Pterostichus coracinus (Newman, 1838) Coleoptera Trogloxene G5 AL, IL ○ ○ ○
Pterostichus lachrymosus (Newman, 1838) Coleoptera Trogloxene G5 ○ ○ ○
Pterostichus relictus (Newman, 1838) Coleoptera Trogloxene GNR AL, GA ○ ○ ○
Pterygoplichthys disjunctivus (Weber, 1991) Vermiculated Sailfin Catfish Siluriformes Stygoxene GNR LC FL ○ ○ ○
Ptomaphagus appalachianus (Peck, 1979) Coleoptera Troglophile AL, GA ○ ○ ○
Ptomaphagus barri Peck, 1973 Coleoptera Troglobiont G2G3 TN ○ 5 12
Ptomaphagus bedfordensis (Peck, 1979) Coleoptera Troglophile TN ○ ○ ○
Ptomaphagus californicus (LeConte, J. L., 1853) Coleoptera Trogloxene GNR CA ○ ○ ○
Ptomaphagus cavernicola Schwarz, 1898 Coleoptera Troglophile G4 AL, AR, FL, GA, IA, IL, MO, OK, TX ○ ○ ○
Ptomaphagus chromolithus Peck, 1984 Coleoptera Troglobiont G2G3 AL, TN ○ ○ 6
Ptomaphagus cocytus Peck, 1973 Coleoptera Troglobiont G1G2 AZ ○ 2 ○
Ptomaphagus consobrinus (LeConte, J. L., 1853) Coleoptera Trogloxene GNR AL ○ ○ ○
Ptomaphagus cumberlandus (Peck, 1979) Coleoptera Troglophile AL ○ ○ ○
Ptomaphagus elabra Peck, 1971 Coleoptera Troglophile ○ ○ ○
Ptomaphagus episcopus Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 3
Ptomaphagus fecundus Barr, 1963 Coleoptera Troglobiont G1 TN ○ ○ 3
Ptomaphagus fiskei Peck, 1973 Coleoptera Troglobiont G1G2 GA ○ 5 ○
Ptomaphagus fumosus (Peck, 1979) Coleoptera Troglophile GA ○ ○ ○
Ptomaphagus giaquintoi Jeannel, 1936 Coleoptera Troglophile ○ ○ ○
Ptomaphagus gypsum Peck, 1973 Coleoptera Troglophile ○ ○ ○
Ptomaphagus hatchi Jeannel, 1933 Hatch's Cave Fungus Beetle Coleoptera Troglobiont G3 AL, TN ○ 4 ○
Ptomaphagus hazelae Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 ○
Ptomaphagus hirtus (Tellkampf, 1844) Kentucky Cave Beetle Coleoptera Troglobiont G4 KY ○ 14 ○
Ptomaphagus hubrichti Barr, 1958 Coleoptera Troglobiont G2G3 TN ○ 4 ○
Ptomaphagus inyoensis Peck & Gnaspini, 1997 Coleoptera Troglobiont CA ○ ○ ○
Ptomaphagus julius Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 6
Ptomaphagus laticornis Jeannel, 1949 Coleoptera Troglobiont G1G2 AL ○ ○ 6
Ptomaphagus leo Peck, 1973 Coleoptera Troglophile ○ ○ ○
Ptomaphagus loedingi (Hatch, 1933) Coleoptera Troglobiont G1G2 AL ○ 5 ○
Ptomaphagus longicornis Jeannel, 1949 Coleoptera Troglobiont G3G4 AL ○ ○ 9
Ptomaphagus nashvillensis (Peck, 1979) Coleoptera Troglophile TN ○ ○ ○
Ptomaphagus neojonesi Peck & Newton, 2017 Coleoptera Troglophile AL ○ ○ ○
Ptomaphagus parashant Peck & Wynne, 2013 Coleoptera Troglobiont AZ ○ 2 ○
Ptomaphagus shapardi Sanderson, 1939 Coleoptera Troglophile AR, OK ○ ○ ○
Ptomaphagus solanum Peck, 1973 Coleoptera Troglobiont G1 AL ○ 2 3
Ptomaphagus spelaeus (Bilimek, 1867) Coleoptera Troglophile ○ ○ ○
Ptomaphagus torodei Peck, 1984 Torode's Cave Fungus Beetle Coleoptera Troglobiont G1G2 AL ○ ○ 6
Ptomaphagus ulkei Horn, 1885 Coleoptera Troglophile AL ○ ○ ○
Ptomaphagus valentinei Jeannel, 1933 Valentine's Cave Fungus Beetle Coleoptera Troglobiont G3G4 AL ○ 3 ○
Ptomaphagus walteri Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 6
Ptomaphagus whiteselli Barr, 1963 Coleoptera Troglobiont G2G3 AL, GA ○ 5 ○
Ptyoiulus impressus (Say, 1821) Flocculent Tailed Millipede Julida Trogloxene G5 ○ ○ ○
Puliciphora suavis Borgmeier, 1963 Diptera Accidental AL ○ ○ ○
Puliciphora virginiensis Malloch, 1912 Diptera Troglophile GA ○ ○ ○
Pupoides albilabris (C. B. Adams, 1841) Stylommatophora Trogloxene IL ○ ○ ○
Pygmarrhopalites altus (Christiansen, 1966) Symphypleona Troglobiont G2G3 KY ○ 2 ○
Pygmarrhopalites arcus (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont GNR ID ○ 5 ○
Pygmarrhopalites ashcraftensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont G1 IN ○ 2 ○
Pygmarrhopalites ater (Christiansen & Bellinger, 1998) Black Medusa Cave Springtail Symphypleona Troglobiont G3 IL, IN ○ ○ ○
Pygmarrhopalites benitus (Folsom, JW, 1896) Symphypleona Troglophile G1 TN ○ ○ ○
Pygmarrhopalites bimus (Christiansen, 1966) Symphypleona Troglobiont G3G4 IA, IN, KY, TN ○ 3 ○
Pygmarrhopalites buffaloensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont AR ○ 2 ○
Pygmarrhopalites caedus (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G3 VA ○ ○ ○
Pygmarrhopalites carolynae (Christiansen & Bellinger, 1996) Carolyn's Cave Springtail Symphypleona Troglobiont G4 IL, IN, VA, WV ○ ○ ○
Pygmarrhopalites clarus (Christiansen, 1966) Clarus Cave Springtail Symphypleona Troglobiont G4 AR, MO, SD, VA, WA, WI, WV ○ 3 ○
Pygmarrhopalites commorus (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G2G3 VA, WV ○ ○ ○
Pygmarrhopalites dubius (Christiansen, 1966) Symphypleona Troglobiont G2G3 AR, IA, MN ○ 3 ○
Pygmarrhopalites fransjanssens Soto-Adames & Taylor, 2013 Symphypleona Troglobiont IL ○ ○ ○
Pygmarrhopalites hirtus (Christiansen, K, 1966) Symphypleona Troglophile GNR IA, IL, WI ○ ○ ○
Pygmarrhopalites hubbardi (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont CO ○ 2 ○
Pygmarrhopalites incantator Soto-Adames & Taylor, 2013 Symphypleona Troglobiont IL ○ 3 ○
Pygmarrhopalites jay Christiansen & Bonet, 1996 Symphypleona Troglobiont G2G3 OK ○ ○ ○
Pygmarrhopalites lacuna (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G1G2 VA ○ ○ ○
Pygmarrhopalites leonardwoodensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont MO ○ 2 ○
Pygmarrhopalites lewisi (Christiansen & Bellinger, 1998) Lewis' Cave Springtail Symphypleona Troglobiont GNR IL, IN ○ 3 ○
Pygmarrhopalites madonnensis (Zeppelini & Christiansen, 2003) Madonna Cave Springtail Symphypleona Troglobiont GNR IL ○ 3 ○
Pygmarrhopalites marshalli (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G3 TN, VA ○ 4 ○
Pygmarrhopalites obtusus (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont G1 VA ○ 7 ○
Pygmarrhopalites pavo (Christiansen & Bellinger, 1998) Symphypleona Troglobiont G3 MO, TN, VA, WV, IN ○ 6 ○
Pygmarrhopalites plethorasari Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont MO ○ 6 ○
Pygmarrhopalites pygmaeus (Wankel, 1860) Symphypleona Troglophile GNR AL, AR, GA, IA, IL, KY, MO, TN, VA ○ ○ ○
Pygmarrhopalites sacer (Christiansen & Bellinger, 1998) Symphypleona Troglobiont G2 VA, WV ○ ○ ○
Pygmarrhopalites salemensis Soto-Adames & Taylor, 2013 Symphypleona Troglobiont IL ○ ○ ○
Pygmarrhopalites sapo (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont IL ○ 8 ○
Pygmarrhopalites sextus (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont G1 VA ○ 7 ○
Pygmarrhopalites shoshoneiensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont NV ○ 4 ○
Pygmarrhopalites silvus (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G2 VA ○ ○ ○
Pygmarrhopalites texensis (Christiansen & Bonet, 1996) Symphypleona Troglobiont G3G4 TX ○ ○ ○
Pygmarrhopalites youngsteadtii Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont AR ○ ○ ○
Pylodictis olivaris (Rafinesque, 1818) Flathead Catfish Siluriformes Stygoxene G5 LC FL ○ ○ ○
Quedius alesi Brunke, 2020 Coleoptera Troglophile OK ○ ○ ○
Quedius capucinus (Gravenhorst, 1806) Coleoptera Troglophile G5 KY ○ ○ ○
Quedius erythrogaster Mannerheim, 1852 Coleoptera Troglophile G5 AL, GA, IL, TN, VA ○ ○ ○
Quedius fulgidus (Fabricius, 1792) Large Rove Beetle Coleoptera Troglophile GNR AL, GA, IL, IN, KY ○ ○ ○
Quedius mesomelinus (Marsham, 1802) Staph Beetle Coleoptera Troglophile GNR IA, MN ○ ○ ○
Quedius spelaeus Horn, 1871 Coleoptera Troglophile GNR IA, IL, IN, WI ○ ○ ○
Remasellus parvus (Steeves, 1964) Isopoda Stygobiont G1G2 FL ○ 13 16
Rhadine austinica Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 82 18
Rhadine babcocki (Barr, 1960) Coleoptera Troglophile TX ○ ○ ○
Rhadine bullis Reddell & Cokendolpher, 2004 Coleoptera Troglobiont GNR TX ○ 22 ○
Rhadine caudata (LeConte, 1863) Coleoptera Troglophile G3 AL, GA, KY, TN ○ ○ ○
Rhadine exilis (Barr & Lawrence, 1960) Coleoptera Troglobiont G3 ESA TX ○ 133 12
Rhadine grubbsi Reddell & Duperre, 2009 Coleoptera Troglobiont TX ○ 7 ○
Rhadine howdeni (Barr & Lawrence, 1960) Coleoptera Troglophile TX ○ ○ ○
Rhadine infernalis (Barr & Lawrence, 1960) Coleoptera Troglobiont G2G3 ESA TX ○ 152 17
Rhadine insolita Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 5 ○
Rhadine ivyi Reddell & Cokendolpher, 2004 Coleoptera Troglobiont TX ○ 2 ○
Rhadine koepkei (Barr, 1960) Coleoptera Troglobiont G1 TX ○ 12 ○
Rhadine larvalis LeConte, 1846 Coleoptera Troglophile GNR AL, FL, GA, IL ○ ○ ○
Rhadine longicollis Benedict, 1927 Coleoptera Troglophile NM, TX ○ ○ ○
Rhadine noctivaga Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 94 14
