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Subterranean biodiversity of North America and around the globe

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Cave Biology Data Portal
Subterranean biodiversity of North America and around the globe

Browse species

1445 species

Species Common name Group Class Status States Photo Spec Seq
Larca aalbui (Muchmore, 1984) Aalbu's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Agastoschizomus texanus Monjarez-Ruedas et al., 2016 Schizomida Troglobiont TX ○ 3 ○
Agyneta gagnei (Gertsch, 1973) Araneae Troglobiont G1G2 HI ○ 1 ○
Agyneta hedini Duperre & Paquin, 2009 Araneae Troglobiont CO ○ 3 ○
Stygobromus alabamensis (Stout, 1911) Alabama cave amphipod Amphipoda Stygobiont G5 AL, AR, KS, LA, MO, OK, TN, TX ○ 207 ○
Pseudanophthalmus alabamae Valentine, 1932 Alabama Cave Beetle Coleoptera Troglobiont G2 AL ○ 8 ○
Spinanycta alabamensis Espinasa, Gutierrez, Hinkle & Niemiller, 2024 Alabama Cave Bristletail Zygentoma Troglobiont AL ○ ○ ○
Cambarus jonesi Hobbs & Barr, 1960 Alabama Cave Crayfish Decapoda Stygobiont G2 VU AL ○ 70 36
Scoterpes alabama Shear, 2010 Alabama Cave Millipede Chordeumatida Troglobiont AL ○ ○ ○
Palaemonias alabamae Smalley, 1961 Alabama Cave Shrimp Decapoda Stygobiont G1 EN ESA AL ○ 5 14
Speoplatyrhinus poulsoni Cooper & Kuehne, 1974 Alabama cavefish Percopsiformes Stygobiont G1 CR ESA AL ○ 9 27
Alabamocreagris mortis (Muchmore, 1969) Alabama pseudoscorpion Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Stygobromus smithi Hubricht, 1943 Alabama Well Amphipod Amphipoda Stygobiont G2G3 VU AL ○ 3 ○
Alabamocreagris pecki (Muchmore, 1969) Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Banksula californica Briggs, 1974 Alabaster Cave Harvestman Opiliones Troglobiont GH CA ○ 3 ○
Albiorix anophthalmus Muchmore & Pape, 1998 Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Chaetaspis aleyorum Lewis, 2002 Aley's Cave Milliped Polydesmida Troglobiont G1 MO ○ ○ ○
Pseudanophthalmus alladini Valentine, 1945 Alladin's Cave Beetle Coleoptera Troglobiont G3G4 AL ○ ○ ○
Caecidotea alleghenyensis Lewis, Bowman, & Feller, 2011 Allegheny Spring Isopod Isopoda Stygobiont G1G2 MD ○ 18 ○
Allomideopsis wichitaensis (Smith, 1990) Trombidiformes Stygobiont TX ○ ○ ○
Allotexiweckelia hirsuta Holsinger, 1980 Amphipoda Stygobiont G2G3 TX ○ 43 ○
Almuerzothyas comalensis Smith & Cook, 2009 Trombidiformes Stygobiont GNR TX ○ ○ ○
Ameractis satis Causey, 1959 Julida Troglobiont G2G4 AL, GA, TN, VA ○ 1 ○
Caecidotea putea Lewis, 2009 American Waterslaters Isopoda Stygobiont G1G3 FL, GA ○ ○ ○
Americocreagris columbiana (Chamberlin, 1962) Pseudoscorpiones Troglobiont G2G3 OR, WA ○ ○ ○
Amerigoniscus centralis Vandel, 1978 Isopoda Troglobiont G1G2 OK ○ ○ ○
Amerigoniscus curvatus Vandel, 1978 Isopoda Troglobiont G1 GA ○ ○ ○
Amerigoniscus georgiensis Vandel, 1978 Isopoda Troglobiont G1 GA ○ ○ ○
Amerigoniscus gipsocolus (Vandel, 1965) Isopoda Troglobiont G1G2 TX ○ 1 ○
Amerigoniscus malheurensis Schultz, 1982 Isopoda Troglobiont G1 OR ○ 2 ○
Amerigoniscus nicholasi (Vandel, 1965) Isopoda Troglobiont G1G2 GA, TN ○ 4 ○
Amerigoniscus proximus Vandel, 1978 Isopoda Troglobiont G1G2 GA ○ ○ ○
Ameroduvalius jeanneli Valentine, 1952 Coleoptera Troglobiont G3 KY ○ 8 6
Amphibiocapillaria texensis Moravec & Huffman, 2000 Trichocephalida Stygobiont TX ○ ○ ○
Amplaria flucticulus Shear, 2021 Chordeumatida Troglobiont CA ○ ○ ○
Amplaria shastae (Causey, 1958) Chordeumatida Troglobiont G1G2 CA ○ 1 ○
Nesidiolestes ana Gagne & Howarth, 1974 Ana Wingless Thread Bug Heteroptera Troglobiont G1G2 NE HI ○ 1 ○
Anachemmis jungi Platnick & Ubick, 2005 Araneae Troglobiont CA ○ ○ ○
Anillaspis explanata (Horn, 1888) Coleoptera Troglobiont CA ○ 1 ○
Anillinus actuipennis Sokolov & Reddell, 2014 Coleoptera Troglobiont TX ○ ○ ○
Anillinus bexarensis Sokolov, 2022 Coleoptera Troglobiont TX ○ ○ ○
Anillinus cavicola Sokolov, 2012 Coleoptera Troglobiont AL ○ ○ ○
Anillinus clinei Sokolov, 2020 Coleoptera Troglobiont AL ○ ○ ○
Anillinus forthoodensis Sokolov & Reddell, 2014 Coleoptera Troglobiont TX ○ 4 ○
Anillinus hildebrandti Sokolov, 2020 Coleoptera Troglobiont AL ○ ○ ○
Anillinus longiceps Jeannel, 1963 Coleoptera Troglobiont TN ○ 2 ○
Anillinus reddelli Sokolov, 2022 Coleoptera Troglobiont TX ○ ○ ○
Anillinus smokiensis Sokolov, 2011 Coleoptera Troglobiont TN ○ ○ 13
Anillinus tombarri Sokolov, 2012 Coleoptera Troglobiont AL ○ ○ ○
Anillinus valentinei (Jeannel, 1963) Coleoptera Troglobiont G3G4 AL ○ 2 ○
Anillinus wisemanensis Sokolov & Kavanaugh, 2014 Coleoptera Troglobiont TX ○ 4 ○
Anisolabis howarthi Brindle, 1980 Dermaptera Troglobiont G1G2 HI ○ 1 ○
Anthrobia coylei Miller, 2005 Araneae Troglobiont G2 VA, WV ○ ○ ○
Antrolana lira Bowman 1964 Isopoda Stygobiont G2G4 VU ESA VA, WV ○ 37 17
Aphrastochthonius pachysetus Muchmore, 1976 Pseudoscorpiones Troglobiont G1G2 NM ○ ○ ○
Aphrastochthonius pecki Muchmore, 1968 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Aphrastochthonius tenax Chamberlin, 1962 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Apocheiridium reddelli Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ 3
Apochthonius coecus (Packard, 1884) Pseudoscorpiones Troglobiont G1G2 VA ○ 2 ○
Apochthonius colecampi Muchmore, 1967 Pseudoscorpiones Troglobiont G1G2 MO ○ ○ ○
Apochthonius diabolus Muchmore, 1967 Pseudoscorpiones Troglobiont G1 AR ○ ○ ○
Apochthonius forbesi Benedict, 1979 Pseudoscorpiones Troglobiont G1G2 OR ○ ○ ○
Apochthonius grubbsi Muchmore, 1980 Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Apochthonius holsingeri Muchmore, 1967 Pseudoscorpiones Troglobiont G1G2 VA ○ ○ ○
Apochthonius minor Muchmore, 1976 Pseudoscorpiones Troglobiont G1 GA, TN ○ ○ ○
Apochthonius titanicus Muchmore, 1976 Pseudoscorpiones Troglobiont G2 AR ○ ○ ○
Orconectes packardi Rhoades, 1944 Appalachian Cave Crayfish Decapoda Stygobiont G2 EN KY ○ 34 33
Porrhomma cavernicola (Keyserling, 1886) Appalachian Cave Spider Araneae Troglobiont G5 AL, AR, GA, IA, IL, IN, KY, MD, MO, NC, OK, ON, PA, TN, VA, WV ○ 14 ○
Tolus appalachius Goodnight & Goodnight, 1942 Appalachian Cave Spider Opiliones Troglobiont G3G4 AL, TN ○ 2 4
Fontigens bottimeri (Walker, 1925) Appalachian Springsnail Littorinimorpha Stygobiont G2G3 DC, MD, VA ○ 68 2
Crangonyx antennatus Packard, 1881 Appalachian Valley crangonyctid Amphipoda Stygobiont G5 AL, GA, TN, VA ○ 222 1
Appaleptoneta barrowsi (Gertsch, 1974) Araneae Troglobiont G1 AL ○ 1 2
Appaleptoneta credula (Gertsch, 1974) Araneae Troglobiont G1 AL ○ 5 2
Appaleptoneta fiskei (Gertsch, 1974) Araneae Troglobiont GNR GA ○ 13 3
Appaleptoneta jonesi (Gertsch, 1974) Araneae Troglobiont G1 AL ○ ○ ○
Horologion speokites Valentine, 1932 Arbuckle Cave Beetle Coleoptera Troglobiont GH WV ○ ○ ○
Liocranoides archeri Platnick, 1999 Archer's Two-clawed Cave Spider Araneae Troglobiont G2 AL, TN ○ ○ 6
Arianops cavernensis Park, 1951 Coleoptera Troglobiont GNR AL ○ 7 ○
Arianops gorei Carlton, 2008 Coleoptera Troglobiont TN ○ 1 ○
Arianops jeanneli Park, 1956 Coleoptera Troglobiont G1 VA ○ ○ ○
Arianops pecki Barr, 1974 Coleoptera Troglobiont TN ○ 1 ○
Arianops steevesi Barr, 1974 Coleoptera Troglobiont GNR AL ○ 1 ○
Arianops stygica Park, 1960 Coleoptera Troglobiont G1 TN ○ ○ ○
Stygobromus arizonensis Holsinger, 1974 Arizona Cave Amphipod Amphipoda Stygobiont G1 VU AZ ○ 5 ○
Artesia subterranea Holsinger, 1980 Amphipoda Stygobiont G1G2 TX ○ 94 ○
Artesia welbourni Holsinger, 1992 Amphipoda Stygobiont G1G2 TX ○ 2 ○
Brackenridgia ashleyi Lewis, 2004 Ashley's Terrestrial Cave Isopod Isopoda Troglobiont G1G3 AR, MO ○ 3 ○
Eurycea waterlooensis Hillis et al., 2001 Austin Blind Salamander Caudata Stygobiont G1 CR ESA TX ○ 14 5
Australinocreagris grahami (Muchmore, 1969) Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Austrotyla awishoshola Wynne & Shear, 2016 Chordeumatida Troglobiont NM ○ ○ ○
Austrotyla coloradensis (Chamberlin, 1910) Chordeumatida Troglobiont CO ○ 47 ○
Austrotyla stephensoni (Chamberlin, 1910) Chordeumatida Troglobiont CO ○ 1 ○
Pseudanophthalmus avernus (Valentine, 1945) Avernus Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Sinella avita Christiansen, 1960 Avita Cave Springtail Entomobryomorpha Troglobiont G3G4 IL, IN, KY, MO ○ 3 ○
Axonopsella bakeri Smith & Cook, 1998 Trombidiformes Stygobiont FL, TX ○ ○ ○
Bactrurus brachycaudus Hubricht & Mackin, 1940 Amphipoda Stygobiont G4 IL, MO ○ 98 8
Bactrurus cellulanus Koenemann & Holsinger, 2001 Amphipoda Stygobiont GX IN ○ 4 ○
Bactrurus mucronatus (Forbes, 1876) Amphipoda Stygobiont G5 IA, IL, IN, MI, OH ○ 126 6
Bactrurus wilsoni Koenemann & Holsinger, 2001 Amphipoda Stygobiont G1G2 AL ○ 4 ○
Pseudanophthalmus insularis Barr, 1959 Baker Station Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Veigaia bakeri Farrier, 1957 Baker's Cave Mite Mesostigmata Troglobiont G1 IN ○ ○ ○
Stygobromus balconis (Hubricht, 1943) Balcones Cave Amphipod Amphipoda Stygobiont G2G3 VU TX ○ 28 ○
Palaemonetes antrorum Benedict, 1896 Balcones Cave Shrimp Decapoda Stygobiont G2G3 EN TX ○ ○ ○
Balconorbis uvaldensis Hershler & Longley, 1986 Balcones ghostsnail Littorinimorpha Stygobiont G1G2 TX ○ 47 9
Cicurina bandida Gertsch, 1992 Bandit Cave Meshweaver Araneae Troglobiont G2 TX ○ 12 ○
Banksula grahami Briggs, 1974 Opiliones Troglobiont G2G3 CA ○ 1 ○
Crangonyx barri Zhang & Holsinger, 2003 Barr's Cave Amphipod Amphipoda Stygobiont G5 IN, KY ○ 141 ○
Stygobromus barri (Holsinger, 1967) Barr's Cave Amphipod Amphipoda Stygobiont G2G3 VU MO ○ 8 ○
Pseudotremia barri Lewis, 2005 Barr's Cave Millipede Chordeumatida Troglobiont G2 TN ○ ○ 9
Scoterpes tombarri Shear, 2010 Barr's Cave Millipede Chordeumatida Troglobiont KY, TN ○ ○ ○
Tyrannochthonius barri Muchmore, 1996 Barr's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Nesticus barri Gertsch, 1984 Barr's Cave Spider Araneae Troglobiont G3 AL, TN ○ 4 57
Sinella barri Christiansen, 1960 Barr's Cave Springtail Entomobryomorpha Troglobiont G5 AL, AR, IL, IN, KY, MO, TN, VA, WI ○ 6 ○
Miktoniscus barri Vandel, 1965 Barr's Terrestrial Cave Isopod Isopoda Troglobiont G2G4 IN, KY, TN ○ 4 ○
Stygopyrgus bartonensis Hershler & Longley, 1986 Barton cavesnail Littorinimorpha Stygobiont G1 TX ○ 30 15
Caecidotea macropropoda Chase & Blair, 1937 Bat Cave Isopod Isopoda Stygobiont G3 VU AR, OK ○ 27 ○
Stygobromus mundus (Holsinger, 1967) Bath County Cave Amphipod Amphipoda Stygobiont G1 VU VA ○ 7 1
Bathynella fraterna Cho & Kim, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Bathynella germanitas Cho & Kim, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Bathynella riparia Pennak & Ward, 1985 Bathynellacea Stygobiont CO ○ 1 ○
Batriasymmodes greeveri Park, 1965 Coleoptera Troglobiont G1G2 TN ○ 2 ○
Batriasymmodes parki Barr, 1987 Coleoptera Troglobiont GH WV ○ 1 ○
Batriasymmodes quisnamus (Park, O., 1951) Coleoptera Troglobiont G3 KY, TN ○ 3 ○
Batriasymmodes spelaeus (Park, 1951) Coleoptera Troglobiont G3G4 AL, GA, TN ○ ○ 2
Batriasymmodes troglodytes Park, 1951 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Batrisodes barri Park, 1958 Coleoptera Troglobiont G1G2 TN ○ 2 ○
Batrisodes cavernosus Park, 1951 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes clypeospecus Park, 1960 Coleoptera Troglobiont G1G2 TN ○ ○ ○
Batrisodes dentifrons Chandler & Reddell, 2009 Coleoptera Troglobiont G1G2 TX ○ 1 ○
Batrisodes fanti Chandler & Reddell, 2009 Coleoptera Troglobiont G1G2 TX ○ 1 ○
Batrisodes feminiclypeus Chandler & Reddell, 2001 Coleoptera Troglobiont G1G2 TX ○ 3 ○
Batrisodes ferulifer Park, 1960 Coleoptera Troglobiont G1 TN ○ ○ ○
Batrisodes gemmoides Park, 1960 Coleoptera Troglobiont G1 TN ○ ○ ○
Batrisodes gemmus Park, 1956 Coleoptera Troglobiont G1 TN ○ ○ ○
Batrisodes gravesi Chandler & Reddell, 2001 Coleoptera Troglobiont G3 TX ○ 14 ○
Batrisodes grubbsi Chandler, 1992 Coleoptera Troglobiont G1G2 TX ○ 3 ○
Batrisodes henroti Park, 1956 Coleoptera Troglobiont G2G3 KY ○ 2 ○
Batrisodes hubrichti Park, 1958 Coleoptera Troglobiont G1 KY ○ 2 ○
Batrisodes incispes Chandler & Reddell, 2009 Coleoptera Troglobiont G1 TX ○ ○ ○
Batrisodes jocuvestus Park, 1960 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes jonesi Park, O., 1951 Coleoptera Troglobiont G2G3 AL ○ ○ ○
Batrisodes krekeleri Park, 1960 Coleoptera Troglobiont IN ○ ○ ○
Batrisodes pannosus Park, 1960 Coleoptera Troglobiont G1 TN ○ 2 ○
Batrisodes pekinsi Chandler & Reddell, 2009 Coleoptera Troglobiont G1 TX ○ 2 ○
Batrisodes profundus Park, 1956 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Batrisodes reyesi Chandler, 1992 Coleoptera Troglobiont G2G3 TX ○ 25 ○
Batrisodes shadeae Chandler & Reddell, 2009 Coleoptera Troglobiont G1 TX ○ 1 ○
Batrisodes specus Park, 1951 Coleoptera Troglobiont G3G4 AL, TN ○ 4 ○
Batrisodes subterraneus Park, 1951 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes tumoris Park, 1960 Coleoptera Troglobiont G1 AL ○ ○ ○
Batrisodes valentinei Park, 1951 Coleoptera Troglobiont G2G4 AL, TN ○ ○ ○
Batrisodes wartoni Chandler & Reddell, 2001 Coleoptera Troglobiont G1G2 TX ○ 1 ○
Phreatodrobia rotunda Hershler & Longley, 1986 Beaked cavesnail Littorinimorpha Stygobiont G1G2 DD TX ○ 9 8
Pseudanophthalmus beakleyi Valentine, 1937 Beakley's Cave Beetle Coleoptera Troglobiont G3G4 KY, TN ○ ○ ○
Caecidotea beattyi Lewis & Bowman, 1981 Beatty's Groundwater Isopod Isopoda Stygobiont G3G4 IL, IN, MO ○ 16 13
Oreonetides beattyi Paquin et al., 2009 Beatty's Sheetweb Cave Spider Araneae Troglobiont GNR IN, MD, PA, TN, VA ○ ○ ○
Pseudanophthalmus major Krekeler, 1973 Beaver Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Belba bulbipedata (Packard, 1888) Oribatida Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus bendermani Barr, 1959 Benderman's Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Caecidotea carolinensis (Lewis & Bowman, 1977 Bennett's Mill Cave Isopod Isopoda Stygobiont G2G3 NC, SC ○ 4 ○
Cambarus aculabrum Hobbs & Brown, 1987 Benton County Cave Crayfish Decapoda Stygobiont G1 CR ESA AR, MO ○ 11 4
Gyrinophilus gulolineatus Brandon, 1965 Berry cave salamander Caudata Stygobiont G1 EN TN ○ 31 34
Bicornucandona fineganensis Kulkoyluoglu et al., 2011 Podocopida Stygobiont GNR TX ○ ○ ○
Stygobromus bifurcatus (Holsinger, 1967) Bifurcated Cave Amphipod Amphipoda Stygobiont G3G4 VU TX ○ 77 4
Procambarus horsti Hobbs & Means, 1972 Big Blue Spring Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 10 5
Procambarus delicatus Hobbs & Franz, 1986 Bigcheek Cave Crayfish Decapoda Stygobiont G1 CR FL ○ 3 2
Sphalloplana subtilis Kenk, 1977 Biggers Cave Flatworm Tricladida Stygobiont G1G2 NE VA ○ 3 ○
Stygobromus biggersi Holsinger, 1978 Biggers' Cave Amphipod Amphipoda Stygobiont G2G4 MD, PA, VA, WV ○ 22 1
Bishopella jonesi Goodnight & Goodnight, 1942 Opiliones Troglobiont G1 AL ○ ○ ○
Pygmarrhopalites ater (Christiansen & Bellinger, 1998) Black Medusa Cave Springtail Symphypleona Troglobiont G3 IL, IN ○ ○ ○
Pseudotremia blacki Lewis, 2000 Black's Cave Milliped Chordeumatida Troglobiont G1 IN ○ ○ ○
Trigenotyla blacki Shear, 2003 Black's cave millipede Chordeumatida Troglobiont G1 OK ○ ○ ○
Blackburnia aaae (Samuelson & Liebherr, 1992) Coleoptera Troglobiont G1G2 HI ○ ○ ○
Blackburnia howarthi (Samuelson & Liebherr, 1992) Coleoptera Troglobiont G1G2 HI ○ ○ ○
Eurycea robusta (Longley, 1978) Blanco Blind Salamander Caudata Stygobiont G1 DD TX ○ 1 ○
Blancosoma scaturgo Shear & Hubbard, 1998 Chordeumatida Troglobiont G1G2 CO ○ ○ ○
Glyphyalinia pecki Hubricht, 1966 Blind Glyph Stylommatophora Troglobiont G1G2 DD AL ○ 5 ○
Stygobromus blinni Wang & Holsinger, 2001 Blinn's Cave Amphipod Amphipoda Stygobiont G1G2 AZ ○ 2 ○
Pseudanophthalmus ventus Barr, 1981 Blowing Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Stygobromus spinosus (Hubricht & Mackin, 1940) Blue Ridge Cave Amphipod Amphipoda Stygobiont G3 VA ○ 16 1