Rhadine ozarkensis Sanderson & Miller, 1941 Coleoptera Troglobiont GH AR ○ 2 ○
Rhadine persephone Barr, 1974 Tooth Cave Beetle Coleoptera Troglobiont G1G2 NE ESA TX ○ 90 9
Rhadine reyesi Reddell & Cokendolpher, 2001 Coleoptera Troglobiont GNR TX ○ 36 ○
Rhadine rubra (Barr, 1960) Coleoptera Troglophile TX ○ ○ ○
Rhadine russelli Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 13 3
Rhadine speca (Barr, 1960) Coleoptera Troglobiont G2 TX ○ ○ ○
Rhadine sprousei Reddell & Cokendolpher, 2004 Coleoptera Troglobiont TX ○ 3 ○
Rhadine subterranea (Van Dyke, 1918) Coleoptera Troglobiont G2 TX ○ 143 15
Rhadine tenebrosa (Barr, 1960) Coleoptera Troglobiont G1G2 TX ○ 22 5
Rhagidia diversicolor (C. L. Koch, 1838) Actinedida Troglophile WV ○ ○ ○
Rhagidia varia Zacharda 1985 Actinedida Troglobiont G5 VA, WV ○ ○ ○
Rhamdia quelen (Quoy & Gaimard, 1824) Silver Catfish Siluriformes Stygophile LC ○ ○ ○
Rhantus suturalis (W. S. MacLeay, 1825) Cosmopolitan Diving Beetle Coleoptera Accidental MO ○ ○ ○
Rheocyclops indiana Reid in Reid et al. 1999 Indiana Groundwater Copepod Cyclopoida Stygobiont G1 IN, TN ○ 2 ○
Rheocyclops virginianus (Reid, 1993) Cyclopoida Stygobiont VA ○ 5 ○
Rhinichthys obtusus Agassiz, 1854 Western Blacknose Dace Cypriniformes Stygoxene G5 LC AL ○ ○ ○
Rhinocricus ocraceus Brölemann, 1900 Spirobolida Troglophile ○ ○ ○
Rhinotus panamanus (Loomis, 1964) Polyzoniida Troglophile ○ ○ ○
Rhyacodrilus sodalis (Eisen, 1879) Aquatic Oligochaete Worm Tubificida Stygophile GNR WV ○ ○ ○
Rhyacophila nigrita Banks, 1907 Trichoptera Stygoxene KY ○ ○ ○
Rhynchomonas nasuta (Stokes) Klebs, 1892 Bodonida Stygoxene AL, KY ○ ○ ○
Robustocheles hilli (Strandtmann, 1971) Actinedida Troglophile AZ, GA, IN, KY, TN, VA, WA, WV ○ ○ ○
Robustocheles occulta Zacharda & Pugsley, 1988 Actinedida Troglobiont G2G3 AB, IA, WA ○ ○ ○
Rotomelita ana Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 3 ○
Rotomelita lokoa Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 13 ○
Rudiloria trimaculata (Wood, 1864) Polydesmida Accidental G5 VA ○ ○ ○
Rudolfina cavernicola Marshall & Fitzgerald, 1997 Diptera Troglophile CO ○ ○ ○
Rugathodes sexpunctatus (Emerton, 1882) Common Six-Spotted Cobweaver Araneae Trogloxene G5 ○ ○ ○
Rugilus dentatus Say, 1831 Coleoptera Accidental GNR MO ○ ○ ○
Rugosuscandona scharfi Kulkoyluoglu et al., 2017 Podocopida Stygobiont TX ○ ○ ○
Ryloviella pilosa (Chappuis, 1929) Harpacticoida Stygophile IN, KY ○ ○ ○
Rymosia fasciata (Meigen, 1804) Diptera Troglophile IN ○ ○ ○
Rymosia filipes Loew, 1870 Diptera Troglophile GNR IN, TN ○ ○ ○
Rymosia triangularis Shaw, 1935 Diptera Troglophile GNR AL, IL ○ ○ ○
Sabacon cavicolens (Packard, 1884) North American Cavernicolous Harvestman Opiliones Troglophile GNR AL, AR, GA, IL, KY, MT, TN ○ ○ ○
Saetigerocreagris phyllisae (J. C. Chamberlin, 1930) Pseudoscorpiones Troglophile G1 CA ○ ○ ○
Sagittocythere barri (Hart & Hobbs, 1961) Podocopida Stygobiont G5 AL, IN, KY, TN ○ 76 ○
Sagittocythere stygia Hart & Hart, 1966 Podocopida Stygobiont G1 KY ○ 1 ○
Salmasellus howarthi Lewis, 2001 Isopoda Stygobiont GNR OR, WA ○ 4 9
Salmasellus steganothrix Bowman, 1975 Isopoda Stygobiont G2 AB, MT ○ 15 ○
Salpingoeca amphoridium James-Clark Craspedida Stygoxene KY ○ ○ ○
Satan eurystomus Hubbs & Bailey, 1947 Widemouth Blindcat Siluriformes Stygobiont G1G2 VU ESA? TX ○ 48 ○
Sayornis phoebe (Latham, 1790) Eastern Phoebe Passeriformes Trogloxene G5 LC AL, AR, FL, GA, IL, KY, OK, TN, VA ○ ○ ○
Scalopus aquaticus (Linnaeus, 1758) Eastern Mole Eulipotyphla Accidental G5 LC KY ○ ○ ○
Scaphinotus andrewsii (T.W.Harris, 1839) Andrew's Snail-Eating Beetle Coleoptera Trogloxene KY ○ ○ ○
Scaphinotus unicolor (Fabricius, 1787) Coleoptera Trogloxene AL ○ ○ ○
Scatopse notata (Linnaeus, 1758) Black Compost Fly Diptera Trogloxene GNR IL ○ ○ ○
Sceloporus undulatus (Bosc & Daudin, 1801) Eastern Fence Lizard Squamata Accidental G5 LC AR, KY ○ ○ ○
Schaefferia asymmetrica (Christiansen & Wang, 2006) Poduromorpha Troglobiont CO ○ ○ ○
Schaefferia steinmanni (Christiansen & Wang, 2006) Poduromorpha Troglobiont CO ○ ○ ○
Schaefferia unica (Christiansen & Wang, 2006) Poduromorpha Troglobiont CO ○ ○ ○
Schizocosa ocreata (Hentz, 1844) Brush-Legged Split Wolf Spider Araneae Accidental G5 IL ○ ○ ○
Schizocosa saltatrix (Hentz, 1844) Araneae Accidental G5 ○ ○ ○
Schornikovdona bellensis Kulkoyluoglu et al., 2017 Podocopida Stygobiont TX ○ ○ ○
Schrankia howarthi Davis & Medeiros, 2009 Lepidoptera Troglobiont HI ○ 3 105
Sciaphilus asperatus (Bonsdorff, 1785) Strawberry Root Weevil Coleoptera Accidental GNR ○ ○ ○
Scincella lateralis (Say, 1822) Ground Skink Squamata Accidental G5 LC AL, AR ○ ○ ○
Sciodrepoides latinotum Peck & Cook, 2002 Coleoptera Troglophile GNR MO ○ ○ ○
Sciodrepoides terminans (LeConte, J. L., 1850) Coleoptera Trogloxene G5 GA ○ ○ ○
Sciurus carolinensis Gmelin, 1788 Eastern Gray Squirrel Rodentia Accidental G5 LC AL, AR, KY ○ ○ ○
Sclerobunus cavicolens (Banks, 1905) Opiliones Troglobiont G1G2 MT ○ 8 24
Sclerobunus jemez Derkarabetian & Hedin, 2014 Opiliones Troglobiont NV ○ ○ ○
Sclerobunus klomax Derkarabetian & Hedin, 2014 Opiliones Troglobiont GNR NM ○ 5 16
Sclerobunus madhousensis (Briggs, 1971) Opiliones Troglobiont G1G2 UT ○ 4 20
Sclerobunus speoventus Derkarabetian & Hedin, 2014 Opiliones Troglobiont CO ○ 15 32
Sclerobunus steinmanni Derkarabetian & Hedin, 2014 Opiliones Troglobiont CO ○ 3 21
Sclerobunus ungulatus (Briggs, 1971) Opiliones Troglobiont G1G2 NV ○ 29 21
Scoliocentra brachypterna (Loew, 1873) Diptera Trogloxene GNR GA, IA, IL, IN, KY ○ ○ ○
Scoliocentra fraterna Loew, 1863 Diptera Trogloxene GNR IN ○ ○ ○
Scoliocentra tincta (Walker, 1849) Diptera Troglophile IN ○ ○ ○
Scoliopteryx libatrix (Linnaeus, 1758) Herald Moth Lepidoptera Trogloxene G5 AL, GA, IA, IL, IN, KY, OK, TN ○ ○ ○
Scolopocryptops sexspinosus Say (1821) Eastern Fire Centipede Scolopendromorpha Trogloxene GNR AL, GA, MO, TN ○ ○ ○
Scoterpes alabama Shear, 2010 Alabama Cave Millipede Chordeumatida Troglobiont AL ○ ○ ○
Scoterpes austrinus Loomis, 1943 Chordeumatida Troglobiont G3G4 AL, GA ○ 3 ○
Scoterpes blountensis Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes copei (Packard, 1871) Chordeumatida Troglobiont G3G4 KY, TN, GA ○ 6 ○
Scoterpes hesperus Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes jackdanieli Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes musicarustica Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes nudus Chamberlin, 1943 Chordeumatida Troglobiont GA ○ ○ ○
Scoterpes sollmani Lewis, 2000 Sollman's Cave Millipede Chordeumatida Troglobiont G1 IN ○ ○ ○
Scoterpes stewartpecki Shear, 2010 Peck's Cave Millipede Chordeumatida Troglobiont AL, TN ○ ○ ○
Scoterpes syntheticus (Shear, 1972) Chordeumatida Troglobiont AL ○ ○ ○
Scoterpes tombarri Shear, 2010 Barr's Cave Millipede Chordeumatida Troglobiont KY, TN ○ ○ ○
Scoterpes tricorner Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes ventus Shear, 1972 Chordeumatida Troglobiont G3 TN ○ 1 ○
Scoterpes willreevesi Shear, 2010 Reeves' Cave Millipede Chordeumatida Troglobiont AL, GA ○ ○ ○
Scotinella redempta (Gertsch, 1941) Araneae Trogloxene GNR IL ○ ○ ○
Scutacarus strinatii Cooreman, 1959 Trombidiformes Accidental IA ○ ○ ○
Scutigera coleoptrata Linnaeus (1758) House Centipede Scutigeromorpha Trogloxene GNR IL ○ ○ ○
Scytodes intricata Banks, 1909 Spitting Spiders Araneae Trogloxene ○ ○ ○
Scytonotus australis Hoffman, 1962 Polydesmida Accidental TN ○ ○ ○
Scytonotus granulatus (Say, 1821) Granulated Millipede Polydesmida Trogloxene G5 AL, IN, KY, PA, TN ○ ○ ○
Seborgia hershleri Holsinger, 1992 Amphipoda Stygobiont G1G2 TX ○ 38 ○
Seborgia relicta Holsinger, 1980 Amphipoda Stygobiont G2G3 TX ○ 96 ○
Sehirus cinctus (Palisot, 1811) White-Margined Burrower Bug Hemiptera Trogloxene GNR IL ○ ○ ○
Selachochthonius heterodentatus (Beier, 1955) Pseudoscorpiones Troglophile ○ ○ ○
Semionellus placidus (Wood, 1864) Salmon Cherry Millipede Polydesmida Accidental G3 IL ○ ○ ○
Semotilus atromaculatus (Mitchill, 1818) Creek Chub Cypriniformes Stygoxene G5 LC AL, KY, TN ○ ○ ○
Sensillanura barberi (Handschin, 1928) Poduromorpha Troglophile GNR AR, TN ○ ○ ○
Sensillanura illina (Christiansen, K & Bellinger, P, 1980) Poduromorpha Trogloxene IL ○ ○ ○
Sepedophilus crassus (Gravenhorst, 1802) Coleoptera Accidental G5 IA ○ ○ ○
Sepedophilus littoreus (Linnaeus, 1758) Staph Beetle Coleoptera Troglophile GNR GA ○ ○ ○
Sequoiadesmus krejcae Shear & Shelley, 2008 Polydesmida Troglobiont CA ○ 1 ○
Shibaia longisensilla (Shiba, 1969) Actinedida Troglophile AR ○ ○ ○
Sicifera bifurca (O.P. Hay, 1882) Amphipoda Stygoxene GNR IL ○ ○ ○
Sicifera dentata (Hubricht, 1943) Amphipoda Stygoxene GNR IL, IN, OH ○ ○ ○
Sigmocheir calaveras Chamberlin, 1951 Polydesmida Accidental CA ○ ○ ○
Sigmoria fumimontis Shelley, 1981 Polydesmida Accidental TN ○ ○ ○
Simulium parnassum Malloch, 1914 Dark Black Fly Diptera Trogloxene GNR GA ○ ○ ○
Simulium tuberosum (Lundstrom, 1911) Tubercled Black Fly Diptera Trogloxene G5 ○ ○ ○
Simulium venustum Say, 1823 Whitestockinged Black Fly Diptera Trogloxene G5 ○ ○ ○
Simulium vernum (Macquart, 1826) Diptera Trogloxene ○ ○ ○
Sinella agna Christiansen & Bellinger, 1980 Entomobryomorpha Troglobiont G3G4 WV ○ 2 ○
Sinella alata Christiansen, 1960 Entomobryomorpha Troglobiont G5 IN, Mexico ○ 1 ○
Sinella avita Christiansen, 1960 Avita Cave Springtail Entomobryomorpha Troglobiont G3G4 IL, IN, KY, MO, Mexico ○ 3 ○
Sinella baca Christiansen & Bonet, 1980 Entomobryomorpha Troglobiont G3G4 BC, CA, TX ○ 2 ○
Sinella barri Christiansen, 1960 Barr's Cave Springtail Entomobryomorpha Troglobiont G5 AL, AR, IL, IN, KY, MO, TN, VA, WI, Mexico ○ 6 ○
Sinella basidens Bonet, 1934 Entomobryomorpha Troglobiont G3G4 IN, KY, TN ○ 3 ○
Sinella caeca Zhao in Wenying, 1992 Entomobryomorpha Troglophile AL, IA, MN, MO, WI ○ ○ ○
Sinella cavernarum (Packard, 1888) Entomobryomorpha Troglobiont G5 IL, IN, KY, MD, MO, OH, PA, TN, VA, LA, WA ○ 11 ○
Sinella hoffmani Wray, 1952 Hoffman's Cave Springtail Entomobryomorpha Troglobiont G5 MD, PA, TN, VA, WV, KY, Mexico ○ 5 ○
Sinella krekeleri Christiansen, 1960 Entomobryomorpha Troglobiont G3G4 KY, WV, Mexico ○ 3 ○
Sinella tecta Christiansen & Bonet, 1980 Entomobryomorpha Troglobiont G1G2 CA ○ 2 ○
Sinella yosiia Bellinger & Christiansen, 1974 Entomobryomorpha Troglobiont G2G3 HI ○ 1 ○
Sitophilus oryzae Hustache, A., 1930 Rice Weevil Coleoptera Accidental GNR IA ○ ○ ○
Skistodiaptomus pallidus (Herrick, 1879) Calanoida Stygoxene GNR IN, KY ○ ○ ○
Sminthurides lepus Mills, HB, 1934 Symphypleona Trogloxene GNR TN ○ ○ ○
Sminthurides malmgreni Janssens, F, 1992 Symphypleona Trogloxene IL ○ ○ ○
Sminthurinus henshawi (Folsom, JW, 1896) Symphypleona Trogloxene GNR IL ○ ○ ○
Soldanellonyx chappuisi Walter, 1917 Actinedida Stygophile G1 IN ○ ○ ○
Soldanellonyx chapuisi Walter, 1917 Actinedida Stygobiont G1 IN ○ ○ ○
Soldanellonyx monardi Walter, 1919 Mite Actinedida Stygophile IN ○ ○ ○
Soldanellonyx morardi Walter, 1919 Actinedida Stygobiont G1 IN ○ ○ ○
Solenopsis geminata (Fabricius, 1804) Tropical Fire Ant Hymenoptera Accidental ○ ○ ○
Somatochlora tenebrosa (Say, 1840) Clamp-Tipped Emerald Odonata Accidental G5 LC AL ○ ○ ○
Sorex cinereus Kerr, 1792 Cinereus Shrew Eulipotyphla Accidental G5 LC ○ ○ ○
Sorex fumeus G. M. Miller, 1895 Smoky Shrew Eulipotyphla Accidental G5 LC KY ○ ○ ○
Sorex hoyi Baird, 1857 American Pygmy Shrew Eulipotyphla Accidental G5 LC ○ ○ ○
Sorex palustris Richardson, 1828 American Water Shrew Eulipotyphla Accidental G5 LC ○ ○ ○
Sparganophilus eiseni Smith, 1895 Crassiclitellata Troglophile AL, NC ○ ○ ○
Sparganophilus tamesis Benham, 1892 Amerikaanse Wortelwroeter Crassiclitellata Troglophile GA ○ ○ ○
Spelaedrilus multiporus Cook 1975 Lumbriculida Stygobiont G1 VA ○ 2 ○
Spelaeorchestia koloana Bousfield & Howarth, 1976 Kauai Cave Amphipod Amphipoda Troglobiont G1G2 EN ESA HI ○ 4 ○
Spelaeorchestia pahoehoe Myers & Lowry, 2023 Amphipoda Troglobiont HI ○ ○ ○
Speleobama vana Park, 1951 Coleoptera Troglobiont G1 AL ○ ○ ○
Speleochus blanchardensis Carlton, 2012 Coleoptera Troglobiont AR, AZ ○ 1 ○
Speleochus croceus Park, 1960 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Speleochus eurous (Park, 1960) Coleoptera Troglobiont G2G3 AL ○ ○ ○
Speleochus ferus Park, 1951 Coleoptera Troglobiont G1G2 AL, TN ○ 1 ○
Speleochus macosar Carlton, 2012 Coleoptera Troglobiont AR, AZ ○ 1 ○
Speleochus steevesi (Park, 1960) Coleoptera Troglobiont G4 AL, TN ○ ○ ○
Speleochus stygicus Park, 1951 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Speleochus synstygicus Park, 1956 Coleoptera Troglobiont G1 AL ○ ○ ○
Speleomaster lexi Briggs, 1974 Opiliones Troglobiont G1G2 ID ○ 2 6
Speleomaster pecki Briggs, 1974 Opiliones Troglobiont G1G2 ID ○ 1 ○
Speleonura kenchristianseni Palacios-Vargas & Benito, 2007 Poduromorpha Troglobiont G1 VA ○ ○ ○
Speleonychia sengeri Briggs, 1974 Opiliones Troglobiont G1G2 WA ○ 27 5
Speleonycta ozarkensis Espinasa et al., 2010 Ozark Cave Bristletail Zygentoma Troglobiont AR, OK ○ ○ 1
Spelobia clunipes (Meigen, 1830) Diptera Troglophile GNR VA ○ ○ ○
Spelobia maculipennis (Spuler, 1925) Diptera Trogloxene GNR CO ○ ○ ○
Spelobia pseudosetaria (Duda, 1918) Diptera Trogloxene GNR CO ○ ○ ○
Spelobia semioculata (Richards, 1965) Diptera Troglophile GNR IA, IL, TN, VA, WI ○ ○ ○
Spelobia tenebrarum (Aldrich, 1897) Cave dung fly Diptera Troglobiont G5 AL, AR, GA, IL, IN, KY, MO, NY, OK, PA, TN, VA, WV ○ 1 ○
Spelobia tuberculosa Marshall, 1985 Diptera Trogloxene GNR CO ○ ○ ○
Spelungula cavernicola Forster, 1987 Nelson Cave Spider Araneae Troglophile DD ○ ○ ○
Speocirolana hardeni Bowman, 1992 Isopoda Stygobiont G2G3 TX ○ 116 4
Speodesmus bicornourus Causey, 1959 Polydesmida Troglobiont G2G3 TX ○ 252 ○
Speodesmus castellanus Elliott, 2004 Polydesmida Troglobiont TX ○ 115 ○
Speodesmus echinourus Loomis, 1939 Polydesmida Troglobiont G2G3 TX ○ 366 ○
Speodesmus falcatus Elliott, 2004 Sickled Cave Millipede Polydesmida Troglobiont GNR TX ○ 29 ○
Speodesmus ivyi Elliott, 2004 Ivy's Cave Millipede Polydesmida Troglobiont GNR TX ○ 18 ○
Speodesmus reddelli Elliott, 2004 Reddell's Cave Millipede Polydesmida Troglobiont GNR TX ○ 71 ○
Speolepta leptogaster (Winnertz, 1864) Diptera Troglophile IN ○ ○ ○
Speoplatyrhinus poulsoni Cooper & Kuehne, 1974 Alabama cavefish Percopsiformes Stygobiont G1 CR ESA AL ○ 9 27
Speorthus reyesi Shear & Reddell, 2017 Polydesmida Troglobiont TX ○ ○ ○
Speorthus tuganbius Chamberlin, 1952 Polydesmida Troglobiont G2G3 NM, TX ○ ○ ○
Speoseya grahami Causey, 1963 Chordeumatida Troglobiont G2G3 CA ○ 1 ○
Speotrombicula trifurca (Ewing, 1933) Trombidiformes Troglophile ○ ○ ○
Spermophora senoculata (Dugès, 1836) Shortbodied Cellar Spider Araneae Trogloxene GNR FL ○ ○ ○
Sphaeridia serrata (Folsom & Mills, 1938) Symphypleona Trogloxene GNR IL ○ ○ ○
Sphaerium simile (Say, 1817) Grooved Fingernailclam Sphaeriida Accidental ○ ○ ○
Sphaerium transversum (Say, 1829) Long Fingernail Clam Sphaeriida Accidental MO ○ ○ ○
Sphaeroderus stenostomus (Weber, 1801) Small Snail-Eating Ground Beetle Coleoptera Trogloxene G5 GA ○ ○ ○
Sphaeromicola moria Hart, 1978 Podocopida Stygobiont G1G2 TX ○ ○ ○
Sphalloplana buchanani (Hyman, 1937) Buchanan's Cave Flatworm Tricladida Stygobiont G1G2 KY, MD, TN ○ 10 ○
Sphalloplana californica Kenk, 1977 California Cave Flatworm Tricladida Stygobiont G1G2 CA ○ 4 ○
Sphalloplana chandleri Kenk, 1977 Chandler's Cave Flatworm Tricladida Stygobiont G1G2 IN, TN, VA ○ 17 ○
Sphalloplana consimilis Kenk, 1977 Powell Valley Cave Flatworm Tricladida Stygobiont G2G3 TN, VA ○ 13 ○
Sphalloplana culveri Kenk, 1977 Culver's Cave Flatworm Tricladida Stygobiont G1 WV ○ 1 ○
Sphalloplana evaginata Kenk, 1977 Perryville Cave Flatworm Tricladida Stygobiont G1G2 MO ○ 20 ○
Sphalloplana georgiana Hyman, 1954 Georgia Cave Flatworm Tricladida Stygobiont G1 GA ○ 2 ○
Sphalloplana hoffmasteri (Hyman, 1954) Hoffmaster's Cave Flatworm Tricladida Stygobiont G3G4 MD, WV ○ 12 ○
Sphalloplana holsingeri Kenk, 1977 Holsinger's Cave Flatworm Tricladida Stygobiont G1G2 NE VA ○ 10 ○
Sphalloplana hubrichti (Hyman, 1945) Hubricht's Cave Flatworm Tricladida Stygobiont G2G4 IL, MO ○ 21 ○
Sphalloplana hypogea Kenk, 1984 Tricladida Stygobiont G1G2 VA ○ 2 ○
Sphalloplana kansensis Hyman, 1945 Kansas Cave Flatworm Tricladida Stygobiont G1G2 KS ○ ○ ○
Sphalloplana mohri Hyman, 1938 Tricladida Stygobiont G3G4 TX ○ 12 ○
Sphalloplana percoeca (Packard, 1879) Tricladida Stygobiont G5 AL, KY, TN, WV ○ 25 ○
Sphalloplana pricei Hyman, 1937 Refton Cave Flatworm Tricladida Stygobiont G2G3 MD, PA ○ 12 ○
Sphalloplana subtilis Kenk, 1977 Biggers Cave Flatworm Tricladida Stygobiont G1G2 NE VA ○ 3 ○
Sphalloplana virginiana Hyman, 1945 Rockbridge County Cave Flatworm Tricladida Stygobiont G1 VA ○ 1 ○
Sphalloplana weingartneri Kenk, 1970 Weingartner's cave flatworm Tricladida Stygobiont G4 IN ○ 3 ○
Sphendononema guildingii Newport G. (1845) Quarenta-Pernas Scutigeromorpha Trogloxene ○ ○ ○