Pseudotremia indianae Hoffman, 1958 Blue River Cave Millipede Chordeumatida Troglobiont G4 IN ○ 5 ○
Diacyclops conversus Reid, 2004 Blue River Groundwater Copepod Cyclopoida Stygobiont G2 IN ○ 3 ○
Pseudanophthalmus audax (Horn, 1883) Bold Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 1 ○
Conotyla bollmani (McNeill, 1887) Bollman's Cave Millipede Chordeumatida Troglobiont G3 IN ○ 1 ○
Texella reyesi Ubick & Briggs, 1992 Bone Cave Harvestman Opiliones Troglobiont G2G3 ESA TX ○ 210 3
Stygobromus limbus Wang & Holsinger, 2001 Border Cave Amphipod Amphipoda Stygobiont G1G2 TX ○ 1 ○
Stygobromus boultoni Wang & Holsinger, 2001 Boulton's Cave Amphipod Amphipoda Stygobiont G1G2 AZ ○ 4 ○
Stygobromus bowmani (Holsinger, 1967) Bowman's Cave Amphipod Amphipoda Stygobiont G1G2 VU OK ○ 3 ○
Brackenridgia cavernarum Ulrich, 1902 Isopoda Troglobiont G3G4 TX ○ 8 ○
Brackenridgia reddelli (Vandel, 1965) Isopoda Troglobiont G2G3 TX ○ 13 ○
Calileptoneta briggsi Ledford, 2004 Briggs' Cave Spider Araneae Troglobiont G1 CA ○ 1 ○
Cambarus setosus Faxon, 1889 Bristly Cave Crayfish Decapoda Stygobiont G3 NT AR, MO ○ 51 11
Sphalloplana buchanani (Hyman, 1937) Buchanan's Cave Flatworm Tricladida Stygobiont G1G2 KY, MD, TN ○ 10 ○
Apochthonius hobbsi Muchmore, 1994 Buckskin Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 OH ○ ○ ○
Pseudotremia burnsorum Lewis, 2000 Burn's Cave Milliped Chordeumatida Troglobiont G1G2 IN ○ ○ ○
Caconemobius albus Otte, 1994 Orthoptera Troglobiont HI ○ ○ ○
Caconemobius paralbus Otte, 1994 Orthoptera Troglobiont HI ○ ○ ○
Caconemobius uuku Otte, 1994 Orthoptera Troglobiont HI ○ 1 ○
Caecidotea acuticarpa Mackin & Hubricht, 1940 Isopoda Stygobiont G2G3 OK ○ 26 ○
Caecidotea adenta Mackin & Hubricht, 1940 Isopoda Stygobiont G1G2 OK ○ 3 ○
Caecidotea alabamensis Stafford, 1911 Isopoda Stygobiont G3G4 AL ○ 128 ○
Caecidotea ancyla (Fleming, 1972) Isopoda Stygobiont G3G4 AR, MO, OK ○ 34 ○
Caecidotea antricola Creaser, 1931 Isopoda Stygobiont G5 AR, MO, OK ○ 191 10
Caecidotea bilineata Lewis & Bowman, 1996 Isopoda Stygobiont G2G3 TX ○ 4 ○
Caecidotea catachaetus (Fleming & Steeves, 1972) Isopoda Stygobiont GA, TN ○ 10 ○
Caecidotea circulus (Steeves & Holsinger, 1968) Isopoda Stygobiont G1G2 TN ○ 3 ○
Caecidotea cyrtorhynchus (Fleming & Steeves, 1972) Isopoda Stygobiont G1 GA ○ 4 ○
Caecidotea dauphina Modlin, 1986 Isopoda Stygobiont AL ○ 2 ○
Caecidotea dimorpha (Mackin & Hubricht, 1940) Isopoda Stygobiont G2G3 AR, MO ○ 17 ○
Caecidotea extensolinguala (Fleming, 1972) Isopoda Stygobiont MO ○ 3 ○
Caecidotea fisherorum Lewis & Lewis, 2023 Isopoda Stygobiont GNR VA ○ ○ 15
Caecidotea fonticulus Lewis, 1983 Isopoda Stygobiont GNR AR ○ 5 ○
Caecidotea lesliei Lewis & Bowman, 1981 Isopoda Stygobiont G1G2 IL ○ 3 ○
Caecidotea mackini Lewis, Graening, Fenolio & Bergey, 2006 Isopoda Stygobiont GNR OK ○ 3 ○
Caecidotea metcalfi (Fleming, 1972) Isopoda Stygobiont G1G2 KS ○ 4 ○
Caecidotea ornatus Lewis & Lewis, 2023 Isopoda Stygobiont G1 VA ○ ○ 21
Caecidotea packardi Mackin & Hubricht, 1940 Isopoda Stygobiont G2G4 IL, MO ○ 34 10
Caecidotea paurotrigona (Fleming, 1972) Isopoda Stygobiont G1G2 LA, MS ○ 3 ○
Caecidotea reddelli (Steeves, 1968) Isopoda Stygobiont G3G4 TX ○ 267 11
Caecidotea scyphus (Steeves & Holsinger, 1968) Isopoda Stygobiont G1G2 TN, WV ○ ○ ○
Caecidotea simonini (Bresson, 1955) Isopoda Stygobiont G1 WV ○ 3 ○
Caecidotea simulator Lewis, 1999 Isopoda Stygobiont G2G3 AR, KS, OK ○ 7 ○
Caecidotea sinuncus (Steeves, 1965) Isopoda Stygobiont G1 WV ○ 3 ○
Caecidotea spatulata Mackin & Hubricht, 1940 Isopoda Stygobiont G3G4 IL, MO ○ 6 ○
Caecidotea teresae Lewis, 1982 Isopoda Stygobiont GX IN ○ 5 ○
Caecidotea tridentata Hungerford, 1922 Isopoda Stygobiont G1G2 IL, KS ○ 38 ○
Calasellus californicus (Miller, 1933) Isopoda Stygobiont G2 CA, OR ○ 20 1
Calasellus longus Bowman, 1981 Isopoda Stygobiont G1 CA ○ 3 ○
Califobathynella noodti Cho, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Califobathynella teucherti Cho, 1997 Bathynellacea Stygobiont CA ○ ○ ○
Sphalloplana californica Kenk, 1977 California Cave Flatworm Tricladida Stygobiont G1G2 CA ○ 4 ○
Californibathynella californica Schminke & Noodt, 1988 Bathynellacea Stygobiont CA ○ ○ ○
Pseudotremia callae Lewis, 2009 Call's Cave Milliped Chordeumatida Troglobiont G1 TN ○ ○ ○
Cambala speobia Chamberlin, 1953 Spirostreptida Troglobiont G4 TX ○ 1.3k ○
Cambarincola alienus Holt, 1963 Branchiobdellida Stygobiont G1G2 KY, TN ○ 9 ○
Cambarincola dubius Holt, 1973 Branchiobdellida Stygobiont G1G2 IN ○ 8 ○
Cambarincola leoni Holt, 1973 Branchiobdellida Stygobiont G3 FL ○ 13 ○
Cambarincola marthae Holt, 1973 Branchiobdellida Stygobiont G1G2 TN ○ 2 ○
Cambarincola sheltensis Holt, 1973 Branchiobdellida Stygobiont G1G2 AL ○ 12 ○
Cambarincola steevesi Holt, 1973 Branchiobdellida Stygobiont G3 KY ○ 6 ○
Faxonius stygocaneyi (Hobbs, 2001) Caney Mountain Cave Crayfish Decapoda Stygobiont G1 LC MO ○ 2 5
Caecidotea cannula (Steeves, 1963) Cannulate Cave Isopod Isopoda Stygobiont G1 WV ○ 12 ○
Stygobromus sextarius Holsinger, 2009 Capital Area Groundwater Amphipod Amphipoda Stygobiont G1 DC, MD, VA ○ 4 ○
Aphrastochthonius similis Muchmore, 1984 Carlow's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Carnarimelita janstocki Bousfield, 1990 Amphipoda Stygobiont HI ○ 6 ○
Stygobromus carolinensis Holsinger, 1978 Carolina Seep Amphipod Amphipoda Stygobiont GH NC ○ 4 ○
Pygmarrhopalites carolynae (Christiansen & Bellinger, 1996) Carolyn's Cave Springtail Symphypleona Troglobiont G4 IL, IN, VA, WV ○ ○ ○
Stygobromus dejectus (Holsinger, 1967) Cascade Cave Amphipod Amphipoda Stygobiont G1G2 VU TX ○ 4 7
Stygobromus canadensis Holsinger, 1980 Castleguard Cave Amphipod Amphipoda Stygobiont G1 AB ○ 2 ○
Pseudanophthalmus catherinae Barr, 1959 Catherine's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Catopocerus alabamae Peck, 1975 Coleoptera Troglobiont GNR AL ○ 1 ○
Catopocerus jonesi Peck, 1975 Coleoptera Troglobiont GNR AL ○ 1 ○
Causeyella causeyae Shear, 2003 Chordeumatida Troglobiont AR ○ ○ ○
Causeyella dendropus (Loomis, 1939) Causeyella Cave Millipede Chordeumatida Troglobiont G3 AR, MO ○ 1 ○
Causeyella youngsteadtorum Shear, 2003 Chordeumatida Troglobiont AR ○ ○ ○
Cavaticovelia aaa (Gagne & Howarth, 1975) Heteroptera Troglobiont G1G2 NE HI ○ 8 ○
Bactrurus speleopolis Holsinger, Sawicki, & Graening, 2006 Cave City Cave Amphipod Amphipoda Stygobiont G1 AR ○ 4 ○
Spelobia tenebrarum (Aldrich, 1897) Cave dung fly Diptera Troglobiont G5 AL, AR, GA, IL, IN, KY, MO, NY, OK, PA, TN, VA, WV ○ 1 ○
Physella spelunca Turner & Clench, 1974 Cave Physa Basommatophora Stygobiont G1 VU WY ○ 11 4
Islandiana cavealis Ivie, 1965 Cave Sheet-web Spider Araneae Troglobiont GNR IN ○ ○ ○
Islandiana lewisi Milne & Wells, 2018 cave sheet-web spider Araneae Troglobiont G1 IN ○ ○ ○
Nesticus jonesi Gertsch, 1984 Cave Spring Cave Spider Araneae Troglobiont G1 AL ○ 1 4
Carychium stygium Call, 1897 Cave thorn Basommatophora Troglobiont G3 AL, KY, TN ○ 72 6
Islandiana speophila Ivie, 1965 Cavern Sheet-web Spider Araneae Troglobiont G1 WV ○ ○ ○
Stygobromus caecilius Holsinger, 2011 Cecil Groundwater Amphipod Amphipoda Stygobiont G1 MD ○ 1 ○
Stygobromus exilis Hubricht, 1943 Central Kentucky Cave Amphipod Amphipoda Stygobiont G5 AL, KY, TN ○ 105 ○
Pseudotremia cerberus Shear, 2011 Cerberus Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Chaetaspis attenuatus Lewis & Slay, 2013 Polydesmida Troglobiont AR ○ ○ ○
Chaetaspis debilis (Causey, 1959) Polydesmida Troglobiont TN ○ ○ ○
Chaetaspis fragilis (Loomis, 1943) Polydesmida Troglobiont GNR KY ○ 2 ○
Chaetaspis mollis (Causey, 1959) Polydesmida Troglobiont G1 TN ○ ○ ○
Sphalloplana chandleri Kenk, 1977 Chandler's Cave Flatworm Tricladida Stygobiont G1G2 IN, TN, VA ○ 17 ○
Cheiridium reyesi Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Chinquipellobunus madlae (Goodnight & Goodnight, 1967) Opiliones Troglobiont G4 TX ○ 284 ○
Chinquipellobunus russelli (Goodnight & Goodnight, 1967) Opiliones Troglobiont G2G3 TX ○ 21 ○
Chitrella archeri Malcolm & J. C. Chamberlin, 1960 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Chitrella elliotti Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Chitrella major Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Chitrella superba Muchmore, 1973 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Chitrella welbourni Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 NM ○ ○ ○
Chitrellina chiricahuae Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Pseudosinella christianseni Salmon, 1965 Christiansen's Cave Springtail Entomobryomorpha Troglobiont G5 AL, GA, KY, TN, VA ○ ○ ○
Cicurina bandera Gertsch, 1992 Araneae Troglobiont G2G3 TX ○ 2 ○
Cicurina barri Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina browni Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina brunsi Cokendolpher, 2004 Araneae Troglobiont TX ○ 4 ○
Cicurina bullis Cokendolpher, 2004 Araneae Troglobiont TX ○ 14 ○
Cicurina buwata Chamberlin & Ivie, 1940 Araneae Troglobiont G1G2 TX ○ 1 107
Cicurina caliga Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 8 3
Cicurina caverna Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina coryelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 5 1
Cicurina delrio Gertsch, 1992 Araneae Troglobiont TX ○ ○ ○
Cicurina ezelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ 1
Cicurina gruta Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina holsingeri Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina hoodensis Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 35 11
Cicurina joya Gertsch, 1992 Araneae Troglobiont TX ○ ○ ○
Cicurina loftini Cokendolpher, 2004 Araneae Troglobiont TX ○ ○ ○
Cicurina machete Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina mckenziei Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina medina Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina menardia Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina mirifica Gertsch, 1992 Araneae Troglobiont TX ○ ○ ○
Cicurina mixmaster Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 3 1
Cicurina neovespera Cokendolpher, 2004 Araneae Troglobiont TX ○ 4 ○
Cicurina obscura Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina orellia Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina pablo Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina pastura Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina patei Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina platypus Cokendolpher, 2004 Araneae Troglobiont TX ○ 1 ○
Cicurina porteri Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina puentecilla Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 4 7
Cicurina rainesi Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 ○
Cicurina reclusa Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina reddelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ 1
Cicurina russelli Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina sansaba Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina selecta Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina serena Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina sheari Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina sprousei Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 ○
Cicurina stowersi Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 1 ○
Cicurina suttoni Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina travisae Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 11 208
Cicurina troglobia Cokendolpher, 2004 Araneae Troglobiont GNR TX ○ 2 ○
Cicurina ubicki Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina uvalde Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 ○
Cicurina venefica Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina vibora Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ 2 4
Cicurina watersi Gertsch, 1992 Araneae Troglobiont G1G2 TX ○ ○ ○
Cicurina wiltoni Gertsch, 1992 Araneae Troglobiont G1 AL ○ ○ ○
Cirolanides texensis Benedict, 1896 Isopoda Stygobiont G4 TX ○ 431 130
Cirolanides wassenichae Schwartz et al., 2019 Isopoda Stygobiont TX ○ 76 38
Stygobromus clantoni (Creaser, 1934) Clanton's Cave Amphipod Amphipoda Stygobiont G4 VU KS, MO ○ 35 ○
Pseudotremia nefanda Shear, 1972 Clark Cave Millipede Chordeumatida Troglobiont G3G4 IN ○ 2 ○
Pygmarrhopalites clarus (Christiansen, 1966) Clarus Cave Springtail Symphypleona Troglobiont G4 AR, MO, SD, VA, WA, WI, WV ○ 3 ○
Pseudanophthalmus caecus Krekeler, 1973 Clifton Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Caecidotea barri (Steeves, 1965) Clifton Cave Isopod Isopoda Stygobiont G1 EN KY ○ 3 ○
Calicina cloughensis (Briggs & Hom, 1967) Clough Cave Harvestman Opiliones Troglobiont G1 CA ○ 5 7
Pseudanophthalmus gracilis Valentine, 1931 Clover Hollow Cave Beetle Coleoptera Troglobiont G1G2 VA ○ 1 ○
Mexistenasellus coahuila Cole & Minckley, 1972 Coahuila Groundwater Isopod Isopoda Stygobiont G2G3 EN TX ○ 44 2
Procambarus leitheuseri Franz & Hobbs, 1983 Coastal Lowland Cave Crayfish Decapoda Stygobiont G1G2 EN FL ○ 12 5
Coecobrya kukae (Christiansen & Bellinger, 1992) Entomobryomorpha Troglobiont G1G2 HI ○ 1 ○
Coecobrya lua (Christiansen & Bellinger, 1992) Entomobryomorpha Troglobiont G1G2 HI ○ 1 ○
Coecobrya nupa (Christiansen & Bellinger, 1992) Entomobryomorpha Troglobiont G1G2 HI ○ 1 ○
Batrisodes cryptotexanus Chandler & Reddell, 2001 Coffin Cave Mold Beetle Coleoptera Troglobiont G2 TX ○ 28 ○
Texella cokendolpheri Ubick & Briggs, 1992 Cokendolpher Cave Harvestman Opiliones Troglobiont G1 ESA TX ○ 5 ○
Pseudanophthalmus colemanensis Barr, 1959 Coleman Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ 7
Coloradesmus aquiliensis (Shear, 1984) Polydesmida Troglobiont G1G2 CO ○ 7 ○
Coloradesmus beckleyi Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 2 ○
Coloradesmus hopkinsae Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 3 ○
Coloradesmus manitou Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 6 ○
Coloradesmus warneri Shear & Steinmann, 2019 Polydesmida Troglobiont CO ○ 7 ○
Stygobromus coloradensis Ward, 1977 Colorado Cave Amphipod Amphipoda Stygobiont G1G2 CO ○ 16 ○
Columbasellus acheron Lewis et al., 2003 Isopoda Stygobiont WA ○ 1 ○
Eurycea tridentifera Mitchell & Reddell, 1965 Comal Blind Salamander Caudata Stygobiont G3 VU TX ○ ○ ○
Comaldessus stygius Spangler & Barr, 1995 Comal Springs Diving Beetle Coleoptera Stygobiont G1 TX ○ 3 3
Stygoparnus comalensis Barr & Spangler, 1992 Comal Springs Dryopid Beetle Coleoptera Stygobiont G1G2 ESA TX ○ 14 ○
Comalcandona mixoni Kulkoyluoglu et al., 2019 Podocopida Stygobiont TX ○ ○ ○
Comalcandona tressleri Kulkoyluoglu & Gibson, 2018 Podocopida Stygobiont TX ○ ○ ○
Conasellus burkensis (Lewis & Lewis, 2021) Isopoda Stygobiont GNR VA ○ ○ ○
Conasellus jeffersoni (Lewis, 2009) Isopoda Stygobiont VA ○ ○ ○
Conasellus kenki (Bowman, 1967) Isopoda Stygobiont G3 DC, MD, PA, VA ○ ○ ○
Conasellus phreaticus (Lewis & Holsinger, 1985) Isopoda Stygobiont G2G3 VA ○ ○ ○
Conasellus washingtoni Lewis & Lewis, 2023 Isopoda Stygobiont DC ○ ○ ○
Pseudanophthalmus abditus Krekeler, 1973 Concealed Cave Beetle Coleoptera Troglobiont G3 KY ○ ○ ○
Condeicampa langei Ferguson, 1996 Rhabdura Troglobiont GNR TN ○ ○ ○
Conotyla vista Shear, 1971 Chordeumatida Troglobiont GH WV ○ 1 ○
Stygobromus conradi (Holsinger, 1967) Conrad's Cave Amphipod Amphipoda Stygobiont G1 VU VA ○ 15 ○
Pseudotremia cookorum Lewis, 2000 Cook's Cave Milliped Chordeumatida Troglobiont G1G2 IN ○ ○ ○
Litocampa cookei (Packard, 1871) Cooke's Cave Dipluran Rhabdura Troglobiont G5 AL, KY, TN, VA ○ 2 ○
Stygobromus cooperi (Holsinger, 1967) Cooper's Cave Amphipod Amphipoda Stygobiont G1G2 VU WV ○ 4 ○