Spinanycta alabamensis Espinasa, Gutierrez, Hinkle & Niemiller, 2024 Alabama Cave Bristletail Zygentoma Troglobiont AL ○ ○ ○
Spiniphora slossonae (Malloch, 1912) Diptera Accidental GNR AL ○ ○ ○
Spongilla lacustris (Linnaeus, 1759) Common Freshwater Sponge Spongillida Accidental G5 KY ○ ○ ○
Steatoda triangulosa (Walckenaer, 1802) Triangulate Bud Spider Araneae Trogloxene GNR IL ○ ○ ○
Steatornis caripensis von Humboldt, 1817 Oilbird Steatornithiformes Trogloxene LC ○ ○ ○
Stegana coleoptrata (Scopoli, 1763) Pomace Fly Diptera Trogloxene ○ ○ ○
Stelidota geminata (Say, 1825) Strawberry Sap Beetle Coleoptera Trogloxene GNR KY ○ ○ ○
Stenistoderus rubripennis (LeConte, J. L., 1880) Red-Winged Rove Beetle Coleoptera Accidental GNR IA ○ ○ ○
Stenochironomus pulchripennis (Coquillett, 1902) Diptera Trogloxene GNR ○ ○ ○
Stenolophus ochropezus (Say, 1823) Coleoptera Trogloxene G5 MO, TN ○ ○ ○
Stenonema femoratum (Say, 1823) Cream Cahill Ephemeroptera Stygoxene G5 AR, IL ○ ○ ○
Stenophilus californicus (Chamberlin, 1930) Geophilomorpha Troglobiont G1G2 CA ○ ○ ○
Stenoponia americana (Baker, 1899) Siphonaptera Trogloxene GNR IA, TN ○ ○ ○
Stenotrema angellum Hubricht, 1958 Kentucky Slitmouth Stylommatophora Trogloxene G4 KY, TN ○ ○ ○
Stenotrema edvardsi (Bland, 1858) Ridge-And-Valley Slitmouth Stylommatophora Trogloxene G4 KY ○ ○ ○
Stenotrema exodon (Pilsbry, 1900) Alabama Slitmouth Stylommatophora Trogloxene G2 AL ○ ○ ○
Stenotrema hirsutum (Say, 1817) Hairy Slitmouth Stylommatophora Trogloxene G5 KY ○ ○ ○
Stenotrema spinosum (I. Lea, 1831) Carinate Slitmouth Stylommatophora Trogloxene G4 AL, TN ○ ○ ○
Stenotrema stenotrema (L. Pfeiffer, 1842) Inland Slitmouth Stylommatophora Trogloxene AL, TN ○ ○ ○
Stenotrema turbinella (Clench & Archer, 1933) Stylommatophora Trogloxene AL ○ ○ ○
Stenus alacer Casey, 1884 Coleoptera Accidental MO ○ ○ ○
Sternopsylla distincta (Rothschild, 1903) Siphonaptera Trogloxene NM ○ ○ ○
Storeria occipitomaculata (Storer, 1839) Mexican Yellowbelly Brown Snake [Hidalgoensis] Squamata Accidental G5 LC AR ○ ○ ○
Striaria eldora Chamberlin, 1953 Chordeumatida Troglobiont G2G3 CA ○ ○ ○
Striatura meridionalis (Pilsbry & Ferriss, 1906) Median Striate Stylommatophora Trogloxene G5 TN ○ ○ ○
Strigamia branneri Bollman C.H. (1888) Geophilomorpha Troglophile IA, VA ○ ○ ○
Strix varia Barton, 1799 Barred Owl Strigiformes Trogloxene G5 LC FL ○ ○ ○
Strobilops affinis Pilsbry, 1893 Eightfold Pinecone Stylommatophora Trogloxene G4 IL ○ ○ ○
Strobilops texasianus Pilsbry & Ferriss, 1906 Southern Pinecone Stylommatophora Trogloxene G5 GA ○ ○ ○
Stygobromus abditus Holsinger, 1978 James Cave Amphipod Amphipoda Stygobiont G2G3 VA ○ 16 ○
Stygobromus acicularis Holsinger & Lewis, 2025 Amphipoda Stygobiont GNR MO ○ ○ ○
Stygobromus ackerlyi Holsinger, 1978 Amphipoda Stygobiont G1G2 GA ○ 7 ○
Stygobromus alabamensis (Stout, 1911) Alabama cave amphipod Amphipoda Stygobiont G5 AL, AR, KS, LA, MO, OK, TN, TX, MS ○ 207 ○
Stygobromus albapinus Taylor & Holsinger, 2011 White Pine amphipod Amphipoda Stygobiont NV ○ 4 ○
Stygobromus allegheniensis (Holsinger, 1967) Amphipoda Stygobiont G5 MD, NY, PA, WV ○ 72 10
Stygobromus amicus Holsinger, 2014 Friendly Cave Amphipod Amphipoda Stygobiont G1 MD ○ 4 ○
Stygobromus anacostensis Cannizzaro et al., 2024 Amphipoda Stygobiont DC ○ ○ 75
Stygobromus araeus (Holsinger, 1969) Tidewater Interstitial Amphipod Amphipoda Stygobiont G3 VU NC, VA ○ 24 ○
Stygobromus arbucklensis Holsinger & Lewis, 2025 Arbuckle Mountains Cave Amphipod Amphipoda Stygobiont GNR OK ○ ○ ○
Stygobromus arizonensis Holsinger, 1974 Arizona Cave Amphipod Amphipoda Stygobiont G1 VU AZ ○ 5 ○
Stygobromus bakeri Gibson & Hutchins, 2021 Amphipoda Stygobiont TX ○ 27 ○
Stygobromus balconis (Hubricht, 1943) Balcones Cave Amphipod Amphipoda Stygobiont G2G3 VU TX ○ 28 ○
Stygobromus baroodyi Holsinger, 1978 Rockbridge County Cave Amphipod Amphipoda Stygobiont G1 VA ○ 11 1
Stygobromus barri (Holsinger, 1967) Barr's Cave Amphipod Amphipoda Stygobiont G2G3 VU MO, TN ○ 8 ○
Stygobromus barryi Holsinger, 1978 Amphipoda Stygobiont G1 TN ○ ○ ○
Stygobromus bifurcatus (Holsinger, 1967) Bifurcated Cave Amphipod Amphipoda Stygobiont G3G4 VU TX ○ 77 4
Stygobromus biggersi Holsinger, 1978 Biggers' Cave Amphipod Amphipoda Stygobiont G2G4 MD, PA, VA, WV ○ 22 1
Stygobromus blinni Wang & Holsinger, 2001 Blinn's Cave Amphipod Amphipoda Stygobiont G1G2 AZ ○ 2 ○
Stygobromus borealis Hoslinger, 1978 Taconic Cave Amphipod Amphipoda Stygobiont G2G3 MA, NY, QC, VT ○ 7 ○
Stygobromus boultoni Wang & Holsinger, 2001 Boulton's Cave Amphipod Amphipoda Stygobiont G1G2 AZ ○ 4 ○
Stygobromus bowmani (Holsinger, 1967) Bowman's Cave Amphipod Amphipoda Stygobiont G1G2 VU OK ○ 3 ○
Stygobromus caecilius Holsinger, 2011 Cecil Groundwater Amphipod Amphipoda Stygobiont G1 MD ○ 1 ○
Stygobromus canadensis Holsinger, 1980 Castleguard Cave Amphipod Amphipoda Stygobiont G1 AB ○ 2 ○
Stygobromus carolinensis Holsinger, 1978 Carolina Seep Amphipod Amphipoda Stygobiont GH NC ○ 4 ○
Stygobromus cherylae Wang & Holsinger, 2001 Amphipoda Stygobiont G1 CA ○ 5 ○
Stygobromus clantoni (Creaser, 1934) Clanton's Cave Amphipod Amphipoda Stygobiont G4 VU KS, MO, AR, OK ○ 35 ○
Stygobromus coloradensis Ward, 1977 Colorado Cave Amphipod Amphipoda Stygobiont G1G2 CO ○ 16 ○
Stygobromus conradi (Holsinger, 1967) Conrad's Cave Amphipod Amphipoda Stygobiont G1 VU VA ○ 15 ○
Stygobromus cooperi (Holsinger, 1967) Cooper's Cave Amphipod Amphipoda Stygobiont G1G2 VU WV ○ 4 ○
Stygobromus cowani Wang & Holsinger, 2001 Cowan's Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus culveri Holsinger, 1978 Culver's Cave Amphipod Amphipoda Stygobiont G1 WV ○ 4 ○
Stygobromus cumberlandus Holsinger, 1978 Cumberland Cave Amphipod Amphipoda Stygobiont G3G4 VA ○ 9 ○
Stygobromus curroae Wang & Holsinger, 2001 Curro's Cave Amphipod Amphipoda Stygobiont G1G2 NM ○ 3 ○
Stygobromus dejectus (Holsinger, 1967) Cascade Cave Amphipod Amphipoda Stygobiont G1G2 VU TX ○ 4 7
Stygobromus dicksoni Holsinger, 1978 Dickson cave stygobromid Amphipoda Stygobiont G5 AL, GA, TN ○ 22 ○
Stygobromus doughertyensis Cannizzaro & Sawicki in Cannizzaro, Balding, Stine & Sawicki, 2019 Dougherty Plain Cave Amphipod Amphipoda Stygobiont G1G2 FL, GA ○ 6 ○
Stygobromus duplus Wang & Holsinger, 2001 Amphipoda Stygobiont G1G2 WA ○ 2 ○
Stygobromus elatus (Holsinger, 1967) Amphipoda Stygobiont G1 VU AR ○ 3 ○
Stygobromus elliotti Holsinger, 1974 Elliott's Cave Amphipod Amphipoda Stygobiont G1G2 WA ○ 11 ○
Stygobromus emarginatus Hubricht, 1943 Greenbrier Cave Amphipod Amphipoda Stygobiont G3 VU MD, WV ○ 82 2
Stygobromus ephemerus (Holsinger, 1969) Ephemeral Cave Amphipod Amphipoda Stygobiont G1G2 VU VA ○ 9 ○
Stygobromus estesi Holsinger, 1978 Craig County Cave Amphipod Amphipoda Stygobiont G4 VA ○ 9 ○
Stygobromus exilis Hubricht, 1943 Central Kentucky Cave Amphipod Amphipoda Stygobiont G5 AL, KY, TN ○ 105 ○
Stygobromus fecundus Holsinger, 1978 Amphipoda Stygobiont G1G2 TN ○ 6 ○
Stygobromus felleri Holsinger, 2011 Feller's Groundwater Amphipod Amphipoda Stygobiont G1 MD ○ 3 ○
Stygobromus fergusoni Holsinger, 1978 Ferguson's Cave Amphipod Amphipoda Stygobiont G1 VA ○ 5 1
Stygobromus finleyi Holsinger, 1978 Finley's Cave Amphipod Amphipoda Stygobiont G3G4 TN, VA ○ 8 ○
Stygobromus flagellatus (Benedict, 1896) Ezell's Cave Amphipod Amphipoda Stygobiont G2G3 VU TX ○ 308 5
Stygobromus floridanus Holsinger & Sawicki, 2016 Florida Panhandle Cave Amphipod Amphipoda Stygobiont G1G2 FL, CA ○ 12 43
Stygobromus foliatus Holsinger, 2011 Rappahannock Spring Amphipod Amphipoda Stygobiont G2 MD, VA ○ 10 2
Stygobromus fontinalis Wang & Holsinger, 2001 Amphipoda Stygobiont G1G2 CO ○ 5 ○
Stygobromus franzi Holsinger, 1978 Franz's Cave Amphipod Amphipoda Stygobiont G3G4 MD, PA, WV ○ 46 ○
Stygobromus gallawayae Wang & Holsinger, 2001 Gallaway's Cave Amphipod Amphipoda Stygobiont G1 CA ○ 3 ○
Stygobromus gardneri Holsinger & Lewis, 2025 Amphipoda Stygobiont GNR AR, MO ○ ○ ○
Stygobromus glacialis Wang & Holsinger, 2001 Glacier National Park Cave Amphipod Amphipoda Stygobiont G1 MT ○ 7 ○
Stygobromus gracilipes (Holsinger, 1967) Amphipoda Stygobiont G3G4 MD, PA, VA, WV ○ 39 8
Stygobromus gradyi Holsinger, 1974 Grady's Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 9 ○
Stygobromus grahami Holsinger, 1974 Graham's Cave Amphipod Amphipoda Stygobiont G2 CA ○ 15 ○
Stygobromus grandis Holsinger, 1978 Amphipoda Stygobiont G1 GA ○ 3 ○
Stygobromus hadenoecus (Holsinger, 1966) Devil's Sinkhole Amphipod Amphipoda Stygobiont G1G2 VU TX ○ 38 ○