Stygobromus cowani Wang & Holsinger, 2001 Cowan's Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Stygobromus estesi Holsinger, 1978 Craig County Cave Amphipod Amphipoda Stygobiont G4 VA ○ 9 ○
Crangonyx caecus Zhang & Holsinger, 2003 Amphipoda Stygobiont G1 KY ○ 19 ○
Crangonyx castellanum Zhang & Holsinger, 2003 Amphipoda Stygobiont G1 KY ○ 13 ○
Crangonyx specus Zhang & Holsinger, 2003 Amphipoda Stygobiont G1 KY ○ 23 ○
Crosbyella distincta Goodnight & Goodnight, 1942 Opiliones Troglobiont G1G2 AR ○ ○ ○
Crosbyella roeweri Goodnight & Goodnight, 1967 Opiliones Troglobiont G1G2 AR ○ ○ ○
Pseudanophthalmus intersectus Barr, 1965 Crossroads Cave Beetle Coleoptera Troglobiont G1G2 VA ○ ○ ○
Cryptocreagris destica Harvey & Muchmore, 2010 Pseudoscorpiones Troglobiont CO ○ 3 ○
Cryptocreagris steinmanni Harvey & Muchmore, 2010 Pseudoscorpiones Troglobiont CO ○ 3 ○
Pseudanophthalmus simulans Barr, 1985 Cub Run Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Stygobromus culveri Holsinger, 1978 Culver's Cave Amphipod Amphipoda Stygobiont G1 WV ○ 4 ○
Sphalloplana culveri Kenk, 1977 Culver's Cave Flatworm Tricladida Stygobiont G1 WV ○ 1 ○
Pseudotremia culveri Shear, 2011 Culver's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Stygobromus cumberlandus Holsinger, 1978 Cumberland Cave Amphipod Amphipoda Stygobiont G3G4 VA ○ 9 ○
Bactrurus angulus Koenemann & Holsinger, 2001 Cumberland Gap Cave Amphipod Amphipoda Stygobiont G1 TN, VA ○ 4 ○
Pseudanophthalmus hirsutus Valentine, 1931 Cumberland Gap Cave Beetle Coleoptera Troglobiont G1G2 TN, VA ○ 3 ○
Caecidotea cumberlandensis Lewis, 2000 Cumberland Gap Cave Isopod Isopoda Stygobiont G1G2 VA ○ 2 3
Orconectes barri Buhay & Crandall, 2008 Cumberland Plateau Cave Crayfish Decapoda Stygobiont G2 DD KY, TN ○ 14 66
Stygobromus curroae Wang & Holsinger, 2001 Curro's Cave Amphipod Amphipoda Stygobiont G1G2 NM ○ 3 ○
Cyclops cavernarum Ulrich, 1902 Cyclopoida Stygobiont GNR TX ○ ○ ○
Cyclops learii Ulrich, 1902 Cyclopoida Stygobiont TX ○ ○ ○
Cypria lacrima Kulkoyluoglu et al., 2017 Podocopida Stygobiont GNR TX ○ ○ ○
Darlingtonea kentuckensis Valentine, 1952 Coleoptera Troglobiont G3G4 KY, TN ○ 15 88
Pseudanophthalmus deceptivus Barr, 1981 Deceptive Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Cambarus subterraneus Hobbs III, 1993 Delaware County Cave Crayfish Decapoda Stygobiont G1 CR OK ○ 7 6
Pseudanophthalmus delicatus Valentine, 1932 Delicate Cave Beetle Coleoptera Troglobiont G3G4 VA ○ ○ ○
Dendrocoelopsis americana (Hyman, 1939) Tricladida Stygobiont G3 AR, MO, OK, TX ○ 30 ○
Dendronucleata americana Moravec & Huffman, 2000 Neoechinorhynchida Stygobiont TX ○ 2 ○
Pseudotremia inexpectata Shear, 2011 Devault's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Stygobromus hadenoecus (Holsinger, 1966) Devil's Sinkhole Amphipod Amphipoda Stygobiont G1G2 VU TX ○ 38 ○
Diacyclops alabamensis Reid, 1992 Cyclopoida Stygobiont AL ○ 2 ○
Diacyclops chrisae Reid, 1992 Cyclopoida Stygobiont MD ○ 5 ○
Diacyclops jeanneli (Chappuis, 1929) Cyclopoida Stygobiont G1G2 IN ○ 6 ○
Diacyclops nikolasarburni Suarez-Morales et al., 2013 Cyclopoida Stygobiont NM ○ 4 ○
Stygobromus dicksoni Holsinger, 1978 Dickson cave stygobromid Amphipoda Stygobiont G5 AL, GA, TN ○ 22 ○
Dinocheirus cavicola Muchmore, 1992 Pseudoscorpiones Troglobiont G2G3 TX ○ ○ ○
Phreatodrobia plana Hershler & Longley, 1986 Disc cavesnail Littorinimorpha Stygobiont G2 TX ○ 7 3
Phreatodrobia nugax (Pilsbry & Ferriss, 1906) Domed Cavesnail Littorinimorpha Stygobiont G1G2 DD TX ○ 103 13
Megacyclops donnaldsoni (Chappuis, 1929) Donaldson's Cave Copepod Cyclopoida Stygobiont G3G4 GA, IN, KY, TN ○ 1 ○
Hamohalacarus subterraneus Walter, 1931 Donaldson's Cave Water Mite Actinedida Stygobiont G1 IN ○ ○ ○
Stygobromus doughertyensis Cannizzaro & Sawicki in Cannizzaro, Balding, Stine & Sawicki, 2019 Dougherty Plain Cave Amphipod Amphipoda Stygobiont G1G2 FL, GA ○ 6 ○
Cambarus cryptodytes Hobbs, 1941 Dougherty Plain Cave Crayfish Decapoda Stygobiont G2G3 LC FL, GA ○ 47 6
Pseudotremia douglasi Lewis, 2009 Douglas' Cave Milliped Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus montanus Barr, 1965 Dry Fork Valley Cave Beetle Coleoptera Troglobiont G1 WV ○ 1 ○
Apochthonius paucispinosus Muchmore, 1967 Dry Fork Valley Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Pseudanophthalmus tullahoma Barr, 1959 Duck River Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Caecidotea dunlaporum Lewis, 2015 Dunlap's Groundwater Isopod Isopoda Stygobiont GNR IN ○ ○ 4
Pseudanophthalmus acherontis Barr, 1959 Echo Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Haideoporus texanus Young & Longley, 1976 Edwards Aquifer Diving Beetle Coleoptera Stygobiont G1G2 NE TX ○ 43 8
Eidmannella bullata Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ 1 ○
Eidmannella delicata Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ ○ ○
Eidmannella nasuta Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ ○ ○
Eidmannella reclusa Gertsch, 1984 Araneae Troglobiont G1G2 TX ○ 13 ○
Eidmannella rostrata Gertsch, 1984 Araneae Troglobiont G4 TX ○ 262 ○
Eidmannella tuckeri Cokendolpher & Reddell, 2001 Araneae Troglobiont GNR TX ○ 1 ○
Eleodes microps Aalbu et al., 2012 Coleoptera Troglobiont CA ○ 12 5
Pseudanophthalmus pusio (Horn, 1868) Ellet Valley Cave Beetle Coleoptera Troglobiont G2G3 VA ○ 5 ○
Pseudotremia cavernarum Cope, 1869 Ellett Valley Pseudotremia Millipede Chordeumatida Troglobiont G2G3 VA ○ 8 ○
Stygobromus elliotti Holsinger, 1974 Elliott's Cave Amphipod Amphipoda Stygobiont G1G2 WA ○ 11 ○
Elliotta howarthi (Elliott, 1976) Actinedida Troglobiont G1G2 WA ○ ○ ○
Fissilicreagris imperialis (Muchmore, 1969) Empire Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VU CA ○ ○ ○
Neochthonius imperialis Muchmore, 1996 Empire Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ 1 ○
Pseudanophthalmus englehardti (Barber, 1928) Englehardt's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Fontigens antroecetes Hubricht, 1972 Enigmatic ghostsnail Littorinimorpha Stygobiont G2 VU IL, MO ○ 35 1
Entomobrya troglodytes Christiansen, 1958 Entomobryomorpha Troglobiont G2G3 CO, SD ○ ○ ○
Stygobromus ephemerus (Holsinger, 1969) Ephemeral Cave Amphipod Amphipoda Stygobiont G1G2 VU VA ○ 9 ○
Eremidrilus allegheniensis (Cook, 1971) Lumbriculida Stygobiont G1G2 TN ○ 3 ○
Eremidrilus coyote Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 19 8
Eremidrilus elegans Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 20 ○
Eremidrilus felini Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 28 ○
Eremidrilus parvitheca Fend, 2023 Lumbriculida Stygobiont TX ○ 6 ○
Eremidrilus ritocsi Fend & Rodriguez, 2003 Lumbriculida Stygobiont CA ○ 10 ○
Eremidrilus samacos Fend, 2023 Lumbriculida Stygobiont TX ○ 8 ○
Ergodesmus remingtoni (Hoffman, 1962) Polydesmida Troglobiont G3 IL ○ 1 ○
Erigone stygius Gertsch, 1973 Araneae Troglobiont G1G2 HI ○ ○ ○
Pseudosinella espana Christiansen, 1961 Espana Cave Springtail Entomobryomorpha Troglobiont G3 MO ○ 1 ○
Pseudanophthalmus exoticus Krekeler, 1973 Exotic Cave Beetle Coleoptera Troglobiont G1 KY ○ 1 ○
Stygobromus flagellatus (Benedict, 1896) Ezell's Cave Amphipod Amphipoda Stygobiont G2G3 VU TX ○ 308 5
Bactrurus pseudomucronatus Koenemann & Holsinger, 2001 False Sword-tailed Cave Amphipod Amphipoda Stygobiont G2G3 AR, MO ○ 25 5
Stygobromus felleri Holsinger, 2011 Feller's Groundwater Amphipod Amphipoda Stygobiont G1 MD ○ 3 ○
Stygobromus fergusoni Holsinger, 1978 Ferguson's Cave Amphipod Amphipoda Stygobiont G1 VA ○ 5 1
Pseudotremia fergusoni Shear, 2011 Ferguson's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Caecidotea filicispeluncae Bowman & Hobbs, 1983 Fern Cave Isopod Isopoda Stygobiont G1 OH ○ 3 ○
Litocampa fieldingae (Conde, 1949) Fielding's Cave Dipluran Rhabdura Troglobiont G2G3 VA, WV ○ ○ ○
Stygobromus finleyi Holsinger, 1978 Finley's Cave Amphipod Amphipoda Stygobiont G3G4 TN, VA ○ 8 ○
Pseudotremia piscator Shear, 2011 Fisher Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Flabellorhagidia pecki Elliott, 1976 Actinedida Troglobiont G1G2 ID ○ ○ ○
Phreatodrobia micra (Pilsbry & Ferriss, 1906) Flattened Cavesnail Littorinimorpha Stygobiont G2G3 TX ○ 573 10
Crangonyx parhobbsi Cannizzaro & Sawicki in Cannizzaro, Balding, Lazo-Wasem & Sawicki, 2020 Florida Big Bend Cave Amphipod Amphipoda Stygobiont G1G2 FL, GA ○ 13 89
Crangonyx grandimanus Bousfield, 1963 Florida Cave Amphipod Amphipoda Stygobiont G2G3 VU FL ○ 67 13
Caecidotea hobbsi (Maloney, 1939) Florida Cave Isopod Isopoda Stygobiont G1G2 FL, GA ○ 17 8
Mexistenasellus floridensis Lewis & Sawicki, 2016 Florida Cave Isopod Isopoda Stygobiont G1 FL ○ 3 ○
Stygobromus floridanus Holsinger & Sawicki, 2016 Florida Panhandle Cave Amphipod Amphipoda Stygobiont G1G2 FL ○ 12 43
Forcipomyia pholetor Wirth & Howarth, 1982 Diptera Troglobiont HI ○ ○ ○
Pseudosinella fonsa Christiansen & Bellinger, 1996 Fountain Cave Springtail Entomobryomorpha Troglobiont G3G4 IN, OH, TN ○ 1 ○
Typhlogastrura fousheensis Christiansen & Wang, 2006 Foushee Cave Springtail Poduromorpha Troglobiont G1 AR ○ 1 ○
Amnicola cora Hubricht, 1979 Foushee Cavesnail Littorinimorpha Stygobiont G1 VU AR ○ 9 ○
Foveacheles auricularia Zacharda & Elliott, 1985 Actinedida Troglobiont CA ○ ○ ○
Foveacheles goffi Zacharda, 1982 Actinedida Troglobiont G1G2 HI ○ 1 ○
Foveacheles tenorioae Zacharda, 1982 Actinedida Troglobiont G1G2 HI ○ ○ ○
Foveacheles titanica Zacharda & Elliott, 1985 Actinedida Troglobiont G2G3 CA ○ ○ ○
Pseudanophthalmus fowlerae Barr, 1980 Fowler's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Stygobromus franzi Holsinger, 1978 Franz's Cave Amphipod Amphipoda Stygobiont G3G4 MD, PA, WV ○ 46 ○
Caecidotea franzi (Holsinger & Steeves, 1971) Franz's Cave Isopod Isopoda Stygobiont G2G4 KY, MD, PA, WV ○ 15 ○
Stygobromus amicus Holsinger, 2014 Friendly Cave Amphipod Amphipoda Stygobiont G1 MD ○ 4 ○
Caecidotea rotunda Bowm& & Lewis, 1984 Frost Cave Isopod Isopoda Stygobiont G2G4 IN, OH ○ 17 32
Caecidotea fustis Lewis, 1981 Fustis Cave Isopod Isopoda Stygobiont G3 MO ○ 15 ○
Banksula galilei Briggs, 1974 Galile's Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Stygobromus gallawayae Wang & Holsinger, 2001 Gallaway's Cave Amphipod Amphipoda Stygobiont G1 CA ○ 3 ○
Galumna alata (Hermann, 1804) Oribatida Troglobiont G1G2 KY ○ ○ ○
Gammarus cohabitus Holsinger & Shafer, 2008 Amphipoda Stygobiont G1G2 PA ○ 10 ○
Gammarus minus tenuipes Shoemaker, 1940 Amphipoda Stygobiont VA, WV ○ ○ ○
Pseudosinella certa Christiansen & Bellinger, 1980 Gandy Creek Cave Springtail Entomobryomorpha Troglobiont G1 WV ○ 1 ○
Pseudanophthalmus hortulanus Barr, 1965 Garden Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudotremia garlandae Lewis, 2005 Garland's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus horni (Garman, 1892) Garman's Cave Beetle Coleoptera Troglobiont G3 KY ○ 8 ○
Geocentrophora cavernicola Carpenter, 1970 Alloecoela Stygobiont G2 KY, VA, WV ○ 2 ○
Eurycea wallacei (Carr, 1939) Georgia Blind Salamander Caudata Stygobiont G1G2 VU FL, GA ○ 175 22
Pseudanophthalmus georgiae Barr, 1981 Georgia Cave Beetle Coleoptera Troglobiont G1G2 GA ○ ○ ○
Sphalloplana georgiana Hyman, 1954 Georgia Cave Flatworm Tricladida Stygobiont G1 GA ○ 2 ○
Nesticus georgia Gertsch, 1984 Georgia Cave Spider Araneae Troglobiont G1G2 GA ○ 2 4
Pseudotremia lusciosa (Loomis, 1939) Germany Valley Cave Millipede Chordeumatida Troglobiont G1 WV ○ 2 ○
Kleptochthonius gertschi Malcolm & Chamberlin, 1961 Gertsch's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Orconectes inermis Cope, 1872 Ghost Crayfish Decapoda Stygobiont G5 LC IN, KY ○ 114 3
Comalcandona gibsoni Kulkoyluoglu, 2019 Gibson's Groundwater Ostracod Podocopida Stygobiont TX ○ ○ ○
Glacicavicola bathyscioides Westcott, 1968 Coleoptera Troglobiont G3 VU ID, WY ○ 4 ○
Stygobromus glacialis Wang & Holsinger, 2001 Glacier National Park Cave Amphipod Amphipoda Stygobiont G1 MT ○ 7 ○
Cicurina vespera Gertsch, 1992 Government Canyon Bat Cave Meshweaver Araneae Troglobiont G1 ESA TX ○ 6 ○
Tayshaneta microps (Gertsch, 1974) Government Canyon Bat Cave Spider Araneae Troglobiont G1 ESA TX ○ 24 ○
Stygobromus gradyi Holsinger, 1974 Grady's Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 9 ○
Stygobromus grahami Holsinger, 1974 Graham's Cave Amphipod Amphipoda Stygobiont G2 CA ○ 15 ○
Larca cavicola (Muchmore, 1981) Grand Canyon Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Zygonopus weyeriensis Causey, 1960 Grand Caverns Blind Cave Millipede Chordeumatida Troglobiont G3G4 VA, WV ○ ○ ○
Grandidierella palama Barnard, 1977 Amphipoda Stygobiont HI ○ 4 ○
Pseudanophthalmus jonesi Valentine, 1945 Grassy Cove Cave Beetle Coleoptera Troglobiont G1G2 TN ○ 3 ○
Nesticus dilutus Gertsch, 1984 Grassy Creek Cave Spider Araneae Troglobiont G1 TN ○ 2 4
Kleptochthonius griseomanus Muchmore, 2000 Gray-handed Cave Pseudoscorpion Pseudoscorpiones Troglobiont G2 IN ○ ○ ○
Microcreagris grandis Muchmore, 1962 Great Basin Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 NV ○ 12 2
Pseudanophthalmus pholeter Krekeler, 1973 Greater Adams Cave Beetle Coleoptera Troglobiont G1 KY ○ 2 ○
Stygobromus emarginatus Hubricht, 1943 Greenbrier Cave Amphipod Amphipoda Stygobiont G3 VU MD, WV ○ 82 2
Cambarus nerterius Hobbs, 1964 Greenbrier Cave Crayfish Decapoda Stygobiont G1 NT WV ○ 26 2
Fontigens turritella Hubricht, 1976 Greenbrier Cavesnail Littorinimorpha Stygobiont G1 NT WV ○ 8 ○
Pseudotremia fulgida Loomis, 1943 Greenbrier Valley Cave Millipede Chordeumatida Troglobiont G3 WV ○ 2 ○
Kleptochthonius henroti (Vachon, 1952) Greenbrier Valley Cave Pseudoscorpion Pseudoscorpiones Troglobiont G2 WV ○ 1 ○
Stygobromus sparsus Holsinger, 1978 Gregory Cave Amphipod Amphipoda Stygobiont G1 TN ○ 8 ○
Cottus specus Adams et al. 2013 Grotto sculpin Scorpaeniformes Stygobiont G2 ESA MO ○ 10 21
Aphrastochthonius grubbsi Muchmore, 1984 Grubb's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Banksula grubbsi Briggs & Ubick, 1981 Grubbs' Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Larca guadalupensis (Muchmore, 1981) Guadalupe Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Hesperochernes occidentalis (Hoff & Bolsterli, 1956) Guano Pseudoscorpion Pseudoscorpiones Troglobiont G5 AR, MO, OK, TX ○ 9 ○
Halocaridina palahemo Kensley & Williams, 1986 Decapoda Stygobiont G1 HI ○ 3 ○
Halocaridina rubra Holthuis, 1963 Decapoda Stygobiont G3 HI ○ 25 337
Haplotaxis brinkhursti Cook, 1975 Haplotaxida Stygobiont G1 WV ○ ○ ○
Stygobromus harai Holsinger, 1974 Hara's Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 6 ○
Pseudotremia jaculohamatum Shear, 2011 Harpoon Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Ptomaphagus hatchi Jeannel, 1933 Hatch's Cave Fungus Beetle Coleoptera Troglobiont G3 AL, TN ○ 4 ○
Hawaiioscia micropthalma Taiti & Howarth, 1997 Isopoda Troglobiont G1G2 HI ○ ○ ○
Hawaiioscia paeninsulae Taiti & Howarth, 1997 Isopoda Troglobiont G1G2 HI ○ ○ 1
Hawaiioscia parvituberculata Schultz, 1973 Isopoda Troglobiont G1G2 HI ○ 7 ○
Hawaiioscia rotundata Taiti & Howarth, 1997 Isopoda Troglobiont G1G2 HI ○ 1 3
Hawinella lava Bellinger & Christiansen, 1974 Entomobryomorpha Troglobiont G2G3 HI ○ 1 ○
Stygobromus hayi (Hubricht & Mackin, 1940) Hay's Spring Amphipod Amphipoda Stygobiont G1G2 EN ESA DC ○ 11 11
Cambarus zophonastes Hobbs & Bedinger, 1964 Hell Creek Cave Crayfish Decapoda Stygobiont G1 CR ESA AR ○ 6 2
Typhlogastrura helleri Christiansen & Wang, 2006 Heller Cave Springtail Poduromorpha Troglobiont GNR PA ○ 1 ○
Batrisodes venyivi Chandler, 1992 Helotes Cave Mold Beetle Coleoptera Troglobiont G1 ESA TX ○ 4 ○
Pseudobaicalasellus henroti (Bresson, 1955) Henrot's Cave Isopod Isopoda Stygobiont G2 VA ○ ○ ○
Stygobromus herbsti Wang & Holsinger, 2001 Herbst's Cave Amphipod Amphipoda Stygobiont G2G3 NV ○ 11 ○
Hesperochernes bradybaughi Harvey & Wynne, 2014 Pseudoscorpiones Troglobiont GNR AZ ○ ○ ○
Hesperonemastoma inops (Packard, 1884) Opiliones Troglobiont G1G2 KY ○ ○ ○
Hesperonemastoma pallidimaculosum (Goodnight & Goodnight, 1945) Opiliones Troglobiont G3 AL ○ ○ ○
Hesperonemastoma smilax Shear, 2010 Opiliones Troglobiont GNR WV ○ ○ ○
Hexabathynella hessleri Cho, 2001 Bathynellacea Stygobiont CA ○ ○ ○
Hexabathynella muliebris Cho, 2001 Bathynellacea Stygobiont CA ○ ○ ○
Hexabathynella otayana Cho, 2001 Bathynellacea Stygobiont CA ○ 2 ○
Hexabathynella virginiae Cho & Schminke, 2006 Bathynellacea Stygobiont VA ○ ○ ○
Pseudanophthalmus conditus Krekeler, 1973 Hidden Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 2 ○
Fontigens cryptica Hubricht, 1963 Hidden springsnail Littorinimorpha Stygobiont G1 IN ○ 7 1
Phreatodrobia punctata Hershler & Longley, 1986 High-hat cavesnail Littorinimorpha Stygobiont G2 TX ○ 12 3
Pseudosinella hirsuta (Delamare Deboutteville, 1949) Hirsute Cave Springtail Entomobryomorpha Troglobiont G5 AL, GA, KY, TN, VA ○ 8 ○
Crangonyx hobbsi Shoemaker, 1941 Hobb's Cave Amphipod Amphipoda Stygobiont G2G3 VU FL ○ 80 146
Hobbsinella edwardensis Camacho et al., 2017 Bathynellacea Stygobiont GNR TX ○ 5 1
Hobbsinella gunnisonensis Camacho & Taylor, 2023 Bathynellacea Stygobiont CO ○ ○ ○
Oncopodura hoffi Christiansen & Bonet, 1980 Hoff's Cave Springtail Entomobryomorpha Troglobiont G1G2 MO ○ 3 ○
Stygobromus hoffmani Holsinger, 1978 Hoffman's Cave Amphipod Amphipoda Stygobiont G2 VA ○ 4 1
Pseudanophthalmus hoffmani Barr, 1965 Hoffman's Cave Beetle Coleoptera Troglobiont G2G3 VA ○ 1 ○
Pseudotremia hoffmani Lewis, 2009 Hoffman's Cave Millipede Chordeumatida Troglobiont GNR TN ○ ○ ○
Sinella hoffmani Wray, 1952 Hoffman's Cave Springtail Entomobryomorpha Troglobiont G5 MD, PA, TN, VA, WV ○ 5 ○
Sphalloplana hoffmasteri (Hyman, 1954) Hoffmaster's Cave Flatworm Tricladida Stygobiont G3G4 MD, WV ○ 12 ○
Phagocata holleri Kenk, 1979 Holler's Cave Flatworm Tricladida Stygobiont G1G2 NC ○ 4 ○
Pseudotremia hollidayi Lewis, 2005 Holliday's Cave Milliped Chordeumatida Troglobiont G1 TN ○ ○ ○
Helicodiscus notius specus Hubricht, 1962 Hollow Coil Stylommatophora Troglobiont G2 KY, TN ○ ○ ○
Glyphyalinia specus Hubricht, 1965 Hollow glyph Stylommatophora Troglobiont G4 AL, GA, KY, TN, WV ○ 28 ○
Stygobromus holsingeri Ward, 1977 Holsinger's Cave Amphipod Amphipoda Stygobiont G1G2 CO ○ 33 ○
Pseudanophthalmus holsingeri Barr, 1965 Holsinger's Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Sphalloplana holsingeri Kenk, 1977 Holsinger's Cave Flatworm Tricladida Stygobiont G1G2 NE VA ○ 10 ○
Pseudobaicalasellus holsingeri (Steeves, 1963) Holsinger's Cave Isopod Isopoda Stygobiont G5 MD, VA, WV ○ ○ ○
Pseudotremia johnholsingeri Shear, 2011 Holsinger's Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Hesperochernes holsingeri Muchmore, 1994 Holsinger's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 IN ○ ○ ○
Nesticus holsingeri Gertsch, 1984 Holsinger's Cave Spider Araneae Troglobiont G3G4 VA ○ 4 16
Holsingerius samacos (Holsinger, 1980) Amphipoda Stygobiont G1G2 TX ○ 78 ○
Amblyopsis hoosieri Niemiller, Prejean & Chakrabarty, 2014 Hoosier Cavefish Percopsiformes Stygobiont G2 IN ○ 63 1
Horologion hubbardi Harden & Davidson, 2024 Coleoptera Troglobiont GNR VA ○ ○ 10
Horreolanus orphanus Mitchell, 1955 Trombidiformes Stygobiont IL ○ 42 5
Caconemobius howarthi Gurney & Rentz, 1978 Howarth's Cave Cricket Orthoptera Troglobiont G1G2 VU HI ○ ○ ○
Pseudanophthalmus hubbardi Barber, 1928 Hubbard's Cave Beetle Coleoptera Troglobiont G1 VA ○ 5 ○
Pseudotremia hubbardi Shear, 2011 Hubbard's Cave Milliped Chordeumatida Troglobiont G1G2 VA ○ ○ ○
Pseudanophthalmus hubrichti Valentine, 1948 Hubricht's Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Sphalloplana hubrichti (Hyman, 1945) Hubricht's Cave Flatworm Tricladida Stygobiont G2G4 IL, MO ○ 21 ○
Allocrangonyx hubrichti Holsinger, 1971 Hubricht's Long-tailed Cave Amphipod Amphipoda Stygobiont G3G4 VU AR, MO ○ 18 ○
Phreatodrobia conica Hershler & Longley, 1986 Hueco Cavesnail Littorinimorpha Stygobiont G1 TX ○ 27 7
Stygobromus hubbardi Holsinger, 2009 Hupp's Hill Cave Amphipod Amphipoda Stygobiont G1 VA ○ 3 ○
Pseudanophthalmus parvicollis Jeannel, 1931 Hupps Hill Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Hyalella cretae Cannizzaro et al., 2023 Amphipoda Stygobiont GNR NV ○ ○ ○
Stygobromus hyporheicus Wang & Holsinger, 2001 Hyporheic Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Pseudanophthalmus frigidus Barr, 1981 Icebox Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Idagona jasperi Shear, 2007 Chordeumatida Troglobiont UT ○ ○ ○
Idagona lehmanensis Shear, 2007 Chordeumatida Troglobiont NV ○ 34 ○
Stygobromus idahoensis Wang & Holsinger, 2001 Idaho Groundwater Amphipod Amphipoda Stygobiont G1 ID ○ 2 ○
Idagona westcotti Buckett & Gardner, 1967 Idaho Lave Tube Millipede Chordeumatida Troglobiont G1G2 ID ○ ○ ○
Illacme tobini Marek et al., 2016 Siphonophorida Troglobiont CA ○ ○ ○
Gammarus acherondytes Hubricht & Mackin, 1940 Illinois Cave Amphipod Amphipoda Stygobiont G2G3 EN ESA IL ○ 37 80
Pseudanophthalmus illinoisensis Barr & Peck, 1966 Illinois Cave Beetle Coleoptera Troglobiont G1 IL ○ 1 ○
Stygobromus imperialis Wang & Holsinger, 2001 Imperial Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Pseudobaicalasellus incurvus (Steeves & Holsinger, 1968) Incurved Cave Isopod Isopoda Stygobiont G2G4 TN, VA ○ ○ ○
Pseudanophthalmus tiresias Barr, 1959 Indian Grave Point Cave Beetle Coleoptera Troglobiont G1 TN ○ 1 ○
Crangonyx indianensis Zhang & Holsinger, 2003 Indiana Cave Amphipod Amphipoda Stygobiont G3 IN ○ 33 ○
Apochthonius indianensis Muchmore, 1967 Indiana Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 IN ○ ○ ○
Diacyclops indianensis Reid, 2004 Indiana Groundwater Copepod Cyclopoida Stygobiont G2 IN, TN ○ ○ ○
Rheocyclops indiana Reid in Reid et al. 1999 Indiana Groundwater Copepod Cyclopoida Stygobiont G1 IN, TN ○ 2 ○
Batrisodes texanus Chandler, 1992 Inner Space Cavern Mold Beetle Coleoptera Troglobiont G1G2 ESA TX ○ 15 ○
Pseudanophthalmus inquisitor Barr, 1980 Inquirer Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus intermedius (Valentine, 1931) Intermediate Cave Beetle Coleoptera Troglobiont G2 TN ○ 2 ○
Stygobromus interstitialis Wang & Holsinger, 2001 Interstitial Amphipod Amphipoda Stygobiont G1G3 CO ○ 3 ○
Iolania frankanstonei Hoch, 2024 Homoptera Troglobiont HI ○ ○ ○
Stygobromus iowae Hubricht, 1943 Iowa Cave Amphipod Amphipoda Stygobiont G2G3 IA, IL ○ 8 ○
Islandiana muma Ivie, 1965 Araneae Troglobiont G1G2 AL, VA ○ 1 ○
Islandiana unicornis Ivie, 1965 Araneae Troglobiont G2G3 OK, TX ○ 4 ○
Speodesmus ivyi Elliott, 2004 Ivy's Cave Millipede Polydesmida Troglobiont GNR TX ○ 18 ○
Crangonyx manubrium Cannizzaro & Sawicki in Cannizzaro, Balding, Lazo-Wasem & Sawicki, 2019 Jackson County Cave Amphipod Amphipoda Stygobiont G1G2 FL ○ 32 123
Stygobromus abditus Holsinger, 1978 James Cave Amphipod Amphipoda Stygobiont G2G3 VA ○ 16 ○
Pseudocandona jeanneli (Klie, 1931) Jeannel's Cave Ostracod Podocopida Stygobiont G3 IN, KY, TN ○ 3 ○
Stygobromus jemezensis Wang & Holsinger, 2001 Jemez Mountains Groundwater Amphipod Amphipoda Stygobiont G1G2 NM ○ 2 ○
Eurycea tonkawae Chippindale et al., 2000 Jollyville Plateau Salamander Caudata Stygobiont G2 EN ESA TX ○ 232 22
Tetracion jonesi Hoffman, 1956 Jones' Cave Millipede Callipodida Troglobiont G3G4 AL, TN ○ 7 17
Tyrannochthonius jonesi Chamberlin, 1995 Jones' Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Pseudotremia salifodina Shear, 2011 Jones' Saltpetre Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Caecidotea jordani (Eberly, 1966) Jordan's Cave Isopod Isopoda Stygobiont G2G3 IN ○ 20 10
Sphalloplana kansensis Hyman, 1945 Kansas Cave Flatworm Tricladida Stygobiont G1G2 KS ○ ○ ○
Bactrurus hubrichti Shoemaker, 1945 Kansas Well Amphipod Amphipoda Stygobiont G4 KS, MO, OK ○ 26 ○
Spelaeorchestia koloana Bousfield & Howarth, 1976 Kauai Cave Amphipod Amphipoda Troglobiont G1G2 EN ESA HI ○ 4 ○
Adelocosa anops Gertsch, 1973 Kauai Cave Wolf Spider Araneae Troglobiont G1G2 EN ESA HI ○ 1 ○
Caecidotea kendeighi (Steeves & Seidenberg, 1971) Kendeigh's Groundwater Isopod Isopoda Stygobiont G5 IA, IL, IN, MO ○ 81 9
Ptomaphagus hirtus (Tellkampf, 1844) Kentucky Cave Beetle Coleoptera Troglobiont G4 KY ○ 14 ○
Caecidotea insula Lewis, 2013 Kindt's Cave Isopod Isopoda Stygobiont G1 OH ○ 2 ○
Banksula tutankhamen Ubick & Briggs, 2002 King Tut Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Kleptochthonius affinis Muchmore, 1976 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius anophthalmus Muchmore, 1970 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius attenuatus Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 KY ○ ○ ○
Kleptochthonius barri Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius binoculatus Muchmore, 1974 Pseudoscorpiones Troglobiont G1G2 VA ○ ○ ○
Kleptochthonius cerberus Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 KY ○ 1 ○
Kleptochthonius charon Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius daemonius Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Kleptochthonius erebicus Muchmore, 1965 Pseudoscorpiones Troglobiont G1 KY ○ ○ ○
Kleptochthonius hageni Muchmore, 1963 Pseudoscorpiones Troglobiont G1G2 KY ○ 7 ○
Kleptochthonius hubrichti Muchmore, 1965 Pseudoscorpiones Troglobiont G1G2 KY ○ ○ ○
Kleptochthonius infernalis Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius krekeleri Muchmore, 1965 Pseudoscorpiones Troglobiont G2G4 KY ○ ○ ○
Kleptochthonius magnus Muchmore, 1966 Pseudoscorpiones Troglobiont G1 GA, TN ○ ○ ○
Kleptochthonius microphthalmus Malcolm & Chamberlin, 1961 Pseudoscorpiones Troglobiont G1G2 KY ○ ○ ○
Kleptochthonius myopius Malcolm & J. C. Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius pluto Muchmore, 1965 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius proximosetus Muchmore, 1976 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius regulus Muchmore, 1970 Pseudoscorpiones Troglobiont G1G2 VA ○ ○ ○
Kleptochthonius rex Malcolm & J. C. Chamberlin, 1961 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius similis Muchmore, 1976 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Kleptochthonius stygius Muchmore, 1965 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Kleptochthonius tantalus Muchmore, 1966 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus krameri Krekeler, 1973 Kramer's Cave Beetle Coleoptera Troglobiont G1 OH ○ 2 ○
Texamaurops reddelli Barr & Steeves, 1963 Kretschmarr Cave Mold Beetle Coleoptera Troglobiont G1G2 ESA TX ○ 17 ○
Caconemobius varius Gurney & Rentz, 1978 Kuamana Cave Cricket Orthoptera Troglobiont G1G2 VU HI ○ 1 ○
Larca laceyi Muchmore, 1981 Lacey's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Cambarus laconensis Buhay & Crandall, 2009 Lacon Exit Cave Crayfish Decapoda Stygobiont G1 CR AL ● 8 12
Lacrimacandona wisei Kulkoyluoglu et al., 2017 Podocopida Stygobiont GNR TX ○ ○ ○
Laelaps cavernicola Packard, 1888 Mesostigmata Troglobiont KY ○ ○ ○
Stygobromus lacicolus Holsinger, 1974 Lake Tahoe Groundwater Amphipod Amphipoda Stygobiont G1 CA, NV ○ 5 ○
Dendrocoelopsis hymanae Kawakatsu, 1968 Lake Tahoe planarian Tricladida Stygobiont GNR CA, NV ○ 3 ○
Stygobromus lanensis Wang & Holsinger, 2001 Lane County Groundwater Amphipod Amphipoda Stygobiont G1G2 OR ○ 2 ○
Larca welbourni (Muchmore, 1981) Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Pseudanophthalmus cnephosus Krekeler, 1973 Larue County Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudotremia laudermilki Lewis, 2009 Laudermilk's Cave Millipede Chordeumatida Troglobiont GNR KY ○ ○ ○
Stygobromus pollostus Holsinger, 1978 Least Cave Amphipod Amphipoda Stygobiont G2G3 WV ○ 8 ○
Stygobromus leensis Holsinger, 1978 Lee County Cave Amphipod Amphipoda Stygobiont G1G2 VA ○ 5 1
Lirceus usdagalun Holsinger & Bowman, 1973 Lee County Cave Isopod Isopoda Stygobiont G1 EN ESA VA ○ 14 11
Pseudanophthalmus catoryctos Krekeler, 1973 Lesser Adams Cave Beetle Coleoptera Troglobiont G1 KY ○ 3 ○
Leucohya texana Muchmore, 1986 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Kleptochthonius lewisorum Muchmore, 2000 Lewis Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 IN ○ ○ ○
Crangonyx lewisi Zhang & Holsinger, 2003 Lewis' Cave Amphipod Amphipoda Stygobiont G2 IN, KY ○ 15 ○
Kenkia lewisi Kenk, 1975 Lewis' cave planarian Tricladida Stygobiont G1 IL, MO ○ 15 ○
Pygmarrhopalites lewisi (Christiansen & Bellinger, 1998) Lewis' Cave Springtail Symphypleona Troglobiont GNR IL, IN ○ 3 ○
Diacyclops lewisi Reid, 2004 Lewis' Groundwater Copepod Cyclopoida Stygobiont G1 IN ○ ○ ○
Liagoceradocus lonomaka (Barnard, 1977) Amphipoda Stygobiont G1G2 HI ○ ○ ○
Procambarus lucifugus (Hobbs, 1940) Light-fleeing Cave Crayfish Decapoda Stygobiont G1G2 LC FL ○ 89 10
Ligidium kofoidi Maloney, 1930 Isopoda Troglobiont G1G2 CA ○ 5 ○
Pseudanophthalmus calcareus Barr, 1981 Limestone Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Linopodes mammouthia Banks, 1897 Trombidiformes Troglobiont KS ○ ○ ○
Liocranoides unicolor Keyserling, 1881 Araneae Troglobiont GU KY, TN ○ 5 ○
Lirceolus bisetus (Steeves, 1968) Isopoda Stygobiont G1G2 TX ○ 14 5
Lirceolus cocytus Lewis, 2001 Isopoda Stygobiont GNR TX ○ 47 10
Lirceolus hardeni Lewis & Bowman, 1996 Isopoda Stygobiont G2G3 TX ○ 227 21
Lirceolus nidulus Lewis, 2001 Isopoda Stygobiont GNR TX ○ 5 ○
Lirceolus pilus (Steeves, 1968) Isopoda Stygobiont G2G3 TX ○ 117 8
Lirceus bisetus Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ 3
Lirceus holsingeri Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ ○
Lirceus littonensis Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ 3
Lirceus orndorffi Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ 9
Lirceus zigleri Lewis & Lewis, 2023 Isopoda Stygobiont GNR VA ○ ○ 3
Lissocreagris eurydice (Muchmore, 1969) Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Lissocreagris nickajackensis (Muchmore, 1966) Pseudoscorpiones Troglobiont G1G2 AL, TN ○ ○ ○
Lissocreagris persephone (Chamberlin, 1962) Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Lissocreagris pluto (Chamberlin, 1962) Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Litocampa henroti (Conde, 1949) Rhabdura Troglobiont G1G2 AL ○ ○ ○
Litocampa hoffmani Ferguson, 2009 Rhabdura Troglobiont G3 VA ○ ○ ○
Litocampa jonesi (Conde, 1949) Rhabdura Troglobiont G2G3 KY, TN ○ ○ ○
Litocampa valentinei (Conde, 1949) Rhabdura Troglobiont G5 AL, TN ○ ○ ○
Pseudanophthalmus cordicollis Barr, 1981 Little Kennedy Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Pseudanophthalmus longiceps Barr, 1981 Long-headed Cave Beetle Coleoptera Troglobiont G1G2 TN, VA ○ ○ ○
Stygobromus longipes (Holsinger, 1966) Long-legged Cave Amphipod Amphipoda Stygobiont G2G3 VU TX ○ 14 7
Pseudotremia loomisi Shear, 2011 Loomis's Rough-backed Millipede Chordeumatida Troglobiont G1 VA ○ ○ ○
Lophomus skamania (Causey, 1972) Chordeumatida Troglobiont WA ○ 1 ○
Pseudanophthalmus troglodytes Krekeler, 1973 Louisville Cave Beetle Coleoptera Troglobiont G1 KY ○ 2 ○
Stygobromus pseudospinosus Holsinger, 1978 Luray Caverns Amphipod Amphipoda Stygobiont G1 VA ○ 7 ○
Zygonopus whitei Causey, 1960 Luray Caverns Blind Cave Millipede Chordeumatida Troglobiont G3G4 VA, WV ○ ○ ○
Kleptochthonius lutzi Malcolm & Chamberlin, 1961 Lutz's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Lycosa howarthi Gertsch, 1973 Araneae Troglobiont G1G2 HI ○ 2 ○
Lymantes nadineae Paquin & Anderson, 2009 Coleoptera Troglobiont GNR TX ○ 5 ○
Stygobromus mackenziei Holsinger, 1974 MacKenzie's Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 4 ○