Stygobromus harai Holsinger, 1974 Hara's Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 6 ○
Stygobromus hayi (Hubricht & Mackin, 1940) Hay's Spring Amphipod Amphipoda Stygobiont G1G2 EN ESA DC, PA ○ 11 11
Stygobromus herbsti Wang & Holsinger, 2001 Herbst's Cave Amphipod Amphipoda Stygobiont G2G3 NV ○ 11 ○
Stygobromus heteropodus Hubricht, 1943 Pickle Springs Amphipod Amphipoda Stygobiont G1G2 VU MO ○ 2 ○
Stygobromus hoffmani Holsinger, 1978 Hoffman's Cave Amphipod Amphipoda Stygobiont G2 VA ○ 4 1
Stygobromus holsingeri Ward, 1977 Holsinger's Cave Amphipod Amphipoda Stygobiont G1G2 CO ○ 33 ○
Stygobromus hubbardi Holsinger, 2009 Hupp's Hill Cave Amphipod Amphipoda Stygobiont G1 VA ○ 3 ○
Stygobromus hubbsi Shoemaker, 1942 Malheur Cave Amphipod Amphipoda Stygobiont G1G2 VU OR ○ 6 ○
Stygobromus hyporheicus Wang & Holsinger, 2001 Hyporheic Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus idahoensis Wang & Holsinger, 2001 Idaho Groundwater Amphipod Amphipoda Stygobiont G1 ID ○ 2 ○
Stygobromus imperialis Wang & Holsinger, 2001 Imperial Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus indentatus (Holsinger, 1967) Tidewater Amphipod Amphipoda Stygobiont G3 VU MD, NC, VA ○ 22 3
Stygobromus inexpectatus Holsinger, 1978 Amphipoda Stygobiont G1 AL ○ 2 ○
Stygobromus interitus Holsinger, 1978 New Castle Murder Hole Cave Amphipod Amphipoda Stygobiont G1G2 VA ○ 2 ○
Stygobromus interstitialis Wang & Holsinger, 2001 Interstitial Amphipod Amphipoda Stygobiont G1G3 CO ○ 3 ○
Stygobromus iowae Hubricht, 1943 Iowa Cave Amphipod Amphipoda Stygobiont G2G3 IA, IL, MN ○ 8 ○
Stygobromus jemezensis Wang & Holsinger, 2001 Jemez Mountains Groundwater Amphipod Amphipoda Stygobiont G1G2 NM ○ 2 ○
Stygobromus kenki Holsinger, 1978 Rock Creek Groundwater Amphipod Amphipoda Stygobiont G2 DC, MD, VA ○ 15 ○
Stygobromus lacicolus Holsinger, 1974 Lake Tahoe Groundwater Amphipod Amphipoda Stygobiont G1 CA, NV ○ 5 ○
Stygobromus lanensis Wang & Holsinger, 2001 Lane County Groundwater Amphipod Amphipoda Stygobiont G1G2 OR ○ 2 ○
Stygobromus latus Wang & Holsinger, 2001 Amphipoda Stygobiont G1G3 WA ○ 4 ○
Stygobromus leensis Holsinger, 1978 Lee County Cave Amphipod Amphipoda Stygobiont G1G2 VA ○ 5 1
Stygobromus limbus Wang & Holsinger, 2001 Border Cave Amphipod Amphipoda Stygobiont G1G2 TX ○ 1 ○
Stygobromus longipes (Holsinger, 1966) Long-legged Cave Amphipod Amphipoda Stygobiont G2G3 VU TX ○ 14 7
Stygobromus lucifugus (Hay, 1882) Rubious Cave Amphipod Amphipoda Stygobiont G1G2 EX IL ○ ○ ○
Stygobromus mackenziei Holsinger, 1974 MacKenzie's Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 4 ○
Stygobromus mackini Hubricht, 1943 Southwestern Virginia Cave Amphipod Amphipoda Stygobiont G5 IN, KY, TN, VA, WV, TX ○ 152 5
Stygobromus mausi Holsinger, 2009 Round Hill Cave Amphipod Amphipoda Stygobiont G1 VA ○ 2 ○
Stygobromus minutus Holsinger, 1978 Amphipoda Stygobiont G2G3 GA ○ 3 ○
Stygobromus montanensis Holsinger, 1974 Amphipoda Stygobiont G1G2 MT ○ 3 ○
Stygobromus montanus (Holsinger, 1967) Mountain Cave Amphipod Amphipoda Stygobiont G1 VU AR ○ 6 ○
Stygobromus morrisoni (Holsinger, 1967) Morrison's Cave Amphipod Amphipoda Stygobiont G1G2 VU VA, WV ○ 12 1
Stygobromus mundus (Holsinger, 1967) Bath County Cave Amphipod Amphipoda Stygobiont G1 VU VA ○ 7 1
Stygobromus myersae Wang & Holsinger, 2001 Myers' Groundwater Amphipod Amphipoda Stygobiont G1 CA, NV ○ 6 ○
Stygobromus mysticus Holsinger, 1974 Secret Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus nanus Holsinger, 1978 Amphipoda Stygobiont G1 WV ○ 3 ○
Stygobromus nortoni (Holsinger, 1969) Norton's Cave Amphipod Amphipoda Stygobiont G2G3 VU TN ○ 4 ○
Stygobromus obrutus Holsinger, 1978 Pittsylvania Well Amphipod Amphipoda Stygobiont G1G2 VA ○ 2 ○
Stygobromus obscurus Holsinger, 1974 Amphipoda Stygobiont G1G2 MT ○ 3 ○
Stygobromus onondagaensis (Hubricht & Mackin, 1940) Onondaga Cave Amphipod Amphipoda Stygobiont G3 VU AR, KS, MO, OK ○ 61 ○
Stygobromus oregonensis Holsinger, 1974 Amphipoda Stygobiont G1G2 OR ○ 3 ○
Stygobromus ozarkensis (Holsinger, 1967) Ozark Cave Amphipod Amphipoda Stygobiont G4 VU AR, MO, OK ○ 59 1
Stygobromus parvus (Holsinger, 1969) Minute Cave Amphipod Amphipoda Stygobiont G2 VU WV, VA ○ 10 ○
Stygobromus paveki Cannizzaro, Sawicki, Schaefer, Castrillo, Maxwell, Burch, Culver & Niemiller, 2026 Pavek's Groundwater Amphipod Amphipoda Stygobiont MD ○ ○ ○
Stygobromus paxillus Holsinger, 2011 Prettyboy Groundwater Amphipod Amphipoda Stygobiont G1 MD ○ 3 ○
Stygobromus pecki (Holsinger, 1967) Peck's Cave Amphipod Amphipoda Stygobiont G1G2 EN ESA TX ○ 36 33
Stygobromus pennaki (Ward, 1977) Pennak Cave Amphipod Amphipoda Stygobiont G2G3 CO ○ 9 ○
Stygobromus phreaticus Holsinger, 1978 Northern Virginia Well Amphipod Amphipoda Stygobiont G1 VA ○ 4 ○
Stygobromus pizzinii (Shoemaker, 1938 Pizzini's Cave Amphipod Amphipoda Stygobiont G3G4 VU DC, MD, PA, VA ○ 89 3
Stygobromus pollostus Holsinger, 1978 Least Cave Amphipod Amphipoda Stygobiont G2G3 WV ○ 8 ○
Stygobromus pseudospinosus Holsinger, 1978 Luray Caverns Amphipod Amphipoda Stygobiont G1 VA ○ 7 ○
Stygobromus putealis (Holmes, 1909) Wisconsin Well Amphipod Amphipoda Stygobiont G2G3 VU WI ○ 2 ○
Stygobromus puteanus Holsinger, 1974 Amphipoda Stygobiont G1G2 MT ○ 2 ○
Stygobromus quatsinensis Holsinger & Shaw, 1987 Quatsino Cave Amphipod Amphipoda Stygobiont G1G2 AK, BC ○ 21 ○
Stygobromus rallus Wang & Holsinger, 2001 Narrow Groundwater Amphipod Amphipoda Stygobiont G1G3 WA ○ 3 ○
Stygobromus redactus Holsinger, 1978 Pattons Cave Amphipod Amphipoda Stygobiont G1 WV ○ 3 1
Stygobromus reddelli (Holsinger, 1966) Amphipoda Stygobiont G1G2 VU TX ○ 6 ○
Stygobromus rudolphi Wang & Holsinger, 2001 Rudolph's Groundwater Amphipod Amphipoda Stygobiont G1 CA ○ 3 ○
Stygobromus russelli (Holsinger, 1967) Russell's Cave Amphipod Amphipoda Stygobiont G4 TX ○ 396 11
Stygobromus saltuaris Wang & Holsinger, 2001 Williamette Natonal Forest Groundwater Amphipod Amphipoda Stygobiont G1G2 OR ○ 2 ○
Stygobromus scottsruni Cannizzaro, Sawicki, Schaefer, Castrillo, Maxwell, Burch, Culver & Niemiller, 2026 Scotts Run Groundwater Amphipod Amphipoda Stygobiont VA ○ ○ ○
Stygobromus secundus Bousfield & Holsinger, 1981 Amphipoda Stygobiont G1 AB ○ 4 ○
Stygobromus sextarius Holsinger, 2009 Capital Area Groundwater Amphipod Amphipoda Stygobiont G1 DC, MD, VA ○ 4 ○
Stygobromus sheldoni Holsinger, 1974 Sheldon's Groundwater Amphipod Amphipoda Stygobiont G1 CA ○ 15 ○
Stygobromus sierrensis Holsinger, 1974 Sierra Groundwater Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus simplex Wang & Holsinger, 2001 Simple Cave Amphipod Amphipoda Stygobiont G1G2 CO ○ 1 ○
Stygobromus smithi Hubricht, 1943 Alabama Well Amphipod Amphipoda Stygobiont G2G3 VU AL, TN ○ 3 ○
Stygobromus sparsus Holsinger, 1978 Gregory Cave Amphipod Amphipoda Stygobiont G1 TN ○ 8 ○
Stygobromus spinatus (Holsinger, 1967) Spiny Cave Amphipod Amphipoda Stygobiont G2G3 VU WV, VA ○ 92 1
Stygobromus spinosus (Hubricht & Mackin, 1940) Blue Ridge Cave Amphipod Amphipoda Stygobiont G3 VA ○ 16 1
Stygobromus stegerorum Holsinger, 1978 Madison Cave Amphipod Amphipoda Stygobiont G1 VA ○ 5 6
Stygobromus stellmacki (Holsinger, 1967) Stellmack's Cave Amphipod Amphipoda Stygobiont G1G2 VU PA ○ 6 ○
Stygobromus subtilis (Hubricht, 1943) Subtle Cave Amphipod Amphipoda Stygobiont G2 VU IL, MI, MO ○ 17 ○
Stygobromus tahoensis Holsinger, 1974 Amphipoda Stygobiont G1 CA, NV ○ 16 ○
Stygobromus tenuis (Smith, 1874) Amphipoda Stygobiont G4 CT, DC, MA, MD, NY, PA, VA, WV ○ 292 102
Stygobromus trinus Wang & Holsinger, 2001 Trinity County Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus tritus Holsinger, 1974 Worn Groundwater Amphipod Amphipoda Stygobiont G1 MT ○ 5 ○
Stygobromus urospinatus Wang & Holsinger, 2001 Spinose Cave Amphipod Amphipoda Stygobiont G1G3 CO ○ 6 ○
Stygobromus utahensis Wang & Holsinger, 2001 Utah Cave Amphipod Amphipoda Stygobiont G1G2 UT ○ 2 ○
Stygobromus vitreus Cope, 1872 Mammoth Cave stygobromid Amphipoda Stygobiont G4 AL, KY, TN, IN ○ 40 ○
Stygobromus wahkeenensis Wang & Holsinger, 2001 Wahkeena Creek Groundwater Amphipod Amphipoda Stygobiont G1G2 OR ○ 3 ○
Stygobromus wardi Wang & Holsinger, 2001 Ward's Groundwater Amphipod Amphipoda Stygobiont G1G2 CO ○ 3 ○
Stygobromus wengerorum Holsinger, 1974 Wenger Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 4 ○
Stygonyx courtneyi Bousfield & Holsinger, 1989 Amphipoda Stygobiont G1G2 OR ○ 13 ○
Stygoparnus comalensis Barr & Spangler, 1992 Comal Springs Dryopid Beetle Coleoptera Stygobiont G1G2 ESA TX ○ 14 ○
Stygoporus oregonensis Larson & LaBonte, 1994 Coleoptera Stygobiont G1G2 OR ○ 4 12
Stygopyrgus bartonensis Hershler & Longley, 1986 Barton cavesnail Littorinimorpha Stygobiont G1 TX ○ 30 15
Stygopyrgus gracilis Perez et al., 2025 Littorinimorpha Stygobiont TX ○ ○ ○
Stygopyrgus variabilis Perez et al., 2025 Littorinimorpha Stygobiont TX ○ ○ ○
Stylodrilus beattiei Cook, 1975 Lumbriculida Stygobiont G1 VA, WV ○ 5 ○
Stylodrilus californianus Rodriguez, 1996 Lumbriculida Stygobiont CA ○ 10 ○
Stylodrilus wahkeenensis Rodriguez & Coates, 1996 Lumbriculida Stygobiont AL, OR, TN ○ 6 ○
Subulina octona (Bruguière, 1789) Miniature Awlsnail Stylommatophora Trogloxene G5 ○ ○ ○
Superodontella cornifer (Mills, HB, 1934) Poduromorpha Trogloxene GNR IL ○ ○ ○
Superodontella substriata (Wray, 1952) Poduromorpha Trogloxene GNR IL ○ ○ ○
Sus scrofa Linnaeus, 1758 Eurasian Wild Pig Artiodactyla Accidental G5 LC KY ○ ○ ○
Sylvilagus floridanus (Allen, 1890) Eastern Cottontail Lagomorpha Accidental G5 LC AL, KY ○ ○ ○
Syngrapha alias (Ottolengui, 1902) Spruce Climbing Cutworm Lepidoptera Trogloxene G5 ○ ○ ○
Syrphus rectus Osten Sacken, 1875 Diptera Accidental ○ ○ ○
Tachinus fimbriatus Gravenhorst, 1802 Coleoptera Trogloxene G5 GA ○ ○ ○
Tachinus fumipennis (Say, 1832) Coleoptera Trogloxene G5 IA ○ ○ ○
Tachycines asynamorus Adelung, 1902 Greenhouse Camel Cricket Orthoptera Trogloxene GNR AL, GA, IL, OK ○ ○ ○
Tachypeza humeralis Melander, 1928 Diptera Trogloxene GNR ○ ○ ○
Tachyporus maculipennis LeConte, J. L., 1866 Coleoptera Trogloxene IL ○ ○ ○
Tachys proximus (Say, 1823) Coleoptera Troglophile GNR NM ○ ○ ○
Tachys pumilus (Dejean, 1831) Coleoptera Accidental GNR MO ○ ○ ○
Tachys scitulus LeConte, 1848 Coleoptera Trogloxene G5 VA ○ ○ ○
Tachyura lutea (Andrewes, 1925) Coleoptera Trogloxene TN ○ ○ ○
Tadarida brasiliensis (I. Geoffroy, 1824) Brazilian Free-Tailed Bat Chiroptera Trogloxene G5 LC FL, KY, NM, TX ○ ○ ○
Taiyutyla loftinae Shear & Krejca, 2011 Chordeumatida Troglobiont CA ○ ○ ○
Tamias striatus (Linnaeus, 1758) Eastern Chipmunk Rodentia Trogloxene G5 LC AR, IN, KY, TN ○ ○ ○
Tamiasciurus hudsonicus (Erxleben, 1777) Red Squirrel Rodentia Accidental G5 LC KY ○ ○ ○
Tanytarsus recurvatus Brundin, 1947 Diptera Trogloxene GNR GA ○ ○ ○
Taracus aspenae Shear, 2018 Opiliones Troglobiont OR ○ 1 ○
Taracus fluvipileus Shear & Warfel, 2016 Opiliones Troglobiont CA ○ 1 ○
Taracus marchingtoni Shear & Warfel, 2016 Opiliones Troglobiont OR ○ ○ ○
Taracus silvestrii Briggs, 1969 Opiliones Troglobiont G1G2 OR ○ 1 ○
Taracus taylori Shear & Warfel, 2016 Opiliones Troglobiont GNR NV ○ 6 ○
Taranucnus ornithes (Barrows, 1940) Araneae Accidental GNR AL ○ ○ ○
Tarebia granifera (Lamarck, 1816) Quilted Melania Cerithiida Stygoxene G5 LC FL ○ ○ ○
Tarlina noorundi Gray, 1987 Araneae Troglophile ○ ○ ○
Tarnania tarnanii (Dziedzicki, 1910) Diptera Troglophile GNR ○ ○ ○
Tartarocreagris altimana Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris amblyopa Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris attenuata Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris domina Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris grubbsi Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris hoodensis Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ 3 ○
Tartarocreagris infernalis (Muchmore, 1969) Pseudoscorpiones Troglobiont G2G3 TX ○ 5 ○
Tartarocreagris intermedia Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris proserpina Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris reyesi Muchmore, 2001 Pseudoscorpiones Troglobiont GNR TX ○ ○ ○
Tartarocreagris texana (Muchmore, 1969) Tooth Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 DD ESA TX ○ 1 ○
Taxidea taxus (Schreber, 1777) American Badger Carnivora Trogloxene G5 LC ○ ○ ○
Tayshaneta anopica (Gertsch, 1974) Araneae Troglobiont G1G2 TX ○ 11 ○
Tayshaneta archambaulti Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 19 ○
Tayshaneta bullis (Cokendolpher, 2004) Araneae Troglobiont GNR TX ○ 12 ○
Tayshaneta concinna (Gertsch, 1974) Araneae Troglobiont G1G2 TX ○ 15 ○
Tayshaneta fawcetti Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 41 ○
Tayshaneta microps (Gertsch, 1974) Government Canyon Bat Cave Spider Araneae Troglobiont G1 ESA TX ○ 24 ○
Tayshaneta myopica (Gertsch, 1974) Tooth Cave Spider Araneae Troglobiont G1G2 DD ESA TX ○ 74 ○
Tayshaneta oconnorae Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 1 ○
Tayshaneta sandersi Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 9 ○
Tayshaneta sprousei Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 14 ○
Tayshaneta vidrio Ledford et al., 2012 Araneae Troglobiont GNR TX ○ ○ ○
Tayshaneta whitei Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 12 ○
Tectocepheus velatus (Michael, 1880) Sarcoptiformes Accidental GNR IA ○ ○ ○
Tegenaria domestica (Clerck, 1757) Barn Funnel Weaver Araneae Troglophile GNR AL, GA, IL, IN ○ ○ ○
Tegenaria pagana C. L. Koch, 1840 Rural Funnel-Web Spider Araneae Trogloxene NM ○ ○ ○
Telomerina cana Marshall & Roháček, 1984 Diptera Troglophile AL, TN ○ ○ ○
Telomerina chillcotti Marshall & Roháček, 1984 Diptera Troglophile GNR TN, TX ○ ○ ○
Telomerina flavipes (Meigen, 1830) Diptera Troglophile GNR AR ○ ○ ○
Tenuiphantes sabulosus (Keyserling, 1886) Araneae Trogloxene GNR AL, IL, TN ○ ○ ○
Terrapene carolina (Linnaeus, 1758) Common Box Turtle Testudines Accidental G5 VU AL, FL, GA, IL, KY, TN ○ ○ ○
Terrapene triunguis (Agassiz, 1857) Three-Toed Box Turtle Testudines Accidental GNR AR ○ ○ ○
Terrilimosina racovitzai (Bezzi, 1911) Diptera Troglophile GNR IA, IL, NY, PA, WI ○ ○ ○
Tethysbaena texana (Maguire, 1965) Thermosbaenacea Stygobiont GNR TX ○ 112 ○
Tetracion jonesi Hoffman, 1956 Jones' Cave Millipede Callipodida Troglobiont G3G4 AL, TN ○ 7 17
Tetracion tennesseensis Causey, 1959 Tennessee Cave Millipede Callipodida Troglobiont G2G3 TN ○ ○ 36
Tetragnatha elongata Walckenaer, 1841 Elongate Stilt Spider Araneae Trogloxene G5 ○ ○ ○
Tetragnatha laboriosa Hentz, 1850 Long-Jawed Orb Weaver Araneae Trogloxene G5 IN ○ ○ ○
Tetragnatha shoshone Levi, 1981 Amerikanische Streckerspinne Araneae Trogloxene GNR IL ○ ○ ○
Tetragnatha versicolor Walckenaer, 1841 Versicolor Long-Jawed Orbweaver Araneae Trogloxene G5 AL, KY ○ ○ ○
Texamaurops reddelli Barr & Steeves, 1963 Kretschmarr Cave Mold Beetle Coleoptera Troglobiont G1G2 ESA TX ○ 17 ○
Texanobathynella aaronswinki Camacho & Hutchins, 2021 Bathynellacea Stygobiont TX ○ ○ ○
Texanobathynella bowmani (Delamare et al., 1975) Bathynellacea Stygobiont GNR TX ○ ○ ○
Texanobathynella coloradoensis Camacho & Hutchins, 2021 Bathynellacea Stygobiont TX ○ ○ ○
Texanobathynella sachi Cho, 1996 Bathynellacea Stygobiont CA ○ 3 ○
Texapyrgus diaboli (Pilsbry & Ferriss, 1906) Devil Tryonia Littorinimorpha Stygobiont TX ○ ○ ○
Texapyrgus longleyi Thompson & Hershler, 1991 Striated hydrobe Littorinimorpha Stygobiont G1 TX ○ 9 1
Texella cokendolpheri Ubick & Briggs, 1992 Cokendolpher Cave Harvestman Opiliones Troglobiont G1 ESA TX ○ 5 ○
Texella elliotti Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 1 ○
Texella hardeni Ubick & Briggs, 1992 Opiliones Troglobiont G1G2 TX ○ 5 ○
Texella hilgerensis Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 2 ○
Texella martensi Ubick, 2021 Opiliones Troglobiont CA ○ 1 ○
Texella mulaiki Goodnight & Goodnight, 1942 Opiliones Troglobiont G2G3 TX ○ 26 ○
Texella reyesi Ubick & Briggs, 1992 Bone Cave Harvestman Opiliones Troglobiont G2G3 ESA TX ○ 210 3
Texella shoshone Ubick & Briggs, 1992 Shoshone Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Texella tuberculata Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 4 ○
Texella welbourni Ubick & Briggs, 1992 Opiliones Troglobiont G1G2 NM ○ 1 ○
Texella whitei Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 3 ○
Texicerberus amistad Hutchins et al., 2022 Isopoda Stygobiont GNR TX ○ 23 ○
Texicerberus castilloi Hutchins et al., 2022 Isopoda Stygobiont TX ○ 6 ○
Texicerberus schotteae Hutchins et al., 2022 Isopoda Stygobiont GNR TX ○ 22 ○
Texiweckelia texensis (Holsinger, 1973) Amphipoda Stygobiont G2G3 TX ○ 268 ○
Texiweckeliopsis insolita (Holsinger, 1980) Amphipoda Stygobiont G3 TX ○ 303 ○
Texoreddellia aquilonalis Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 8
Texoreddellia capitesquameo Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 1