Pseudanophthalmus macradyi Valentine, 1948 Macrady's Cave Beetle Coleoptera Troglobiont G1G2 TN ○ 1 ○
Macrocheles stygius (Packard, 1888) Mesostigmata Troglobiont G1G2 KY ○ ○ ○
Macrocheles troglodytes (Packard, 1888) Mesostigmata Troglobiont G1G2 KY ○ ○ ○
Macromastus umpqua Shear, 1982 Chordeumatida Troglobiont G1G2 OR ○ 1 ○
Pseudanophthalmus limicola Jeannel, 1931 Maddens Cave Beetle Coleoptera Troglobiont G1G2 VA ○ 1 ○
Stygobromus stegerorum Holsinger, 1978 Madison Cave Amphipod Amphipoda Stygobiont G1 VA ○ 5 6
Cicurina madla Gertsch, 1992 Madla Cave Meshweaver Araneae Troglobiont G1 ESA TX ○ 23 22
Pygmarrhopalites madonnensis (Zeppelini & Christiansen, 2003) Madonna Cave Springtail Symphypleona Troglobiont GNR IL ○ 3 ○
Pseudanophthalmus virginicus (Barr, 1960) Maiden Spring Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Stygobromus hubbsi Shoemaker, 1942 Malheur Cave Amphipod Amphipoda Stygobiont G1G2 VU OR ○ 6 ○
Kenkia rhynchida Hyman, 1937 Malheur Cave Flatworm Tricladida Stygobiont G1G2 NE OR ○ 2 ○
Apochthonius malheuri Benedict & Malcolm, 1973 Malheur Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 OR ○ ○ ○
Orconectes pellucidus (Tellkampf, 1844) Mammoth Cave Crayfish Decapoda Stygobiont G4 LC KY, TN ○ 139 2
Palaemonias ganteri Hay, 1901 Mammoth Cave Shrimp Decapoda Stygobiont G1 VU ESA KY ○ 11 ○
Anthrobia monmouthia Tellkampf, 1844 Mammoth Cave Spider Araneae Troglobiont G5 AL, KY, TN, VA ○ 3 ○
Stygobromus vitreus Cope, 1872 Mammoth Cave stygobromid Amphipoda Stygobiont G4 AL, KY, TN ○ 40 ○
Antrorbis breweri Hershler & Thompson, 1990 Manitou cavesnail Littorinimorpha Stygobiont G1 VU AL ○ 7 1
Pseudotremia manni Lewis, 2005 Mann's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus stricticollis Jeannel, 1931 Marengo Cave Beetle Coleoptera Troglobiont G4 IN ○ 5 ○
Pseudocandona marengoensis (Klie, 1931) Marengo Cave Ostracod Podocopida Stygobiont G1G2 IN ○ ○ ○
Banksula martinorum Briggs & Ubick, 1981 Martins' Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Mecyclothorax aa Liebherr, 2008 Coleoptera Troglobiont HI ○ ○ ○
Banksula melones Briggs, 1974 Melones Cave Harvestman Opiliones Troglobiont G1 VU CA ○ 1 ○
Pseudanophthalmus sidus Barr, 1965 Meredith Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Prietella phreatophila Carranza, 1954 Mexican Blindcat Siluriformes Stygobiont GNR VU ESA TX ○ 142 8
Mexichthonius exoticus Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 TN ○ ○ ○
Mexiweckelia hardeni Holsinger, 1992 Amphipoda Stygobiont G2G3 TX ○ 21 ○
Procambarus milleri Hobbs, 1971 Miami Cave Crayfish Decapoda Stygobiont G1 EN ESA? FL ○ 14 6
Phreatodrobia imitata Hershler & Longley, 1986 Mimic cavesnail Littorinimorpha Stygobiont G1 VU TX ○ 42 2
Stygobromus parvus (Holsinger, 1969) Minute Cave Amphipod Amphipoda Stygobiont G2 VU WV ○ 10 ○
Lethemurus missus Mills, 1948 Missus Cave Springtail Entomobryomorpha Troglobiont G4 CO, IL, IN, KY, MO, TN ○ 5 ○
Mixojapyx reddelli Muegge, 1992 Dicellurata Troglobiont G3G4 TX ○ 9 ○
Montanabathynella pecosensis Camacho & Hutchins, 2021 Bathynellacea Stygobiont TX ○ ○ ○
Montanabathynella salish Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Nesticus pecki Hedin & Dellinger, 2005 Monteagle Cave Spider Araneae Troglobiont GNR TN ○ 3 4
Stygobromus morrisoni (Holsinger, 1967) Morrison's Cave Amphipod Amphipoda Stygobiont G1G2 VU VA, WV ○ 12 1
Bryocamptus morrisoni (Chappuis, 1929) Morrison's Cave Copepod Harpacticoida Stygobiont G3G4 IN ○ 2 ○
Stygobromus montanus (Holsinger, 1967) Mountain Cave Amphipod Amphipoda Stygobiont G1 VU AR ○ 6 ○
Mundochthonius cavernicola Muchmore, 1968 Pseudoscorpiones Troglobiont G1G2 IL, MO ○ ○ ○
Mundochthonius holsingeri Benedict & Malcolm, 1974 Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Pseudogarypus orpheus Muchmore, 1981 Music Hall Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 CA ○ ○ ○
Stygobromus myersae Wang & Holsinger, 2001 Myers' Groundwater Amphipod Amphipoda Stygobiont G1 CA, NV ○ 6 ○
Apochthonius mysterius Muchmore, 1976 Mystery Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 MO ○ ○ ○
Namiotkocypria haysensis Kulkoyluoglu, 2018 Podocopida Stygobiont TX ○ ○ ○
Nampabius turbator Crabill, 1952 Lithobiomorpha Troglobiont G1 VA ○ ○ ○
Nannopetalum fontis Shear, 2003 Chordeumatida Troglobiont AL ○ ○ ○
Nannopetalum vespertilo Shear, 2003 Chordeumatida Troglobiont NC ○ ○ ○
Stygobromus rallus Wang & Holsinger, 2001 Narrow Groundwater Amphipod Amphipoda Stygobiont G1G3 WA ○ 3 ○
Pseudanophthalmus pontis Barr, 1965 Natural Bridge Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudobaicalasellus bowmani (Lewis, 1980) Natural Bridge Cave Isopod Isopoda Stygobiont G1 VA ○ ○ ○
Ereboporus naturaconservatus Miller et al., 2019 Nature Conservancy Diving Beetle Coleoptera Stygobiont GNR TX ○ 1 8
Neanura hawaiiensis (Bellinger & Christiansen, 1974) Poduromorpha Troglobiont G1G2 HI ○ ○ ○
Neaphaenops tellkampfi (Erichson, 1844) Coleoptera Troglobiont G3G4 KY ○ ○ ○
Pseudanophthalmus nelsoni Barr, 1965 Nelson's Cave Beetle Coleoptera Troglobiont G2 VA ○ 1 ○
Nelsonites jonesi Valentine, 1952 Coleoptera Troglobiont G2G3 KY ○ ○ ○
Nelsonites walteri Valentine, 1952 Coleoptera Troglobiont G3 TN ○ 3 ○
Neoallochernes incertus Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 NM ○ ○ ○
Neoallochernes stercoreus (Turk, 1949) Pseudoscorpiones Troglobiont G4 TX ○ 10 ○
Neochthonius troglodytes Muchmore, 1969 Pseudoscorpiones Troglobiont G2G3 CA ○ ○ ○
Nesticus barrowsi Gertsch, 1984 Araneae Troglobiont G1G2 TN ○ 4 9
Nesticus brimleyi Gertsch, 1984 Araneae Troglobiont G1G2 NC ○ 7 9
Nesticus cressleri Zigler & Milne, 2022 Araneae Troglobiont GNR GA ○ 2 8
Nesticus lula Zigler & Milne, 2022 Araneae Troglobiont GNR GA ○ 2 6
Nesticus paynei Gertsch, 1984 Araneae Troglobiont G3G4 TN, VA ○ 18 27
Nesticus potterius (Chamberlin, 1933) Araneae Troglobiont G1G2 CA ○ ○ ○
Nesticus stygius Gertsch, 1984 Araneae Troglobiont G1G2 TN ○ 3 7
Nevadesmus hubbsi (Chamberlin, 1943) Polydesmida Troglobiont GNR NV ○ ○ ○
Nevadesmus ophimontis Shear, 2009 Polydesmida Troglobiont NV ○ 1 ○
Stygobromus interitus Holsinger, 1978 New Castle Murder Hole Cave Amphipod Amphipoda Stygobiont G1G2 VA ○ 2 ○
Pseudanophthalmus egberti Barr, 1965 New River Valley Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Pseudanophthalmus nickajackensis Barr, 1981 Nickajack Cave Beetle Coleoptera Troglobiont G1 AL, TN ○ ○ ○
Caecidotea nickajackensis Packard, 1881 Nickajack Cave Isopod Isopoda Stygobiont G1 VU AL, TN ○ 9 ○
Niptus arcanus Aalbu & Andrews, 1992 Coleoptera Troglobiont CA ○ 18 ○
Nitocrellopsis texana Fiers & Iliffe, 2000 Harpacticoida Stygobiont GNR TX ○ 1 ○
Pseudanophthalmus paulus Barr, 1981 Nobletts Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Caecidotea nordeni Lewis & Bowman, 2010 Norden's Groundwater Isopod Isopoda Stygobiont GH MD ○ 2 ○
Troglocambarus maclanei Hobbs, 1942 North Florida Spider Cave Crayfish Decapoda Stygobiont G1G2 NT FL ○ 42 6
Amblyopsis spelaea DeKay, 1842 Northern Cavefish Percopsiformes Stygobiont G2G3 NT KY ○ 93 384
Cauloxenus stygius Cope, 1872 Northern Cavefish Copepod Siphonostomatoida Stygobiont G1G2 IN, KY ○ ○ ○
Eurycea nerea (Bishop, 1944) Northern Grotto Salamander Caudata Troglobiont GNR LC AR, MO ○ 67 144
Stygobromus phreaticus Holsinger, 1978 Northern Virginia Well Amphipod Amphipoda Stygobiont G1 VA ○ 4 ○
Stygobromus nortoni (Holsinger, 1969) Norton's Cave Amphipod Amphipoda Stygobiont G2G3 VU TN ○ 4 ○
Pseudanophthalmus nortoni Barr, 1981 Norton's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Caecidotea nortoni (Steeves, 1966) Norton's Cave Isopod Isopoda Stygobiont G2G3 TN ○ 6 ○
Phreatoceras taylori (Hershler & Longley, 1986) Nymph trumpet Littorinimorpha Stygobiont G1G2 DD TX ○ 20 5
Oaphantes prometheus Paquin et al., 2020 Araneae Troglobiont CA ○ 2 ○
Occasjapyx kofoidi (Silvestri, 1928) Dicellurata Troglobiont CA ○ ○ ○
Crangonyx ohioensis Zhang & Holsinger, 2003 Ohio Cave Amphipod Amphipoda Stygobiont G1G2 IN, KY, OH ○ 32 ○
Pseudanophthalmus ohioensis Krekeler, 1973 Ohio Cave Beetle Coleoptera Troglobiont G1 OH ○ 2 ○
Allocrangonyx pellucidus (Mackin, 1935) Oklahoma Cave Amphipod Amphipoda Stygobiont G2G3 VU OK ○ 12 ○
Cambarus tartarus Hobbs & Cooper, 1972 Oklahoma Cave Crayfish Decapoda Stygobiont G1 CR OK ○ 11 10
Pseudanophthalmus puteanus Krekeler, 1973 Old Well Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 1 ○
Oliarus gagneri Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus kalaupapae Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus lorettae Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus makaiki Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oliarus polyphemus Fennah, 1973 Homoptera Troglobiont G1G2 HI ○ 4 29
Oliarus priola Fennah, 1973 Homoptera Troglobiont G1G2 HI ○ 2 ○
Oliarus waikau Hoch & Howarth, 1999 Homoptera Troglobiont G1G2 HI ○ ○ ○
Oncopodura cruciata Bonet, 1943 Entomobryomorpha Troglobiont G1G2 MT ○ 10 ○
Oncopodura fenestra Christiansen & Bonet, 1996 Entomobryomorpha Troglobiont G2G3 TX ○ 1 ○
Oncopodura hubbardi Christiansen & Bellinger 1996 Entomobryomorpha Troglobiont G2 VA ○ 1 ○
Oncopodura iowae Christiansen, 1980 Entomobryomorpha Troglobiont G3G4 IA, IL, MO ○ 3 ○
Oncopodura prietoi Bonet, 1943 Entomobryomorpha Troglobiont G1G2 NM ○ 2 ○
Oncopodura subhoffi Christiansen & Bellinger, 1998 Entomobryomorpha Troglobiont G3 CO ○ 2 ○
Stygobromus onondagaensis (Hubricht & Mackin, 1940) Onondaga Cave Amphipod Amphipoda Stygobiont G3 VU AR, KS, MO, OK ○ 61 ○
Onychiurus gelus Christiansen & Bonet, 1980 Poduromorpha Troglobiont G2G3 IA, WA, WI ○ ○ ○
Onychiurus janus Christiansen & Bellinger, 1980 Poduromorpha Troglobiont G2G3 AL, VA, WV ○ ○ ○
Onychiurus obesus Mills, 1934 Poduromorpha Troglobiont G3G4 IA, MN, MO, WI ○ ○ ○
Onychiurus oregonensis Bonet, 1931 Poduromorpha Troglobiont G1G2 OR ○ ○ ○
Onychiurus paro Christiansen & Bellinger, 1980 Poduromorpha Troglobiont G1 AL ○ ○ ○
Onychiurus pipistrellae Soto-Adames & Taylor, 2013 Poduromorpha Troglobiont IL ○ ○ ○
Onychiurus reluctus Christiansen, 1961 Poduromorpha Troglobiont GNR WV ○ 9 ○
Opiona catorycha Shear & Crawford, 2019 Chordeumatida Troglobiont OR ○ ○ ○
Opiona graeningi Shear, 2011 Chordeumatida Troglobiont CA ○ 1 ○
Procambarus franzi Hobbs & Lee, 1976 Orange Lake Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 13 5
Phagocata oregonensis Hyman, 1963 Oregon Cave Flatworm Tricladida Stygobiont G1G2 OR ○ ○ ○
Oregonasellus elliotti Lewis, 2008 Isopoda Stygobiont OR ○ ○ ○
Kleptochthonius hetricki Muchmore, 1974 Organ Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Fontigens tartarea Hubtricht 1963 Organ cavesnail Littorinimorpha Stygobiont G2 VA, WV ○ 42 2
Procambarus acherontis (Lonnberg, 1894) Orlando Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 18 5
Pseudotremia orndorffi Shear, 2011 Orndorff's Cave Milliped Chordeumatida Troglobiont G2 VA ○ ○ ○
Kleptochthonius orpheus Muchmore, 1965 Orpheus Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Pseudanophthalmus praetermissus Barr, 1981 Overlooked Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Stygobromus ozarkensis (Holsinger, 1967) Ozark Cave Amphipod Amphipoda Stygobiont G4 VU AR, MO, OK ○ 59 1
Speleonycta ozarkensis Espinasa et al., 2010 Ozark Cave Bristletail Zygentoma Troglobiont AR, OK ○ ○ 1
Troglichthys rosae (Eigenmann, 1898) Ozark cavefish Percopsiformes Stygobiont G3 NT ESA AR, MO, OK ○ 4 ○
Ozarkia alabama (Gertsch, 1974) Araneae Troglobiont GNR AL ○ 27 ○
Ozarkia arkansa (Gertsch, 1974) Araneae Troglobiont AR ○ 8 ○
Ozarkia blanda (Gertsch, 1974) Araneae Troglobiont GNR AL ○ 1 ○
Ozarkia georgia (Gertsch, 1974) Araneae Troglobiont GNR GA ○ 25 ○
Ozarkia novaegalleciae (Brignoli, 1979) Araneae Troglobiont NM ○ ○ ○
Ozarkia serena (Gertsch, 1974) Araneae Troglobiont G1G2 AL ○ 15 ○
Pacificabathynella kalispellensis Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Pacificabathynella ruthae Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Pacificabathynella sequoiae Schminke & Noodt, 1988 Bathynellacea Stygobiont GNR CA ○ 5 ○
Pacificabathynella stanfordi Camacho et al., 2009 Bathynellacea Stygobiont GNR MT ○ ○ ○
Pacificabathynella yupik Camacho et al., 2016 Bathynellacea Stygobiont AK ○ 47 ○
Zygonopus packardi Causey, 1960 Packard's Blind Cave Millipede Chordeumatida Troglobiont G4 VA, WV ○ ○ ○
Crangonyx packardi Smith, 1888 Packard's Cave Amphipod Amphipoda Stygobiont G4 IL, IN, KS, KY, MO ○ 103 ○
Pseudanophthalmus packardi Barr, 1959 Packard's Cave Beetle Coleoptera Troglobiont G1G2 KY ○ ○ ○
Kleptochthonius packardi (Hagen, 1879) Packard's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G2 IN ○ 6 ○
Pseudanophthalmus pallidus Barr, 1981 Pale Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Paleonura petebellingeri Palacios-Vargas & Benito, 2007 Poduromorpha Troglobiont G1 VA ○ ○ ○
Procambarus pallidus (Hobbs, 1940) Pallid Cave Crayfish Decapoda Stygobiont G1G2 NT FL, GA ○ 84 5
Parabogidiella americana Holsinger, 1980 Amphipoda Stygobiont G2G3 TX ○ 65 4
Paraholsingerius smaragdinus (Holsinger, 1992) Amphipoda Stygobiont GNR TX ○ 17 ○
Paramoera lokowai Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 6 ○
Paramoera paakai Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 17 ○
Paramoera rua Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 11 ○
Pararhynchelmis texana Fend, 2023 Lumbriculida Stygobiont TX ○ 7 ○
Paratachys aaa Liebherr, 2021 Coleoptera Troglobiont HI ○ ○ ○
Parobisium charlotteae Chamberlin, 1962 Pseudoscorpiones Troglobiont G1G2 OR ○ 3 6
Parobisium petilum Harvey & Cullen, 2020 Pseudoscorpiones Troglobiont CA ○ 1 ○
Parobisium yosemite Cokendolpher & Krejca, 2010 Pseudoscorpiones Troglobiont CA ○ ○ ○
Parvidrilus strayeri Erseus, 1999 Lumbriculida Stygobiont G1G2 AL ○ 17 ○
Stygobromus redactus Holsinger, 1978 Pattons Cave Amphipod Amphipoda Stygobiont G1 WV ○ 3 1
Pseudanophthalmus paynei Barr, 1981 Payne's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Stygobromus pecki (Holsinger, 1967) Peck's Cave Amphipod Amphipoda Stygobiont G1G2 EN ESA TX ○ 36 33
Scoterpes stewartpecki Shear, 2010 Peck's Cave Millipede Chordeumatida Troglobiont AL, TN ○ ○ ○
Tyrannochthonius pecki Muchmore, 1996 Peck's Cave Peudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Pseudosinella pecki Christiansen & Bellinger, 1980 Peck's Cave Springtail Entomobryomorpha Troglobiont G2G3 GA, NC, TN ○ 1 ○
Stygobromus pennaki (Ward, 1977) Pennak Cave Amphipod Amphipoda Stygobiont G2G3 CO ○ 9 ○
Crangonyx dearolfi Shoemaker, 1942 Pennsylvania Cave Amphipod Amphipoda Stygobiont G2 EN MD, PA ○ 14 ○
Sphalloplana evaginata Kenk, 1977 Perryville Cave Flatworm Tricladida Stygobiont G1G2 MO ○ 20 ○
Pseudanophthalmus petrunkevitchi Valentine, 1945 Petrunkevitch's Cave Beetle Coleoptera Troglobiont G1G2 VA ○ 1 ○
Phagocata angusta Kenk, 1977 Tricladida Stygobiont G1 WV ○ 7 ○
Phagocata carolinensis Kenk, 1979 Tricladida Stygobiont G1G2 NC ○ 4 ○
Phagocata notorchis Kenk, 1987 Tricladida Stygobiont TN ○ 4 ○
Phagocata procera Kenk, 1984 Tricladida Stygobiont G1G2 NC ○ 14 ○
Phagocata pygmaea Kenk, 1987 Tricladida Stygobiont NC ○ 2 ○
Phagocata spuria Kenk, 1987 Tricladida Stygobiont NC ○ 3 ○
Phalangodes armata Tellkampf, 1844 Opiliones Troglobiont G3G4 KY ○ 15 4
Cambarus pecki (Hobbs, 1967) Phantom Cave Crayfish Decapoda Stygobiont G1G2 EN AL ○ 16 10
Phreatodrobia bulla Perez & Castañeda, 2023 Littorinimorpha Stygobiont TX ○ ○ 4
Phreatodrobia coronae Hershler, 1987 Littorinimorpha Stygobiont G1G2 DD TX ○ 9 ○
Stygobromus heteropodus Hubricht, 1943 Pickle Springs Amphipod Amphipoda Stygobiont G1G2 VU MO ○ 2 ○
Kenkia glandulosa Hyman, 1956 Pink Cave Flatworm Tricladida Stygobiont G2G3 IA, MO ○ 11 ○
Stygobromus obrutus Holsinger, 1978 Pittsylvania Well Amphipod Amphipoda Stygobiont G1G2 VA ○ 2 ○