Texoreddellia coahuilensis Espinasa & Giribet, 2009 Zygentoma Troglobiont TX, Mexico ○ ○ 5
Texoreddellia media Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 17
Texoreddellia occasus Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ ○
Texoreddellia texensis (Ulrich, 1902) Zygentoma Troglobiont G4 TX ○ ○ 17
Thalassaphorura encarpata (Denis, 1931) Springtail Poduromorpha Trogloxene IL ○ ○ ○
Thalkethops grallatrix Crabill, 1960 Scolopendromorpha Troglobiont G1G2 NM ○ 2 ○
Thamnophis proximus (Say, 1823) Western Ribbon Snake Squamata Accidental G5 LC ○ ○ ○
Thaumatogryllus cavicola Gurney & Rentz, 1978 Vovanoes Cave Cricket Orthoptera Troglobiont G1G2 VU HI ○ 4 ○
Theatops phanus Chamberlin, 1951 Scolopendromorpha Troglobiont G2G3 TX ○ 30 ○
Theotima makua Gertsch, 1973 Araneae Troglobiont G2G3 HI ○ 1 ○
Theridion flavonotatum Becker, 1879 Araneae Trogloxene AL ○ ○ ○
Theridion glaucescens Becker, 1879 Araneae Trogloxene GNR AL ○ ○ ○
Theridiosoma gemmosum (Koch, 1877) Common Eastern Ray Spider Araneae Trogloxene G5 AL, GA, IL, IN ○ ○ ○
Theridula opulenta (Walckenaer, 1841) Cobweb Spiders Araneae Accidental GNR AL ○ ○ ○
Thermobia domestica (Packard, 1873) Firebrat Archaeognatha Accidental GNR IA ○ ○ ○
Thesiastes fossulatus (Brendel, 1890) Coleoptera Trogloxene IL ○ ○ ○
Thymoites marxi (Crosby, 1906) Araneae Trogloxene GNR AL, GA ○ ○ ○
Thymoites minero Roth, 1992 Araneae Troglobiont G1G2 AZ ○ 1 ○
Tidarren sisyphoides (Walckenaer, 1841) Tent Cobweb Weaver Araneae Trogloxene AL ○ ○ ○
Tingupa pallida Loomis, 1939 Tingupa Cave Millipede Chordeumatida Troglobiont G4 AR, IL, MO, TN ○ 13 ○
Tipula abdominalis (Say, 1823) Giant Crane Fly Diptera Trogloxene GNR GA ○ ○ ○
Tipula algonquin Alexander, 1915 Diptera Trogloxene GNR AR ○ ○ ○
Tipula caloptera Loew, 1863 Diptera Trogloxene GNR ○ ○ ○
Tmesiphorus costalis LeConte, J. L., 1849 Coleoptera Trogloxene GNR AL ○ ○ ○
Tokophrya cyclopum (Claparède & Lachmann, 1859) Bütschli, 1889 Endogenida Stygoxene KY ○ ○ ○
Tolus appalachius Goodnight & Goodnight, 1942 Appalachian Cave Spider Opiliones Troglobiont G3G4 AL, TN ○ 2 4
Tomocerus curtus Christiansen, 1965 Entomobryomorpha Troglophile IA ○ ○ ○
Tomocerus dubius Christiansen, 1965 Entomobryomorpha Troglophile FL, KY ○ ○ ○
Tomocerus flavescens Gisin, 1944 Entomobryomorpha Troglophile GA, IL, TN ○ ○ ○
Tomocerus vulgaris (Tullberg, 1871) Entomobryomorpha Troglophile GNR TN, WI ○ ○ ○
Tomoderus impressulus Casey, 1895 Coleoptera Accidental TN ○ ○ ○
Toxidium compressum Zimmermann, C., 1869 Coleoptera Accidental GNR IA ○ ○ ○
Trachelipus rathkii (Brandt, 1833) Rathke's Woodlouse Isopoda Trogloxene GNR KY, WV ○ ○ ○
Trachemys scripta (Thunberg, 1792) Pond Slider Testudines Accidental G5 LC FL ○ ○ ○
Traegaardhia gracilis Zacharda, 2010 Actinedida Troglophile ○ ○ ○
Traegaardhia holsingeri (Zacharda, 1980) Actinedida Troglobiont AR, KY, MO, OK ○ ○ ○
Traegaardhia paralleloseta (Zacharda, 1985) Actinedida Troglobiont G1 VA ○ ○ ○
Traegaardhia similis Zacharda et al., 2010 Actinedida Troglobiont AR, MO ○ 12 ○
Trechus cumberlandus Barr, 1962 Coleoptera Troglophile G2 KY, TN ○ ○ ○
Trechus tennesseensis Barr, 1962 Coleoptera Troglophile GNR TN ○ ○ ○
Trechus tuckaleechee Barr, 1962 Coleoptera Troglophile GNR TN ○ ○ ○
Triacanthella copelandi (Wray, DL, 1963) Poduromorpha Trogloxene G1 TN ○ ○ ○
Trichobius major Coquillett, 1899 Diptera Trogloxene GNR FL, NM ○ ○ ○
Trichocera fattigiana Alexander, 1952 Diptera Trogloxene GA ○ ○ ○
Trichocera hiemalis (De Geer, 1776) Winterpluimantennemug Diptera Trogloxene GNR ○ ○ ○
Trichocera maculipennis Meigen, 1818 Diptera Trogloxene GNR ○ ○ ○
Trichocera regelationis (Linnaeus, 1758) Dooimug Diptera Trogloxene GNR IA, IL ○ ○ ○
Trichodrilus comalensis Fend, 2023 Lumbriculida Stygobiont TX ○ 6 ○
Trichodrilus culveri Cook, 1975 Lumbriculida Stygobiont G1 WV ○ 2 ○
Trichomorpha capillata Loomis, 1964 Polydesmida Trogloxene ○ ○ ○
Trichoniscus provisorius Racovitza, 1908 Ansarisiiranen Isopoda Troglophile VA ○ ○ ○
Trichoniscus pusillus Brandt, 1833 Common Pygmy Woodlouse Isopoda Troglophile KY, OK, TN, VA ○ ○ ○
Trichopetalum dux (Chamberlin, 1940) Chordeumatida Troglophile G4 KY ○ ○ ○
Trichopetalum stannardi (Causey, 1951) Chordeumatida Troglophile MD ○ ○ ○
Trichopetalum uncum Cook & Collins, 1895 Chordeumatida Troglobiont GNR IN, AR, MO ○ ○ ○
Trichophya pilicornis (Gyllenhal, 1810) Staph Beetle Coleoptera Trogloxene GNR MN ○ ○ ○
Trichotichnus dichrous (Dejean, 1829) Coleoptera Accidental GNR AL ○ ○ ○
Trichotichnus fulgens (Csiki, 1932) Coleoptera Accidental MO ○ ○ ○
Tridenchthonius juxtlahuaca J. C. Chamberlin & R. V. Chamberlin, 1945 Pseudoscorpiones Troglophile ○ ○ ○
Trigenotyla blacki Shear, 2003 Black's cave millipede Chordeumatida Troglobiont G1 OK ○ ○ ○
Trigenotyla parca Causey, 1951 Chordeumatida Troglophile G2 AR ○ ○ ○
Trimiomelba dubia (LeConte, J. L., 1849) Coleoptera Troglophile GNR TN ○ ○ ○
Trinectes maculatus (Bloch & Schneider, 1801) Freshwater Flounder Carangiformes Stygoxene G5 LC FL ○ ○ ○
Triodopsis complanata (Pilsbry, 1898) Glossy Threetooth Stylommatophora Trogloxene G2 KY, TN ○ ○ ○
Triodopsis hopetonensis (Shuttleworth, 1852) Magnolia Threetooth Stylommatophora Trogloxene G4 FL ○ ○ ○
Triodopsis tridentata (Say, 1817) Northern Threetooth Stylommatophora Trogloxene G5 AL ○ ○ ○
Triodopsis vulgata Pilsbry, 1940 Dished Threetooth Stylommatophora Trogloxene G5 IL ○ ○ ○
Triphosa haesitata (Guenée, [1858]) Tissue Moth Lepidoptera Trogloxene G5 ○ ○ ○
Trisetobisium fallax (J. C. Chamberlin, 1962) Pseudoscorpiones Troglophile GNR AL ○ ○ ○
Trochosa terricola Thorell, 1856 Gewone Nachtwolfspin Araneae Accidental G5 ○ ○ ○
Troglichthys rosae (Eigenmann, 1898) Ozark cavefish Percopsiformes Stygobiont G3 NT ESA AR, MO, OK ○ 4 ○
Troglocambarus maclanei Hobbs, 1942 North Florida Spider Cave Crayfish Decapoda Stygobiont G1G2 NT FL ○ 42 6
Trogloglanis pattersoni Eigenmann, 1919 Toothless Blindcat Siluriformes Stygobiont G1G2 VU ESA? TX ○ 104 4
Trombicula cavernarum Ewing, 1933 Trombidiformes Troglophile ○ ○ ○
Trombicula tibbettsi Brennan & White, 1960 Trombidiformes Trogloxene AL, TX ○ ○ ○
Tropisternus collaris (Fabricius, 1775) Collared Water Scavenger Beetle Coleoptera Accidental GNR AR ○ ○ ○
Tropocyclops extensus Kiefer, 1931 Cyclopoida Stygoxene IN ○ ○ ○
Tropocyclops prasinus (Fischer, 1860) Cyclopoida Stygoxene IL, IN, KY ○ ○ ○
Trox aequalis Say, 1832 Even Hide Scarab Beetle Coleoptera Trogloxene G5 GA ○ ○ ○
Trox laticollis LeConte, 1854 Coleoptera Trogloxene GNR IA ○ ○ ○
Trox scaber (Linnaeus, 1767) Hide Beetle Coleoptera Trogloxene GNR ○ ○ ○
Tuberochernes aalbui Muchmore, 1997 Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Tuberochernes cohni Harvey & Wynne, 2014 Pseudoscorpiones Troglobiont GNR AZ ○ ○ ○
Tuberochernes ubicki Muchmore, 1997 Pseudoscorpiones Troglobiont G1 AZ ○ 1 ○
Tuberona leonidasi (Kulkoyluoglu et al., 2023) Podocopida Stygobiont TX ○ ○ ○
Tullbergia clavata Wills, 1934 Poduromorpha Troglophile IA, IL ○ ○ ○
Tullbergia granulata Mills, HB, 1934 Poduromorpha Troglophile IA ○ ○ ○
Tullbergia macrochaeta (Rusek, 1976) Poduromorpha Accidental IA ○ ○ ○
Tychobythinus bythinioides (Brendel, 1865) Coleoptera Troglophile GNR IL ○ ○ ○
Tychobythinus hubrichti (Park, 1960) Coleoptera Troglobiont G1G2 KY ○ 1 ○
Tychobythinus jonesi (Park, 1951) Coleoptera Troglobiont G1G2 AL ○ ○ ○
Tychobythinus strinatii Besuchet, 1982 Coleoptera Troglobiont G1G2 TN ○ ○ ○
Tylparua troglodytes Evenhuis, 2024 Diptera Troglobiont HI ○ ○ ○
Typhlichthys eigenmanni Charlton, 1933 Salem Plateau Cavefish Percopsiformes Stygobiont G3 AR, MO ○ 33 148
Typhlichthys subterraneus Girard, 1859 Southern Cavefish Percopsiformes Stygobiont G4 NT AL, GA, KY, TN ○ 190 716
Typhlobius caecus Bollman (1888) Lithobiomorpha Troglophile AL, GA ○ ○ ○
Typhloelmis caroline Barr et al., 2015 Coleoptera Stygobiont GNR TX ○ 3 ○
Typhloelmis finegan Barr et al., 2015 Coleoptera Stygobiont GNR TX ○ 3 ○
Typhloelmis sanfelipe Barr et al., 2015 Coleoptera Stygobiont GNR TX ○ 3 ○
Typhloelmis spangleri Barr, 2023 Coleoptera Stygobiont GNR TX ○ ○ ○
Typhlogastrura alabamensis (Thibaud, 1975) Poduromorpha Troglobiont G1G2 AL ○ ○ ○
Typhlogastrura christianseni Thibaud, 1975 Poduromorpha Troglobiont G1 AL, TN ○ ○ ○
Typhlogastrura fousheensis Christiansen & Wang, 2006 Foushee Cave Springtail Poduromorpha Troglobiont G1 AR ○ 1 ○
Typhlogastrura helleri Christiansen & Wang, 2006 Heller Cave Springtail Poduromorpha Troglobiont GNR PA ○ 1 ○