Stygobromus pizzinii (Shoemaker, 1938 Pizzini's Cave Amphipod Amphipoda Stygobiont G3G4 VU DC, MD, PA, VA ○ 89 3
Plumatyla humerosa (Loomis, 1943) Chordeumatida Troglobiont G2G3 CA ○ 2 ○
Plutomurus californicus (Folson, 1913) Entomobryomorpha Troglobiont G2G3 CA ○ 13 ○
Plutomurus grahami (Christiansen, 1980) Entomobryomorpha Troglobiont G1 NM ○ 2 ○
Poecilophysis extraneostella Zacharda 1985 Actinedida Troglobiont G2 VA, WV ○ ○ ○
Poecilophysis weyerensis (Packard, 1888) Actinedida Troglobiont G3 KY, TN, VA, WV ○ ○ ○
Sphalloplana consimilis Kenk, 1977 Powell Valley Cave Flatworm Tricladida Stygobiont G2G3 TN, VA ○ 13 ○
Amerigoniscus henroti Vandel, 1950 Powell Valley Terrestrial Cave Isopod Isopoda Troglobiont G3 VA ○ 10 ○
Pratherodesmus despaini Shear, 2009 Polydesmida Troglobiont CA ○ 1 ○
Pratherodesmus ecclesia Shear, 2009 Polydesmida Troglobiont AZ ○ 1 ○
Pratherodesmus voylesi Shear, 2009 Polydesmida Troglobiont AZ ○ 1 ○
Stygobromus paxillus Holsinger, 2011 Prettyboy Groundwater Amphipod Amphipoda Stygobiont G1 MD ○ 3 ○
Conasellus pricei (Levi, 1949) Price's Cave Isopod Isopoda Stygobiont G5 MD, PA, VA, WV ○ ○ ○
Cambarus hamulatus (Cope & Packard, 1881) Prickly Cave Crayfish Decapoda Stygobiont G3G4 LC AL, TN ○ 46 58
Procaris hawaiana Holthuis, 1973 Decapoda Stygobiont G1 ESA HI ○ 14 4
Kleptochthonius proserpinae Muchmore, 1965 Proserpina Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Fontigens proserpina Hubricht, 1940 Proserpine Springsnail Littorinimorpha Stygobiont G1 MO ○ 20 ○
Proteocephalus poulsoni Whitaker & Zober, 1978 Proteocephalidea Stygobiont KY ○ ○ ○
Protolimnesia ventriplacophora Smith & Cook, 1996 Trombidiformes Stygobiont TX ○ ○ ○
Pseudanophthalmus barberi Jeannel, 1931 Coleoptera Troglobiont G2G3 KY ○ 2 3
Pseudanophthalmus barri Krekeler, 1973 Coleoptera Troglobiont G1G2 IN ○ 2 ○
Pseudanophthalmus cerberus Barr, 1985 Coleoptera Troglobiont G3G4 KY, TN ○ 27 ○
Pseudanophthalmus chthonius Krekeler, 1973 Coleoptera Troglobiont G3 IN ○ 2 ○
Pseudanophthalmus ciliaris Valentine, 1937 Coleoptera Troglobiont G3G4 KY, TN ○ 5 15
Pseudanophthalmus cumberlandus Valentine, 1937 Coleoptera Troglobiont G1G2 TN ○ 1 ○
Pseudanophthalmus darlingtoni Barr, 1985 Coleoptera Troglobiont G3 KY ○ ○ ○
Pseudanophthalmus desertus Krekeler, 1973 Coleoptera Troglobiont G3 KY ○ 4 ○
Pseudanophthalmus digitus Valentine, 1932 Coleoptera Troglobiont G1G2 GA, TN ○ 1 ○
Pseudanophthalmus distinguens Valentine, 1948 Coleoptera Troglobiont G1G2 AL, VA ○ ○ ○
Pseudanophthalmus elongatus Krekeler, 1973 Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus emersoni Krekeler, 1958 Coleoptera Troglobiont G1 IN ○ 2 ○
Pseudanophthalmus eremita (Horn, 1871) Coleoptera Troglobiont G1G2 IN ○ 2 ○
Pseudanophthalmus exiguus Krekeler, 1973 Coleoptera Troglobiont G3G4 KY ○ 4 ○
Pseudanophthalmus farrelli Barr, 1959 Coleoptera Troglobiont G1G2 TN ○ 2 ○
Pseudanophthalmus fluviatilis Valentine, 1948 Coleoptera Troglobiont G3 AL ○ ○ ○
Pseudanophthalmus fulleri Valentine, 1932 Coleoptera Troglobiont G2G3 GA, TN ○ 1 ○
Pseudanophthalmus fuscus Valentine, 1931 Coleoptera Troglobiont G4 WV ○ 5 ○
Pseudanophthalmus globiceps Barr, 1985 Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus grandis Valentine, 1931 Coleoptera Troglobiont G4 WV ○ 9 ○
Pseudanophthalmus henroti Jeannel, 1949 Coleoptera Troglobiont GNR WV ○ ○ ○
Pseudanophthalmus hesperus Barr, 1959 Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus higginbothami Valentine, 1931 Coleoptera Troglobiont G2 WV ○ 2 ○
Pseudanophthalmus humeralis Valentine, 1931 Coleoptera Troglobiont G2 TN ○ 2 ○
Pseudanophthalmus hypertrichosis Valentine, 1932 Coleoptera Troglobiont G3 WV ○ 2 ○
Pseudanophthalmus hypolithos Barr, 1981 Coleoptera Troglobiont G1 KY ○ ○ ○
Pseudanophthalmus lallemanti Jeannel, 1949 Coleoptera Troglobiont G1 WV ○ ○ ○
Pseudanophthalmus leonae Barr, 1960 Coleoptera Troglobiont G1 IN ○ ○ ○
Pseudanophthalmus lodingi Valentine, 1931 Coleoptera Troglobiont G1G2 AL ○ 6 ○
Pseudanophthalmus loganensis Barr, 1959 Coleoptera Troglobiont G2G3 KY, TN ○ ○ 10
Pseudanophthalmus menetriesi (Motschulsky, 1862) Coleoptera Troglobiont G3 KY ○ ○ ○
Pseudanophthalmus meridionalis Valentine, 1945 Coleoptera Troglobiont G2 AL ○ ○ ○
Pseudanophthalmus orthosulcatus Valentine, 1932 Coleoptera Troglobiont G1 WV ○ 1 ○
Pseudanophthalmus pilosus Barr, 1985 Coleoptera Troglobiont G2G3 KY ○ ○ ○
Pseudanophthalmus princeps Barr, 1979 Coleoptera Troglobiont G1G2 KY, TN ○ 3 4
Pseudanophthalmus productus Barr, 1980 Coleoptera Troglobiont G3G4 TN ○ ○ ○
Pseudanophthalmus profundus Valentine, 1945 Coleoptera Troglobiont G2 AL ○ ○ ○
Pseudanophthalmus pubescens (Horn, 1868) Coleoptera Troglobiont G3 KY ○ 24 6
Pseudanophthalmus rittmani Krekeler, 1973 Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus scutilis Barr, 1981 Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus solivagus Krekeler, 1973 Coleoptera Troglobiont G1G2 KY ○ 2 ○
Pseudanophthalmus striatus (Motschulsky, 1862) Coleoptera Troglobiont G2G3 KY ○ 5 ○
Pseudanophthalmus tennesseensis Valentine, 1937 Coleoptera Troglobiont G3 TN ○ 1 ○
Pseudanophthalmus tenuis (Horn, 1871) Coleoptera Troglobiont G4 IN ○ 12 ○
Pseudanophthalmus transfluvialis Barr, 1985 Coleoptera Troglobiont G1G2 KY ○ ○ ○
Pseudanophthalmus umbratilis Krekeler, 1973 Coleoptera Troglobiont G2G3 KY ○ 2 ○
Pseudanophthalmus vanburenensis Barr, 1962 Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudobaicalasellus magniezi Lewis & Lewis, 2023 Isopoda Stygobiont VA ○ ○ ○
Pseudobaicalasellus mausi (Lewis, 2009) Isopoda Stygobiont G2 MD, VA ○ ○ ○
Pseudogarypus hypogeus Muchmore, 1981 Pseudoscorpiones Troglobiont G1G2 AZ ○ ○ ○
Pseudogarypus speleus Benedict & Malcolm, 1978 Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Pseudosinella bona Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G2 VA ○ 1 ○
Pseudosinella dubia Christiansen, 1961 Entomobryomorpha Troglobiont G3 AR, OK ○ 10 ○
Pseudosinella erehwon Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G2 VA ○ 2 ○
Pseudosinella espanita Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G1 KY, TN ○ 2 ○
Pseudosinella extra Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G1G2 VA ○ 1 ○
Pseudosinella flatua Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G1G2 NC, VA ○ 2 ○
Pseudosinella gisini Charistiansen, 1960 Entomobryomorpha Troglobiont G3G4 NC, VA, WV ○ 15 ○
Pseudosinella granda Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G3 VA ○ 1 ○
Pseudosinella josemarii Soto-Adames 2010 Entomobryomorpha Troglobiont GNR WV ○ ○ ○
Pseudosinella meganporteri Soto-Adames 2010 Entomobryomorpha Troglobiont G2 WV ○ ○ ○
Pseudosinella nata Christiansen & Bellinger, 1980 Entomobryomorpha Troglobiont G1 AL ○ 2 ○
Pseudosinella ops Christiansen & Bellinger, 1998 Entomobryomorpha Troglobiont VA ○ 1 ○
Pseudosinella orba Christiansen, 1960 Entomobryomorpha Troglobiont G3G4 KY, NC, TN, VA, WV ○ 65 ○
Pseudosinella vespera Christiansen & Bellinger, 1996 Entomobryomorpha Troglobiont G2 NC, VA ○ 1 ○
Pseudosinella vita Christiansen & Bonet, 1980 Entomobryomorpha Troglobiont G1G2 NM ○ 14 ○
Pseudotremia acheron Shear, 1972 Chordeumatida Troglobiont G1G2 TN ○ 2 ○
Pseudotremia aeacus Shear, 1972 Chordeumatida Troglobiont G1G2 GA ○ 3 ○
Pseudotremia amphiorax Shear, 1972 Chordeumatida Troglobiont G1G2 KY ○ 2 ○
Pseudotremia carterensis (Packard, 1883) Chordeumatida Troglobiont G2G3 KY ○ 2 ○
Pseudotremia cercops Shear, 1972 Chordeumatida Troglobiont G1 TN ○ 3 ○
Pseudotremia deprehendor Shear, 1972 Chordeumatida Troglobiont G1 TN, VA ○ 2 ○
Pseudotremia eburnea Loomis, 1939 Chordeumatida Troglobiont G2G4 AL, GA, TN ○ 3 ○
Pseudotremia fongi Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia handi Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia kerboi Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia lethe Shear, 1972 Chordeumatida Troglobiont G1 TN ○ 1 ○
Pseudotremia lictor Shear, 1972 Chordeumatida Troglobiont G1 TN ○ 2 ○
Pseudotremia merops Shear, 1972 Chordeumatida Troglobiont G1 KY ○ 2 ○
Pseudotremia minos Shear, 1972 Chordeumatida Troglobiont G1 AL, TN ○ 2 10
Pseudotremia nodosa Loomis, 1939 Chordeumatida Troglobiont G4 TN, VA ○ 2 ○
Pseudotremia nyx Shear, 1972 Chordeumatida Troglobiont G1 AL ○ 3 ○
Pseudotremia rhadamanthus Shear, 1972 Chordeumatida Troglobiont G1G2 AL, TN ○ 2 ○
Pseudotremia schneiderae Shear, 2008 Chordeumatida Troglobiont GNR WV ○ ○ ○
Pseudotremia spira Shear, 1972 Chordeumatida Troglobiont G1 KY ○ 2 ○
Pseudotremia styx Lewis, 2009 Chordeumatida Troglobiont GNR KY ○ ○ ○
Pseudotremia tuberculata Loomis, 1939 Chordeumatida Troglobiont G3 VA ○ 1 ○
Pseudotremia unca Shear, 1972 Chordeumatida Troglobiont G1 KY ○ 2 ○
Pseudotremia valga Loomis, 1943 Chordeumatida Troglobiont G1G2 TN, VA ○ 2 ○
Psychopomporus felipi Jean et al., 2012 Coleoptera Stygobiont GNR TX ○ 3 3
Ptomaphagus barri Peck, 1973 Coleoptera Troglobiont G2G3 TN ○ 5 12
Ptomaphagus chromolithus Peck, 1984 Coleoptera Troglobiont G2G3 AL, TN ○ ○ 6
Ptomaphagus cocytus Peck, 1973 Coleoptera Troglobiont G1G2 AZ ○ 2 ○
Ptomaphagus episcopus Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 3
Ptomaphagus fecundus Barr, 1963 Coleoptera Troglobiont G1 TN ○ ○ 3
Ptomaphagus fiskei Peck, 1973 Coleoptera Troglobiont G1G2 GA ○ 5 ○
Ptomaphagus hazelae Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 ○
Ptomaphagus hubrichti Barr, 1958 Coleoptera Troglobiont G2G3 TN ○ 4 ○
Ptomaphagus inyoensis Peck & Gnaspini, 1997 Coleoptera Troglobiont CA ○ ○ ○
Ptomaphagus julius Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 6
Ptomaphagus laticornis Jeannel, 1949 Coleoptera Troglobiont G1G2 AL ○ ○ 6
Ptomaphagus loedingi (Hatch, 1933) Coleoptera Troglobiont G1G2 AL ○ 5 ○
Ptomaphagus longicornis Jeannel, 1949 Coleoptera Troglobiont G3G4 AL ○ ○ 9
Ptomaphagus parashant Peck & Wynne, 2013 Coleoptera Troglobiont AZ ○ 2 ○
Ptomaphagus solanum Peck, 1973 Coleoptera Troglobiont G1 AL ○ 2 3
Ptomaphagus walteri Peck, 1973 Coleoptera Troglobiont G1G2 AL ○ 2 6
Ptomaphagus whiteselli Barr, 1963 Coleoptera Troglobiont G2G3 AL, GA ○ 5 ○
Litocampa pucketti Conde & Bareth, 1996 Puckett's Cave Dipluran Rhabdura Troglobiont G1G2 VA ○ ○ ○
Pseudotremia peponocranium Shear, 2011 Pumpkin-headed Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Calathaemon holthuisi (Strenth, 1976) Purgatory Cave Shrimp Decapoda Stygobiont G1G2 EN TX ○ 11 ○
Pseudotremia purselli Lewis, 2000 Pursell's Cave Milliped Chordeumatida Troglobiont G1 IN ○ ○ ○
Procambarus morrisi Hobbs & Franz, 1991 Putnum County Cave Crayfish Decapoda Stygobiont G1 CR FL ○ 13 5
Pygmarrhopalites altus (Christiansen, 1966) Symphypleona Troglobiont G2G3 KY ○ 2 ○
Pygmarrhopalites arcus (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont GNR ID ○ 5 ○
Pygmarrhopalites ashcraftensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont G1 IN ○ 2 ○
Pygmarrhopalites bimus (Christiansen, 1966) Symphypleona Troglobiont G3G4 IA, IN, KY ○ 3 ○
Pygmarrhopalites buffaloensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont AR ○ 2 ○
Pygmarrhopalites caedus (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G3 VA ○ ○ ○
Pygmarrhopalites commorus (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G2G3 VA, WV ○ ○ ○
Pygmarrhopalites dubius (Christiansen, 1966) Symphypleona Troglobiont G2G3 AR, IA, MN ○ 3 ○
Pygmarrhopalites fransjanssens Soto-Adames & Taylor, 2013 Symphypleona Troglobiont IL ○ ○ ○
Pygmarrhopalites hubbardi (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont CO ○ 2 ○
Pygmarrhopalites incantator Soto-Adames & Taylor, 2013 Symphypleona Troglobiont IL ○ 3 ○
Pygmarrhopalites jay Christiansen & Bonet, 1996 Symphypleona Troglobiont G2G3 OK ○ ○ ○
Pygmarrhopalites lacuna (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G1G2 VA ○ ○ ○
Pygmarrhopalites leonardwoodensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont MO ○ 2 ○
Pygmarrhopalites marshalli (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G3 TN, VA ○ 4 ○
Pygmarrhopalites obtusus (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont G1 VA ○ 7 ○
Pygmarrhopalites pavo (Christiansen & Bellinger, 1998) Symphypleona Troglobiont G3 MO, TN, VA, WV ○ 6 ○
Pygmarrhopalites plethorasari Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont MO ○ 6 ○
Pygmarrhopalites sacer (Christiansen & Bellinger, 1998) Symphypleona Troglobiont G2 VA, WV ○ ○ ○
Pygmarrhopalites salemensis Soto-Adames & Taylor, 2013 Symphypleona Troglobiont IL ○ ○ ○
Pygmarrhopalites sapo (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont IL ○ 8 ○
Pygmarrhopalites sextus (Zeppelini & Christiansen, 2003) Symphypleona Troglobiont G1 VA ○ 7 ○
Pygmarrhopalites shoshoneiensis Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont NV ○ 4 ○
Pygmarrhopalites silvus (Christiansen & Bellinger, 1996) Symphypleona Troglobiont G2 VA ○ ○ ○
Pygmarrhopalites texensis (Christiansen & Bonet, 1996) Symphypleona Troglobiont G3G4 TX ○ ○ ○
Pygmarrhopalites youngsteadtii Zeppelini, Taylor, & Slay, 2009 Symphypleona Troglobiont AR ○ ○ ○
Stygobromus quatsinensis Holsinger & Shaw, 1987 Quatsino Cave Amphipod Amphipoda Stygobiont G1G2 AK, BC ○ 21 ○
Helicodiscus barri Hubricht, 1962 Raccoon coil Stylommatophora Troglobiont G3G4 AL, GA, TN ○ 13 106
Nesticus furtivus Gertsch, 1984 Racoon Mountain Caverns Cave Spider Araneae Troglobiont G1 TN ○ 2 7
Miktoniscus oklahomensis Vandel, 1965 Racovitza's Cave Isopod Isopoda Troglobiont G1G2 OK ○ ○ ○
Miktoniscus racovitzai Vandel, 1950 Racovitza's Terrestrial Cave Isopod Isopoda Troglobiont G3G4 VA ○ 20 ○
Stygobromus foliatus Holsinger, 2011 Rappahannock Spring Amphipod Amphipoda Stygobiont G2 MD, VA ○ 10 2
Speodesmus reddelli Elliott, 2004 Reddell's Cave Millipede Polydesmida Troglobiont GNR TX ○ 71 ○
Scoterpes willreevesi Shear, 2010 Reeves' Cave Millipede Chordeumatida Troglobiont AL, GA ○ ○ ○
Sphalloplana pricei Hyman, 1937 Refton Cave Flatworm Tricladida Stygobiont G2G3 MD, PA ○ 12 ○
Remasellus parvus (Steeves, 1964) Isopoda Stygobiont G1G2 FL ○ 13 16
Pseudotremia reynoldsae Lewis, 2003 Reynolds Cave Millipede Chordeumatida Troglobiont G1 IN ○ ○ ○
Rhadine austinica Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 82 18
Rhadine bullis Reddell & Cokendolpher, 2004 Coleoptera Troglobiont GNR TX ○ 22 ○
Rhadine exilis (Barr & Lawrence, 1960) Coleoptera Troglobiont G3 ESA TX ○ 133 12
Rhadine grubbsi Reddell & Duperre, 2009 Coleoptera Troglobiont TX ○ 7 ○
Rhadine infernalis (Barr & Lawrence, 1960) Coleoptera Troglobiont G2G3 ESA TX ○ 152 17
Rhadine insolita Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 5 ○
Rhadine ivyi Reddell & Cokendolpher, 2004 Coleoptera Troglobiont TX ○ 2 ○
Rhadine koepkei (Barr, 1960) Coleoptera Troglobiont G1 TX ○ 12 ○
Rhadine noctivaga Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 94 14
Rhadine ozarkensis Sanderson & Miller, 1941 Coleoptera Troglobiont GH AR ○ 2 ○
Rhadine reyesi Reddell & Cokendolpher, 2001 Coleoptera Troglobiont GNR TX ○ 36 ○
Rhadine russelli Barr, 1974 Coleoptera Troglobiont G1G2 TX ○ 13 3
Rhadine speca (Barr, 1960) Coleoptera Troglobiont G2 TX ○ ○ ○
Rhadine sprousei Reddell & Cokendolpher, 2004 Coleoptera Troglobiont TX ○ 3 ○
Rhadine subterranea (Van Dyke, 1918) Coleoptera Troglobiont G2 TX ○ 143 15
Rhadine tenebrosa (Barr, 1960) Coleoptera Troglobiont G1G2 TX ○ 22 5
Rhagidia varia Zacharda 1985 Actinedida Troglobiont G5 VA, WV ○ ○ ○
Rheocyclops virginianus (Reid, 1993) Cyclopoida Stygobiont VA ○ 5 ○
Pseudanophthalmus krekeleri Barr, 1965 Rich Mountain Cave Beetle Coleoptera Troglobiont GX WV ○ 1 ○
Pseudanophthalmus paradoxus Barr, 1981 Ridetop Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Cicurina baronia Gertsch, 1992 Robber Baron Cave Meshweaver Araneae Troglobiont G1 ESA TX ○ 12 ○