Typhlogastrura hubbardi Thibaud, 1995 Poduromorpha Troglobiont G2G3 VA ○ ○ ○
Typhlogastrura valentini Thibaud, 1996 Valentine's Cave Springtail Poduromorpha Troglobiont G1 VA ○ ○ ○
Typhlonesticus silvestrii (Fage, 1929) Silvestri's Cave Spider Araneae Troglophile CA, OR ○ ○ ○
Tyrannochthonius alabamensis Muchmore, 1996 Pseudoscorpiones Troglophile G1 AL ○ ○ ○
Tyrannochthonius aladdinensis Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius aralu Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius archeri Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius attenuatus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius avernicolus Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius barri Muchmore, 1996 Barr's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius binoculatus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius chamberlini Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius charon Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius diabolus Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius erebicus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius felix Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius fiskei Muchmore, 1996 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Tyrannochthonius gnomus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius halopotamus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL, TN ○ ○ ○
Tyrannochthonius hypogeus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 KY ○ ○ ○
Tyrannochthonius infernalis Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius jonesi Chamberlin, 1995 Jones' Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius muchmoreorum Cokendolpher, 2009 Pseudoscorpiones Troglobiont GNR TX ○ ○ ○
Tyrannochthonius nergal Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius orpheus Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius osiris Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius parvus Chamberlin, 1995 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius pecki Muchmore, 1996 Peck's Cave Peudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius pholeter Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius pluto Chamberlin, 1995 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius pupukeanus Muchmore, 1983 Pseudoscorpiones Troglobiont G1 HI ○ 1 ○
Tyrannochthonius satan Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius sheltae Muchmore, 1996 Shelta Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius skeletonis Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius steevesi Muchmore, 1996 Steeves' Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Tyrannochthonius stonei Muchmore, 1989 Pseudoscorpiones Troglobiont HI ○ 1 ○
Tyrannochthonius stygius Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius tartarus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius tenuis Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius torodei Muchmore, 1996 Torode's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius troglodytes Muchmore, 1986 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Ufocandona hannaleeae Kulkoyluoglu et al., 2017 Podocopida Stygobiont TX ○ ○ ○
Ula elegans Osten Sacken, 1869 Diptera Trogloxene GNR ○ ○ ○
Uloborus glomosus (Walckenaer, 1841) Featherlegged Orbweaver Araneae Trogloxene GNR AL, NM ○ ○ ○
Ulomorpha pilosella (Osten Sacken, 1860) Diptera Trogloxene GNR KY ○ ○ ○
Uncinocythere ambophora (Walton & Hobbs, 1959) Podocopida Stygobiont G2G3 FL ○ 10 ○
Uncinocythere equicurva (Hoff, 1944) Podocopida Stygophile ○ ○ ○
Uncinocythere pholetera (Hart & Hobbs, 1961) Podocopida Stygobiont G1G2 MO ○ 14 ○
Uncinocythere simondsi (Hobbs & Walton, 1960) Podocopida Stygophile KY ○ ○ ○
Uncinocythere warreni Hobbs & Walton, 1968 Podocopida Stygobiont G1 GA, AR, FL ○ 6 ○
Uncinocythere xania (Hart & Hobbs, 1961) Podocopida Stygobiont G3 IN, MO, AR, GA ○ 21 ○
Uncinocythere zancla Hobbs & Walton, 1963 Podocopida Stygophile KY ○ ○ ○
Uniomerus carolinianus (Bosc, 1801) Eastern Pondhorn Unionoida Accidental G4 FL ○ ○ ○
Uroblaniulus canadensis (Newport, 1844) Sickle Tailed Millipede Julida Accidental G5 ○ ○ ○
Uroctonus grahami Gertsch & Soleglad, 1972 Scorpiones Troglobiont G1G2 CA ○ ○ 1
Uroseius infirmus (Berlese, 1887) Mesostigmata Trogloxene TN ○ ○ ○
Ursus americanus Pallas, 1780 American Black Bear Carnivora Trogloxene G5 LC AR, IN, KY, TN ○ ○ ○
Ursus arctos Linnaeus, 1758 Brown Bear Carnivora Trogloxene G4 LC ○ ○ ○
Usofila gracilis Keyserling, 1891 Araneae Troglobiont G1G2 CA ○ ○ ○
Uvarus texanus (Sharp, 1882) Coleoptera Stygoxene TX ○ ○ ○
Vachonium boneti Chamberlin, 1947 Pseudoscorpiones Troglophile ○ ○ ○
Vachonium maya Chamberlin, 1947 Pseudoscorpiones Troglophile ○ ○ ○
Vaejovis carolinianus (Beauvois, 1805) Southern Unstriped Scorpion Scorpiones Accidental G5 AL ○ ○ ○
Vallonia pulchella (O. F. Müller, 1774) Beautiful Grass Snail Stylommatophora Trogloxene G5 KY ○ ○ ○
Varacosa avara (Keyserling, 1877) Spurred Secretive Wolf Spider Araneae Accidental GNR IL ○ ○ ○
Veigaia bakeri Farrier, 1957 Baker's Cave Mite Mesostigmata Troglobiont G1 IN, NM ○ ○ ○
Veigaia kochi (Trägårdh, 1901) Mesostigmata Troglophile GNR ○ ○ ○
Veigaia wyandottensis (Packard, 1888) Wyandotte Cave Mite Mesostigmata Troglobiont G1 IN ○ ○ ○
Ventridens demissus (A. Binney, 1843) Perforate Dome Stylommatophora Trogloxene G5 FL, TN ○ ○ ○
Ventridens gularis (Say, 1821) Throaty Dome Stylommatophora Troglophile G5 AL, GA, KY, TN ○ ○ ○
Ventridens intertextus (A. Binney, 1840) Pyramid Dome Stylommatophora Trogloxene G5 IL ○ ○ ○
Ventridens ligera (Say, 1821) Globose Dome Stylommatophora Trogloxene G5 GA, KY ○ ○ ○
Vetericaris chaceorum Kensley & Williams, 1986 Decapoda Stygobiont G1 ESA HI ○ 4 ○
Virginia valeriae Baird & Girard, 1853 Smooth Earth Snake Squamata Accidental G5 LC AR ○ ○ ○
Vitronura giselae Cassagnau & Palacios-Vargas, 1983 Poduromorpha Trogloxene IL, TN ○ ○ ○
Vitropyrgus lillianae Perez & Castañeda, 2023 Littorinimorpha Stygobiont TX ○ ○ 6
Vorticella abreviata Keiser, 1921 Sessilida Stygoxene AL ○ ○ ○
Vorticella microstoma Ehrenberg, 1830 Sessilida Stygoxene KY ○ ○ ○
Vulcanochthonius aa Muchmore, 2000 Pseudoscorpiones Troglobiont HI ○ 1 ○
Vulcanochthonius howarthi (Muchmore, 1979) Pseudoscorpiones Troglobiont G1G2 HI ○ 1 ○
Vulcanochthonius pohakuloae Muchmore, 2000 Pseudoscorpiones Troglobiont HI ○ 1 ○
Wadotes calcaratus (Keyserling, 1887) Araneae Trogloxene G5 GA, TN ○ ○ ○
Wadotes saturnus Bennett, 1987 Araneae Trogloxene GA ○ ○ ○
Watobius anderisus Chamberlin, 1911 Lithobiomorpha Accidental AL ○ ○ ○
Wormaldia moesta (Banks, 1914) Trichoptera Stygoxene G4 KY ○ ○ ○
Wulfila albens (Hentz, 1847) Araneae Accidental AL ○ ○ ○
Xenolimosina sicula Marshall, 1985 Diptera Trogloxene GNR CO ○ ○ ○
Xenotrechus condei Barr & Krekeler, 1967 Coleoptera Troglobiont G1G2 MO ○ ○ ○
Xenotrechus denticollis Barr & Krekeler, 1967 Coleoptera Troglobiont G1G2 MO ○ 2 9
Xenylla welchi Folsom, JW, 1916 Springtail Poduromorpha Trogloxene GNR IL ○ ○ ○
Xironodrilus formosus Ellis, 1918 Branchiobdellida Stygophile TN ○ ○ ○
Xironogiton instabilius (Moore, 1894) Branchiobdellida Stygophile VA, WV ○ ○ ○
Xolotrema denotatum (A. Férussac, 1823) Velvet Wedge Stylommatophora Trogloxene G4 IL ○ ○ ○
Xolotrema fosteri (F. C. Baker, 1932) Bladetooth Wedge Stylommatophora Trogloxene G5 IL ○ ○ ○
Xolotrema obstrictum (Say, 1821) Sharp Wedge Stylommatophora Trogloxene G4 TN ○ ○ ○
Yunohamella lyrica (Walckenaer, 1841) Lyric Cobweaver Araneae Trogloxene GNR AL, IN ○ ○ ○
Zealeuctra narfi Ricker & Ross, 1969 Northern Needlefly Plecoptera Stygoxene G4 IL ○ ○ ○
Zonitoides arboreus (Say, 1817) Bark Snail Stylommatophora Troglophile AL, FL, GA, IL, KY, MO, TN, VA ○ ○ ○
Zonitoides limatulus (A. Binney, 1840) Dull Gloss Stylommatophora Trogloxene G4 KY ○ ○ ○
Zonitoides nitidus (O. F. Müller, 1774) Black Gloss Stylommatophora Trogloxene IL ○ ○ ○
Zosteractis interminata Loomis, 1943 Zosteractis Cave Millipede Julida Troglobiont G1 IL, MO ○ 2 ○
Zygonopus krekeleri Causey, 1960 West Virginia Blind Cave Millipede Chordeumatida Troglobiont G4 WV ○ ○ ○
Zygonopus packardi Causey, 1960 Packard's Blind Cave Millipede Chordeumatida Troglobiont G4 VA, WV ○ ○ ○
Zygonopus weyeriensis Causey, 1960 Grand Caverns Blind Cave Millipede Chordeumatida Troglobiont G3G4 VA, WV ○ ○ ○
Zygonopus whitei Causey, 1960 Luray Caverns Blind Cave Millipede Chordeumatida Troglobiont G3G4 VA, WV ○ ○ ○

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