Pseudanophthalmus robustus Valentine, 1931 Robust Cave Beetle Coleoptera Troglobiont G3G4 TN ○ 7 4
Robustocheles occulta Zacharda & Pugsley, 1988 Actinedida Troglobiont G2G3 AB, IA, WA ○ ○ ○
Stygobromus kenki Holsinger, 1978 Rock Creek Groundwater Amphipod Amphipoda Stygobiont G2 DC, MD, VA ○ 15 ○
Stygobromus baroodyi Holsinger, 1978 Rockbridge County Cave Amphipod Amphipoda Stygobiont G1 VA ○ 11 1
Sphalloplana virginiana Hyman, 1945 Rockbridge County Cave Flatworm Tricladida Stygobiont G1 VA ○ 1 ○
Pseudotremia roebuckorum Lewis, 2005 Roebuck's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Pseudanophthalmus rogersae Barr, 1981 Roger's Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Rotomelita ana Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 3 ○
Rotomelita lokoa Barnard, 1977 Amphipoda Stygobiont G1G2 HI ○ 13 ○
Pseudanophthalmus rotundatus Valentine, 1932 Rotund Cave Beetle Coleoptera Troglobiont G2 TN, VA ○ ○ ○
Stygobromus mausi Holsinger, 2009 Round Hill Cave Amphipod Amphipoda Stygobiont G1 VA ○ 2 ○
Chitrella regina Malcolm & Chamberline, 1960 Royal Syarinid Pseudoscorpion Pseudoscorpiones Troglobiont G1 WV ○ ○ ○
Stygobromus lucifugus (Hay, 1882) Rubious Cave Amphipod Amphipoda Stygobiont G1G2 EX IL ○ ○ ○
Banksula rudolphi Briggs & Ubick, 1981 Rudolph's Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Stygobromus rudolphi Wang & Holsinger, 2001 Rudolph's Groundwater Amphipod Amphipoda Stygobiont G1 CA ○ 3 ○
Paramexiweckelia ruffoi Holsinger, 1993 Ruffo's Cave Amphipod Amphipoda Stygobiont G1G2 TX ○ 40 ○
Rugosuscandona scharfi Kulkoyluoglu et al., 2017 Podocopida Stygobiont TX ○ ○ ○
Apochthonius russelli Muchmore, 1976 Russell Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Stygobromus russelli (Holsinger, 1967) Russell's Cave Amphipod Amphipoda Stygobiont G4 TX ○ 396 11
Pseudanophthalmus seclusus Barr, 1981 Rye Cove Cave Beetle Coleoptera Troglobiont G2G3 VA ○ ○ ○
Lirceus culveri Estes & Holsinger, 1976 Rye Cove Cave Isopod Isopoda Stygobiont G1 VU VA ○ 9 9
Pseudotremia ryensis Shear, 2011 Rye Cove Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Sagittocythere barri (Hart & Hobbs, 1961) Podocopida Stygobiont G5 AL, IN, KY, TN ○ 76 ○
Sagittocythere stygia Hart & Hart, 1966 Podocopida Stygobiont G1 KY ○ 1 ○
Pseudanophthalmus sanctipauli Barr, 1981 Saint Paul Cave Beetle Coleoptera Troglobiont G1G2 VA ○ ○ ○
Cambarus hubrichti Hobbs, 1952 Salem Cave Crayfish Decapoda Stygobiont G4 DD MO ○ 21 23
Caecidotea salemensis Lewis, 1981 Salem Cave Isopod Isopoda Stygobiont G4 AR, MO ○ 33 4
Typhlichthys eigenmanni Charlton, 1933 Salem Plateau Cavefish Percopsiformes Stygobiont G3 AR, MO ○ 33 148
Pseudotremia salisae Lewis, 2000 Salisa's Cave Milliped Chordeumatida Troglobiont G2 IN ○ 3 ○
Diacyclops salisae Reid, 2004 Salisa's Groundwater Copepod Cyclopoida Stygobiont G1 IN ○ ○ ○
Salmasellus howarthi Lewis, 2001 Isopoda Stygobiont GNR OR, WA ○ 4 9
Salmasellus steganothrix Bowman, 1975 Isopoda Stygobiont G2 AB, MT ○ 15 ○
Procambarus erythrops Relyea & Sutton, 1975 Santa Fe Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 27 6
Schaefferia asymmetrica (Christiansen & Wang, 2006) Poduromorpha Troglobiont CO ○ ○ ○
Schaefferia steinmanni (Christiansen & Wang, 2006) Poduromorpha Troglobiont CO ○ ○ ○
Schaefferia unica (Christiansen & Wang, 2006) Poduromorpha Troglobiont CO ○ ○ ○
Pseudanophthalmus scholasticus Barr, 1981 Schoolhouse Cave Beetle Coleoptera Troglobiont G1 KY ○ ○ ○
Schornikovdona bellensis Kulkoyluoglu et al., 2017 Podocopida Stygobiont TX ○ ○ ○
Schrankia howarthi Davis & Medeiros, 2009 Lepidoptera Troglobiont HI ○ 3 105
Sclerobunus cavicolens (Banks, 1905) Opiliones Troglobiont G1G2 MT ○ 8 24
Sclerobunus jemez Derkarabetian & Hedin, 2014 Opiliones Troglobiont NV ○ ○ ○
Sclerobunus klomax Derkarabetian & Hedin, 2014 Opiliones Troglobiont GNR NM ○ 5 16
Sclerobunus madhousensis (Briggs, 1971) Opiliones Troglobiont G1G2 UT ○ 4 20
Sclerobunus speoventus Derkarabetian & Hedin, 2014 Opiliones Troglobiont CO ○ 15 32
Sclerobunus steinmanni Derkarabetian & Hedin, 2014 Opiliones Troglobiont CO ○ 3 21
Sclerobunus ungulatus (Briggs, 1971) Opiliones Troglobiont G1G2 NV ○ 29 21
Scoterpes austrinus Loomis, 1943 Chordeumatida Troglobiont G3G4 AL, GA ○ 3 ○
Scoterpes blountensis Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes copei (Packard, 1871) Chordeumatida Troglobiont G3G4 KY, TN ○ 6 ○
Scoterpes hesperus Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes jackdanieli Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes musicarustica Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes nudus Chamberlin, 1943 Chordeumatida Troglobiont GA ○ ○ ○
Scoterpes syntheticus (Shear, 1972) Chordeumatida Troglobiont AL ○ ○ ○
Scoterpes tricorner Shear, 2010 Chordeumatida Troglobiont TN ○ ○ ○
Scoterpes ventus Shear, 1972 Chordeumatida Troglobiont G3 TN ○ 1 ○
Seborgia hershleri Holsinger, 1992 Amphipoda Stygobiont G1G2 TX ○ 38 ○
Seborgia relicta Holsinger, 1980 Amphipoda Stygobiont G2G3 TX ○ 96 ○
Stygobromus mysticus Holsinger, 1974 Secret Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Pseudanophthalmus senecae Valentine, 1932 Seneca Cave Beetle Coleoptera Troglobiont G1 WV ○ 1 ○
Bowmanasellus sequoiae (Bowman, 1975) Sequoia Cave Isopod Isopoda Stygobiont G2 CA ○ ○ ○
Sequoiadesmus krejcae Shear & Shelley, 2008 Polydesmida Troglobiont CA ○ 1 ○
Pseudanophthalmus sequoyah Barr, 1981 Sequoyah Cave Beetle Coleoptera Troglobiont G1 AL ○ ○ ○
Caecidotea serrata (Fleming, 1972) Serrated Cave Isopod Isopoda Stygobiont G1G2 MO ○ 4 ○
Antroselates spiralis Hubricht, 1963 Shaggy cavesnail Littorinimorpha Stygobiont G3 IN, KY ○ 34 3
Dasyscias franzi F. G. Thompson & Hershler, 1991 Shaggy ghostsnail Littorinimorpha Stygobiont G1 FL ○ 8 ○
Pseudotremia sheari Lewis, 2009 Shear's Cave Millipede Chordeumatida Troglobiont GNR TN ○ ○ ○
Stygobromus sheldoni Holsinger, 1974 Sheldon's Groundwater Amphipod Amphipoda Stygobiont G1 CA ○ 15 ○
Pseudosinella testa Christiansen & Bellinger, 1980 Shelled Cave Springtail Entomobryomorpha Troglobiont G2G3 AR, VA, WV ○ 4 ○
Pseudotremia shelleyi Lewis, 2009 Shelley's Cave Millipede Chordeumatida Troglobiont GNR NC ○ 3 ○
Cambarus sheltae (Cooper & Cooper, 1997) Shelta Cave Crayfish Decapoda Stygobiont G1 CR AL ○ ○ ○
Tyrannochthonius sheltae Muchmore, 1996 Shelta Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Pseudanophthalmus shilohensis Krekeler, 1958 Shiloh Cave Beetle Coleoptera Troglobiont G2 IN ○ 2 ○
Texella shoshone Ubick & Briggs, 1992 Shoshone Cave Harvestman Opiliones Troglobiont G1 CA ○ 1 ○
Hubbardia shoshonensis Briggs & Hom, 1972 Shoshone Cave Whipscorpion Schizomida Troglobiont G1 CA ○ 1 ○
Speodesmus falcatus Elliott, 2004 Sickled Cave Millipede Polydesmida Troglobiont GNR TX ○ 29 ○
Stygobromus sierrensis Holsinger, 1974 Sierra Groundwater Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Pseudanophthalmus sericus Barr, 1981 Silken Cave Beetle Coleoptera Troglobiont G1 VA ○ ○ ○
Procambarus attiguus Hobbs & Franz, 1992 Silver Glen Springs Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 8 5
Stygobromus simplex Wang & Holsinger, 2001 Simple Cave Amphipod Amphipoda Stygobiont G1G2 CO ○ 1 ○
Pseudanophthalmus simplex Barr, 1980 Simple Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Sinella agna Christiansen & Bellinger, 1980 Entomobryomorpha Troglobiont G3G4 WV ○ 2 ○
Sinella alata Christiansen, 1960 Entomobryomorpha Troglobiont G5 IN ○ 1 ○
Sinella baca Christiansen & Bonet, 1980 Entomobryomorpha Troglobiont G3G4 BC, CA, TX ○ 2 ○
Sinella basidens Bonet, 1934 Entomobryomorpha Troglobiont G3G4 IN, KY, TN ○ 3 ○
Sinella cavernarum (Packard, 1888) Entomobryomorpha Troglobiont G5 IL, IN, KY, MD, MO, OH, PA, TN, VA ○ 11 ○
Sinella krekeleri Christiansen, 1960 Entomobryomorpha Troglobiont G3G4 KY, WV ○ 3 ○
Sinella tecta Christiansen & Bonet, 1980 Entomobryomorpha Troglobiont G1G2 CA ○ 2 ○
Sinella yosiia Bellinger & Christiansen, 1974 Entomobryomorpha Troglobiont G2G3 HI ○ 1 ○
Caecidotea stiladactyla Mackin & Hubricht, 1940 Slender-fingered Cave Isopod Isopoda Stygobiont G3G4 AR, MO, OK ○ 64 ○
Soldanellonyx chapuisi Walter, 1917 Actinedida Stygobiont G1 IN ○ ○ ○
Soldanellonyx morardi Walter, 1919 Actinedida Stygobiont G1 IN ○ ○ ○
Scoterpes sollmani Lewis, 2000 Sollman's Cave Millipede Chordeumatida Troglobiont G1 IN ○ ○ ○
Pseudanophthalmus potamaca Valentine, 1932 South Branch Valley Cave Beetle Coleoptera Troglobiont G3G4 VA, WV ○ ○ ○
Hesperochernes mirabilis (Banks, 1895) Southeastern Cave Pseudoscorpion Pseudoscorpiones Troglobiont G5 AL, GA, IN, KY, OH, TN, VA ○ 2 ○
Orconectes australis (Rhoades, 1941) Southern Cave Crayfish Decapoda Stygobiont G5 LC AL, TN ○ 158 152
Typhlichthys subterraneus Girard, 1859 Southern Cavefish Percopsiformes Stygobiont G4 NT AL, GA, KY, TN ○ 190 716
Eurycea braggi (Smith, 1968) Southern Grotto Salamander Caudata Troglobiont GNR LC AR ○ 15 137
Caecidotea recurvata (Steeves, 1963) Southwest Virginia Cave Isopod Isopoda Stygobiont G5 TN, VA ○ 120 76
Stygobromus mackini Hubricht, 1943 Southwestern Virginia Cave Amphipod Amphipoda Stygobiont G5 IN, KY, TN, VA, WV ○ 152 5
Spelaedrilus multiporus Cook 1975 Lumbriculida Stygobiont G1 VA ○ 2 ○
Spelaeorchestia pahoehoe Myers & Lowry, 2023 Amphipoda Troglobiont HI ○ ○ ○
Speleobama vana Park, 1951 Coleoptera Troglobiont G1 AL ○ ○ ○
Speleochus blanchardensis Carlton, 2012 Coleoptera Troglobiont AR ○ 1 ○
Speleochus croceus Park, 1960 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Speleochus eurous (Park, 1960) Coleoptera Troglobiont G2G3 AL ○ ○ ○
Speleochus ferus Park, 1951 Coleoptera Troglobiont G1G2 AL, TN ○ 1 ○
Speleochus macosar Carlton, 2012 Coleoptera Troglobiont AR ○ 1 ○
Speleochus steevesi (Park, 1960) Coleoptera Troglobiont G4 AL, TN ○ ○ ○
Speleochus stygicus Park, 1951 Coleoptera Troglobiont G1G2 AL ○ ○ ○
Speleochus synstygicus Park, 1956 Coleoptera Troglobiont G1 AL ○ ○ ○
Speleomaster lexi Briggs, 1974 Opiliones Troglobiont G1G2 ID ○ 2 6
Speleomaster pecki Briggs, 1974 Opiliones Troglobiont G1G2 ID ○ 1 ○
Speleonura kenchristianseni Palacios-Vargas & Benito, 2007 Poduromorpha Troglobiont G1 VA ○ ○ ○
Speleonychia sengeri Briggs, 1974 Opiliones Troglobiont G1G2 WA ○ 27 5
Speocirolana hardeni Bowman, 1992 Isopoda Stygobiont G2G3 TX ○ 116 4
Speodesmus bicornourus Causey, 1959 Polydesmida Troglobiont G2G3 TX ○ 252 ○
Speodesmus castellanus Elliott, 2004 Polydesmida Troglobiont TX ○ 115 ○
Speodesmus echinourus Loomis, 1939 Polydesmida Troglobiont G2G3 TX ○ 366 ○
Speorthus reyesi Shear & Reddell, 2017 Polydesmida Troglobiont TX ○ ○ ○
Speorthus tuganbius Chamberlin, 1952 Polydesmida Troglobiont G2G3 NM, TX ○ ○ ○
Speoseya grahami Causey, 1963 Chordeumatida Troglobiont G2G3 CA ○ 1 ○
Sphaeromicola moria Hart, 1978 Podocopida Stygobiont G1G2 TX ○ ○ ○
Sphalloplana hypogea Kenk, 1984 Tricladida Stygobiont G1G2 VA ○ 2 ○
Sphalloplana mohri Hyman, 1938 Tricladida Stygobiont G3G4 TX ○ 12 ○
Sphalloplana percoeca (Packard, 1879) Tricladida Stygobiont G5 AL, KY, TN, WV ○ 25 ○
Phreatodrobia spica Perez & Alvear, 2020 Spiky Cavesnail Littorinimorpha Stygobiont GNR TX ○ 7 1
Stygobromus urospinatus Wang & Holsinger, 2001 Spinose Cave Amphipod Amphipoda Stygobiont G1G3 CO ○ 6 ○
Stygobromus spinatus (Holsinger, 1967) Spiny Cave Amphipod Amphipoda Stygobiont G2G3 VU WV ○ 92 1
Pseudosinella spinosa (Delamare Deboutteville, 1949) Spiny Cave Springtail Entomobryomorpha Troglobiont G5 AL, GA, TN ○ 5 ○
Pseudanophthalmus punctatus Valentine, 1931 Spotted Cave Beetle Coleoptera Troglobiont G2 VA ○ ○ ○
Palaemonetes cummingi Chace, 1954 Squirrel Chimney Cave Shrimp Decapoda Stygobiont GH CR ESA FL ○ ○ ○
Pseudanophthalmus steevesi Barr, 1981 Steeves' Cave Beetle Coleoptera Troglobiont G1G2 AL ○ ○ ○
Caecidotea steevesi (Fleming, 1972) Steeves' Cave Isopod Isopoda Stygobiont G3G4 AR, KS, MO, OK ○ 10 ○
Tyrannochthonius steevesi Muchmore, 1996 Steeves' Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Stygobromus stellmacki (Holsinger, 1967) Stellmack's Cave Amphipod Amphipoda Stygobiont G1G2 VU PA ○ 6 ○
Stenophilus californicus (Chamberlin, 1930) Geophilomorpha Troglobiont G1G2 CA ○ ○ ○
Pseudanophthalmus tenebrosus Krekeler, 1973 Stevens Creek Cave Beetle Coleoptera Troglobiont G1 KY ○ 2 ○
Apochthonius typhlus Muchmore, 1967 Stone County Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 MO ○ ○ ○
Pseudanophthalmus quadratus Barr, 1965 Straley's Cave Beetle Coleoptera Troglobiont G1 VA ○ 1 ○
Striaria eldora Chamberlin, 1953 Chordeumatida Troglobiont G2G3 CA ○ ○ ○
Texapyrgus longleyi Thompson & Hershler, 1991 Striated hydrobe Littorinimorpha Stygobiont G1 TX ○ 9 1
Caecidotea stygia Packard, 1871 Stygian Cave Isopod Isopoda Stygobiont G5 IL, IN, KY, MO, OH, TN ○ 190 111
Amnicola stygius Hubricht, 1971 Stygian cavesnail Littorinimorpha Stygobiont G1 VU MO ○ 6 ○
Stygobromus ackerlyi Holsinger, 1978 Amphipoda Stygobiont G1G2 GA ○ 7 ○
Stygobromus allegheniensis (Holsinger, 1967) Amphipoda Stygobiont G5 MD, NY, PA, WV ○ 72 10
Stygobromus anacostensis Cannizzaro et al., 2024 Amphipoda Stygobiont DC ○ ○ 75
Stygobromus bakeri Gibson & Hutchins, 2021 Amphipoda Stygobiont TX ○ 27 ○
Stygobromus barryi Holsinger, 1978 Amphipoda Stygobiont G1 TN ○ ○ ○
Stygobromus cherylae Wang & Holsinger, 2001 Amphipoda Stygobiont G1 CA ○ 5 ○
Stygobromus duplus Wang & Holsinger, 2001 Amphipoda Stygobiont G1G2 WA ○ 2 ○
Stygobromus elatus (Holsinger, 1967) Amphipoda Stygobiont G1 VU AR ○ 3 ○
Stygobromus fecundus Holsinger, 1978 Amphipoda Stygobiont G1G2 TN ○ 6 ○
Stygobromus fontinalis Wang & Holsinger, 2001 Amphipoda Stygobiont G1G2 CO ○ 5 ○
Stygobromus gracilipes (Holsinger, 1967) Amphipoda Stygobiont G3G4 MD, PA, VA, WV ○ 39 8
Stygobromus grandis Holsinger, 1978 Amphipoda Stygobiont G1 GA ○ 3 ○
Stygobromus inexpectatus Holsinger, 1978 Amphipoda Stygobiont G1 AL ○ 2 ○
Stygobromus latus Wang & Holsinger, 2001 Amphipoda Stygobiont G1G3 WA ○ 4 ○
Stygobromus minutus Holsinger, 1978 Amphipoda Stygobiont G2G3 GA ○ 3 ○
Stygobromus montanensis Holsinger, 1974 Amphipoda Stygobiont G1G2 MT ○ 3 ○
Stygobromus nanus Holsinger, 1978 Amphipoda Stygobiont G1 WV ○ 3 ○
Stygobromus obscurus Holsinger, 1974 Amphipoda Stygobiont G1G2 MT ○ 3 ○
Stygobromus oregonensis Holsinger, 1974 Amphipoda Stygobiont G1G2 OR ○ 3 ○
Stygobromus puteanus Holsinger, 1974 Amphipoda Stygobiont G1G2 MT ○ 2 ○
Stygobromus reddelli (Holsinger, 1966) Amphipoda Stygobiont G1G2 VU TX ○ 6 ○
Stygobromus secundus Bousfield & Holsinger, 1981 Amphipoda Stygobiont G1 AB ○ 4 ○
Stygobromus tahoensis Holsinger, 1974 Amphipoda Stygobiont G1 CA, NV ○ 16 ○
Stygobromus tenuis (Smith, 1874) Amphipoda Stygobiont G4 CT, DC, MA, MD, NY, PA, VA, WV ○ 292 102
Stygonyx courtneyi Bousfield & Holsinger, 1989 Amphipoda Stygobiont G1G2 OR ○ 13 ○
Stygoporus oregonensis Larson & LaBonte, 1994 Coleoptera Stygobiont G1G2 OR ○ 4 12
Stylodrilus beattiei Cook, 1975 Lumbriculida Stygobiont G1 VA, WV ○ 5 ○
Stylodrilus californianus Rodriguez, 1996 Lumbriculida Stygobiont CA ○ 10 ○
Stylodrilus wahkeenensis Rodriguez & Coates, 1996 Lumbriculida Stygobiont AL, OR, TN ○ 6 ○
Phanetta subterranea (Emerton, 1875) Subterranean Sheetweb Spider Araneae Troglobiont G5 AL, AR, GA, IL, IN, KY, MO, NC, OH, TN, VA, WV ○ 9 47
Stygobromus subtilis (Hubricht, 1943) Subtle Cave Amphipod Amphipoda Stygobiont G2 VU IL, MI, MO ○ 17 ○
Crangonyx sulphurium Sawicki & Holsinger, 2017 Sulphurous Cave Amphipod Amphipoda Stygobiont G1 FL ○ 6 ○
Pseudanophthalmus inexpectatus Barr, 1959 Surprising Cave Beetle Coleoptera Troglobiont G1G2 KY ○ 5 ○
Pseudanophthalmus orientalis Krekeler, 1973 Surprising Cave Beetle Coleoptera Troglobiont G2G3 KY ○ ○ ○
Cambarus speleocoopi Buhay & Crandall, 2009 Sweet Home Alabama Cave Crayfish Decapoda Stygobiont G1 EN AL ○ 6 12
Stygobromus borealis Hoslinger, 1978 Taconic Cave Amphipod Amphipoda Stygobiont G2G3 MA, NY, QC, VT ○ 7 ○
Taiyutyla loftinae Shear & Krejca, 2011 Chordeumatida Troglobiont CA ○ ○ ○
Pseudanophthalmus fastigatus Barr, 1981 Tapered Cave Beetle Coleoptera Troglobiont G1 GA ○ ○ ○
Taracus aspenae Shear, 2018 Opiliones Troglobiont OR ○ 1 ○
Taracus fluvipileus Shear & Warfel, 2016 Opiliones Troglobiont CA ○ 1 ○
Taracus marchingtoni Shear & Warfel, 2016 Opiliones Troglobiont OR ○ ○ ○
Taracus silvestrii Briggs, 1969 Opiliones Troglobiont G1G2 OR ○ 1 ○
Taracus taylori Shear & Warfel, 2016 Opiliones Troglobiont GNR NV ○ 6 ○
Tartarocreagris altimana Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris amblyopa Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris attenuata Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris domina Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris grubbsi Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris hoodensis Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ 3 ○
Tartarocreagris infernalis (Muchmore, 1969) Pseudoscorpiones Troglobiont G2G3 TX ○ 5 ○
Tartarocreagris intermedia Muchmore, 1992 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris proserpina Muchmore, 2001 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Tartarocreagris reyesi Muchmore, 2001 Pseudoscorpiones Troglobiont GNR TX ○ ○ ○
Pseudanophthalmus parvus Krekeler, 1973 Tatum Cave Beetle Coleoptera Troglobiont GH KY ○ 2 ○
Tayshaneta anopica (Gertsch, 1974) Araneae Troglobiont G1G2 TX ○ 11 ○
Tayshaneta archambaulti Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 19 ○
Tayshaneta bullis (Cokendolpher, 2004) Araneae Troglobiont GNR TX ○ 12 ○
Tayshaneta concinna (Gertsch, 1974) Araneae Troglobiont G1G2 TX ○ 15 ○
Tayshaneta fawcetti Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 41 ○
Tayshaneta oconnorae Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 1 ○
Tayshaneta sandersi Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 9 ○
Tayshaneta sprousei Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 14 ○
Tayshaneta vidrio Ledford et al., 2012 Araneae Troglobiont GNR TX ○ ○ ○
Tayshaneta whitei Ledford et al., 2012 Araneae Troglobiont GNR TX ○ 12 ○
Pseudanophthalmus templetoni Valentine, 1948 Templeton's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Orconectes incomptus Hobbs & Barr, 1972 Tennessee Cave Crayfish Decapoda Stygobiont G2 VU TN ○ 8 17
Tetracion tennesseensis Causey, 1959 Tennessee Cave Millipede Callipodida Troglobiont G2G3 TN ○ ○ 36
Gyrinophilus palleucus McCrady, 1954 Tennessee Cave Salamander Caudata Stygobiont G2G3 NT AL, GA, TN ○ 206 67
Nesticus tennesseensis (Petrunkevitch, 1925) Tennessee Cave Spider Araneae Troglobiont G3G4 TN, VA, WV ○ 10 25
Antrorbis tennesseensis Perez, Shoobs, Gladstone, & Niemiller, 2019 Tennessee Cavesnail Littorinimorpha Stygobiont GNR TN ○ 3 6
Caecidotea richardsonae Hay, 1901 Tennessee Valley Cave Isopod Isopoda Stygobiont G5 AL, GA, TN, VA ○ 59 ○
Tethysbaena texana (Maguire, 1965) Thermosbaenacea Stygobiont GNR TX ○ 112 ○
Texanobathynella aaronswinki Camacho & Hutchins, 2021 Bathynellacea Stygobiont TX ○ ○ ○
Texanobathynella bowmani (Delamare et al., 1975) Bathynellacea Stygobiont GNR TX ○ ○ ○
Texanobathynella coloradoensis Camacho & Hutchins, 2021 Bathynellacea Stygobiont TX ○ ○ ○
Texanobathynella sachi Cho, 1996 Bathynellacea Stygobiont CA ○ 3 ○
Eurycea rathbuni (Stejneger, 1896) Texas Blind Salamander Caudata Stygobiont G1 CR ESA TX ○ 380 9
Lirceolus smithii (Ulrich, 1902) Texas Troglobitic Water Slater Isopoda Stygobiont G1G2 TX ○ 432 ○
Texella elliotti Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 1 ○
Texella hardeni Ubick & Briggs, 1992 Opiliones Troglobiont G1G2 TX ○ 5 ○
Texella hilgerensis Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 2 ○
Texella martensi Ubick, 2021 Opiliones Troglobiont CA ○ 1 ○
Texella mulaiki Goodnight & Goodnight, 1942 Opiliones Troglobiont G2G3 TX ○ 26 ○
Texella tuberculata Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 4 ○
Texella welbourni Ubick & Briggs, 1992 Opiliones Troglobiont G1G2 NM ○ 1 ○
Texella whitei Ubick & Briggs, 2004 Opiliones Troglobiont TX ○ 3 ○
Texicerberus amistad Hutchins et al., 2022 Isopoda Stygobiont GNR TX ○ 23 ○
Texicerberus castilloi Hutchins et al., 2022 Isopoda Stygobiont TX ○ 6 ○
Texicerberus schotteae Hutchins et al., 2022 Isopoda Stygobiont GNR TX ○ 22 ○
Texiweckelia texensis (Holsinger, 1973) Amphipoda Stygobiont G2G3 TX ○ 268 ○
Texiweckeliopsis insolita (Holsinger, 1980) Amphipoda Stygobiont G3 TX ○ 303 ○
Texoreddellia aquilonalis Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 8
Texoreddellia capitesquameo Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 1
Texoreddellia coahuilensis Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 5
Texoreddellia media Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ 17
Texoreddellia occasus Espinasa & Giribet, 2009 Zygentoma Troglobiont TX ○ ○ ○
Texoreddellia texensis (Ulrich, 1902) Zygentoma Troglobiont G4 TX ○ ○ 17
Thalkethops grallatrix Crabill, 1960 Scolopendromorpha Troglobiont G1G2 NM ○ 2 ○
Holsingeria unthanksensis Hershler, 1989 Thankless ghostsnail Littorinimorpha Stygobiont G2 VA ○ 20 1
Theatops phanus Chamberlin, 1951 Scolopendromorpha Troglobiont G2G3 TX ○ 30 ○
Theotima makua Gertsch, 1973 Araneae Troglobiont G2G3 HI ○ 1 ○
Pseudanophthalmus thomasi Barr, 1981 Thomas' Cave Beetle Coleoptera Troglobiont G1G2 VA ○ ○ ○
Thymoites minero Roth, 1992 Araneae Troglobiont G1G2 AZ ○ 1 ○
Stygobromus indentatus (Holsinger, 1967) Tidewater Amphipod Amphipoda Stygobiont G3 VU MD, NC, VA ○ 22 3
Stygobromus araeus (Holsinger, 1969) Tidewater Interstitial Amphipod Amphipoda Stygobiont G3 VU NC, VA ○ 24 ○
Pseudanophthalmus hadenoecus Barr, 1965 Timber Ridge Cave Beetle Coleoptera Troglobiont G1 WV ○ 1 ○
Tingupa pallida Loomis, 1939 Tingupa Cave Millipede Chordeumatida Troglobiont G4 AR, IL, MO ○ 13 ○
Pseudanophthalmus pusillus Barr, 1981 Tiny Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ ○
Pseudotremia conservata Hoffman & Lewis, 1997 TNC Cave Milliped Chordeumatida Troglobiont G1G2 IN ○ ○ ○
Rhadine persephone Barr, 1974 Tooth Cave Beetle Coleoptera Troglobiont G1G2 NE ESA TX ○ 90 9
Tartarocreagris texana (Muchmore, 1969) Tooth Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1G2 DD ESA TX ○ 1 ○
Tayshaneta myopica (Gertsch, 1974) Tooth Cave Spider Araneae Troglobiont G1G2 DD ESA TX ○ 74 ○
Trogloglanis pattersoni Eigenmann, 1919 Toothless Blindcat Siluriformes Stygobiont G1G2 VU ESA? TX ○ 104 4
Ptomaphagus torodei Peck, 1984 Torode's Cave Fungus Beetle Coleoptera Troglobiont G1G2 AL ○ ○ 6
Tyrannochthonius torodei Muchmore, 1996 Torode's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Traegaardhia holsingeri (Zacharda, 1980) Actinedida Troglobiont AR, KY, MO, OK ○ ○ ○
Traegaardhia paralleloseta (Zacharda, 1985) Actinedida Troglobiont G1 VA ○ ○ ○
Traegaardhia similis Zacharda et al., 2010 Actinedida Troglobiont AR ○ 12 ○
Trichodrilus comalensis Fend, 2023 Lumbriculida Stygobiont TX ○ 6 ○
Trichodrilus culveri Cook, 1975 Lumbriculida Stygobiont G1 WV ○ 2 ○
Trichopetalum uncum Cook & Collins, 1895 Chordeumatida Troglobiont GNR IN ○ ○ ○
Stygobromus trinus Wang & Holsinger, 2001 Trinity County Cave Amphipod Amphipoda Stygobiont G1 CA ○ 2 ○
Tuberochernes aalbui Muchmore, 1997 Pseudoscorpiones Troglobiont G1G2 CA ○ ○ ○
Tuberochernes cohni Harvey & Wynne, 2014 Pseudoscorpiones Troglobiont GNR AZ ○ ○ ○
Tuberochernes ubicki Muchmore, 1997 Pseudoscorpiones Troglobiont G1 AZ ○ 1 ○
Tuberona leonidasi (Kulkoyluoglu et al., 2023) Podocopida Stygobiont TX ○ ○ ○
Antrobia culveri Hubricht, 1971 Tumbling Creek cavesnail Littorinimorpha Stygobiont G1 EN ESA MO ○ 7 ○
Banksula tuolumne Briggs, 1974 Tuolumne Cave Harvestman Opiliones Troglobiont G2 CA ○ 1 ○
Pseudotremia contorta Shear, 2011 Twisted-gonopod Cave Milliped Chordeumatida Troglobiont G1 VA ○ ○ ○
Caecidotea bicrenata (Steeves, 1963) Two-toothed Cave Isopod Isopoda Stygobiont G5 AL, IL, KY, TN ○ 66 28
Tychobythinus hubrichti (Park, 1960) Coleoptera Troglobiont G1G2 KY ○ 1 ○
Tychobythinus jonesi (Park, 1951) Coleoptera Troglobiont G1G2 AL ○ ○ ○
Tychobythinus strinatii Besuchet, 1982 Coleoptera Troglobiont G1G2 TN ○ ○ ○
Tylparua troglodytes Evenhuis, 2024 Diptera Troglobiont HI ○ ○ ○
Typhloelmis caroline Barr et al., 2015 Coleoptera Stygobiont GNR TX ○ 3 ○
Typhloelmis finegan Barr et al., 2015 Coleoptera Stygobiont GNR TX ○ 3 ○
Typhloelmis sanfelipe Barr et al., 2015 Coleoptera Stygobiont GNR TX ○ 3 ○
Typhloelmis spangleri Barr, 2023 Coleoptera Stygobiont GNR TX ○ ○ ○
Typhlogastrura alabamensis (Thibaud, 1975) Poduromorpha Troglobiont G1G2 AL ○ ○ ○
Typhlogastrura christianseni Thibaud, 1975 Poduromorpha Troglobiont G1 AL, TN ○ ○ ○
Typhlogastrura hubbardi Thibaud, 1995 Poduromorpha Troglobiont G2G3 VA ○ ○ ○
Tyrannochthonius aladdinensis Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius aralu Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius archeri Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius attenuatus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius avernicolus Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius binoculatus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius chamberlini Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius charon Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius diabolus Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius erebicus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius felix Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius fiskei Muchmore, 1996 Pseudoscorpiones Troglobiont G1 TN ○ ○ ○
Tyrannochthonius gnomus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius halopotamus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL, TN ○ ○ ○
Tyrannochthonius hypogeus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 KY ○ ○ ○
Tyrannochthonius infernalis Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius muchmoreorum Cokendolpher, 2009 Pseudoscorpiones Troglobiont GNR TX ○ ○ ○
Tyrannochthonius nergal Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius orpheus Muchmore, 1996 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius osiris Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius parvus Chamberlin, 1995 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius pholeter Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius pluto Chamberlin, 1995 Pseudoscorpiones Troglobiont G1G2 AL ○ ○ ○
Tyrannochthonius pupukeanus Muchmore, 1983 Pseudoscorpiones Troglobiont G1 HI ○ 1 ○
Tyrannochthonius satan Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius skeletonis Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius stonei Muchmore, 1989 Pseudoscorpiones Troglobiont HI ○ 1 ○
Tyrannochthonius stygius Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius tartarus Muchmore, 1996 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius tenuis Chamberlin, 1995 Pseudoscorpiones Troglobiont G1 AL ○ ○ ○
Tyrannochthonius troglodytes Muchmore, 1986 Pseudoscorpiones Troglobiont G1G2 TX ○ ○ ○
Ufocandona hannaleeae Kulkoyluoglu et al., 2017 Podocopida Stygobiont TX ○ ○ ○
Uncinocythere ambophora (Walton & Hobbs, 1959) Podocopida Stygobiont G2G3 FL ○ 10 ○
Uncinocythere pholetera (Hart & Hobbs, 1961) Podocopida Stygobiont G1G2 MO ○ 14 ○
Uncinocythere warreni Hobbs & Walton, 1968 Podocopida Stygobiont G1 GA ○ 6 ○
Uncinocythere xania (Hart & Hobbs, 1961) Podocopida Stygobiont G3 IN, MO ○ 21 ○
Pseudanophthalmus unionis Barr, 1981 Union County Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Uroctonus grahami Gertsch & Soleglad, 1972 Scorpiones Troglobiont G1G2 CA ○ ○ 1
Usofila gracilis Keyserling, 1891 Araneae Troglobiont G1G2 CA ○ ○ ○
Stygobromus utahensis Wang & Holsinger, 2001 Utah Cave Amphipod Amphipoda Stygobiont G1G2 UT ○ 2 ○
Pseudanophthalmus valentinei Jeannel, 1949 Valentine's Cave Beetle Coleoptera Troglobiont G3G4 TN ○ ○ ○
Ptomaphagus valentinei Jeannel, 1933 Valentine's Cave Fungus Beetle Coleoptera Troglobiont G3G4 AL ○ 3 ○
Lissocreagris valentinei (Chamberlin, 1962) Valentine's Cave Pseudoscorpion Pseudoscorpiones Troglobiont G1 VA ○ ○ ○
Typhlogastrura valentini Thibaud, 1996 Valentine's Cave Springtail Poduromorpha Troglobiont G1 VA ○ ○ ○
Pseudobaicalasellus vandeli (Bresson, 1955) Vandel's Cave Isopod Isopoda Stygobiont G3G4 MD, VA ○ ○ ○
Vetericaris chaceorum Kensley & Williams, 1986 Decapoda Stygobiont G1 ESA HI ○ 4 ○
Pseudanophthalmus vicarius Barr, 1965 Vicariant Cave Beetle Coleoptera Troglobiont G2G3 VA ○ 1 ○
Fontigens morrisoni Hershler, Holsinger & Hubricht, 1990 Virginia springsnail Littorinimorpha Stygobiont G2 VA ○ 8 ○
Vitropyrgus lillianae Perez & Castañeda, 2023 Littorinimorpha Stygobiont TX ○ ○ 6
Thaumatogryllus cavicola Gurney & Rentz, 1978 Vovanoes Cave Cricket Orthoptera Troglobiont G1G2 VU HI ○ 4 ○
Vulcanochthonius aa Muchmore, 2000 Pseudoscorpiones Troglobiont HI ○ 1 ○
Vulcanochthonius howarthi (Muchmore, 1979) Pseudoscorpiones Troglobiont G1G2 HI ○ 1 ○
Vulcanochthonius pohakuloae Muchmore, 2000 Pseudoscorpiones Troglobiont HI ○ 1 ○
Stygobromus wahkeenensis Wang & Holsinger, 2001 Wahkeena Creek Groundwater Amphipod Amphipoda Stygobiont G1G2 OR ○ 3 ○
Pseudanophthalmus wallacei Barr, 1981 Wallace's Cave Beetle Coleoptera Troglobiont G1 TN ○ ○ 2
Pseudotremia wallaceae Lewis, 2005 Wallace's Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Stygobromus wardi Wang & Holsinger, 2001 Ward's Groundwater Amphipod Amphipoda Stygobiont G1G2 CO ○ 3 ○
Cicurina wartoni Gertsch, 1992 Warton Cave Meshweaver Araneae Troglobiont G1 TX ○ ○ ○
Sphalloplana weingartneri Kenk, 1970 Weingartner's cave flatworm Tricladida Stygobiont G4 IN ○ 3 ○
Stygobromus wengerorum Holsinger, 1974 Wenger Cave Amphipod Amphipoda Stygobiont G1 VU CA ○ 4 ○
Zygonopus krekeleri Causey, 1960 West Virginia Blind Cave Millipede Chordeumatida Troglobiont G4 WV ○ ○ ○
Gyrinophilus subterraneus Besharse & Holsinger, 1977 West Virginia Spring Salamander Caudata Stygobiont G1 CR ESA? WV ○ 30 45
Pseudanophthalmus assimilis Barr, 1981 West Wills Valley Cave Beetle Coleoptera Troglobiont G1G2 AL ○ ○ ○
Pseudanophthalmus occidentalis Barr, 1959 Western Cave Beetle Coleoptera Troglobiont G1G2 TN ○ ○ ○
Eurycea spelaea (Stejneger, 1892) Western Grotto Salamander Caudata Troglobiont G4 LC AR, KS, MO, OK ○ 2.1k 236
Stygobromus albapinus Taylor & Holsinger, 2011 White Pine amphipod Amphipoda Stygobiont NV ○ 4 ○
Cambarus veitchorum Cooper & Cooper, 1997 White Spring Cave Crayfish Decapoda Stygobiont G1 CR AL ○ 5 ○
Bathyphantes weyeri (Emerton, 1875) Wide Shield Sheetweaver Araneae Troglobiont G4 AR, IN, KY, ON, PA, QC, VA, WI, WV ○ 9 ○
Satan eurystomus Hubbs & Bailey, 1947 Widemouth Blindcat Siluriformes Stygobiont G1G2 VU ESA? TX ○ 48 ○
Stygobromus saltuaris Wang & Holsinger, 2001 Williamette Natonal Forest Groundwater Amphipod Amphipoda Stygobiont G1G2 OR ○ 2 ○
Stygobromus putealis (Holmes, 1909) Wisconsin Well Amphipod Amphipoda Stygobiont G2G3 VU WI ○ 2 ○
Pseudotremia withersi Lewis, 2009 Withers' Cave Millipede Chordeumatida Troglobiont G1 TN ○ ○ ○
Procambarus orcinus Hobbs & Means, 1972 Woodville Karst Cave Crayfish Decapoda Stygobiont G1 EN FL ○ 25 6
Stygobromus tritus Holsinger, 1974 Worn Groundwater Amphipod Amphipoda Stygobiont G1 MT ○ 5 ○
Veigaia wyandottensis (Packard, 1888) Wyandotte Cave Mite Mesostigmata Troglobiont G1 IN ○ ○ ○
Ceratophysella lucifuga (Packard, 1888) Wyandotte Cave Springtail Poduromorpha Troglobiont GNR IN ○ 2 ○
Xenotrechus condei Barr & Krekeler, 1967 Coleoptera Troglobiont G1G2 MO ○ ○ ○
Xenotrechus denticollis Barr & Krekeler, 1967 Coleoptera Troglobiont G1G2 MO ○ 2 9
Diacyclops yeatmani Reid, 1988 Yeatman's Groundwater Copepod Cyclopoida Stygobiont G2G3 IL, IN, MO, OK, TN ○ 5 ○
Pseudanophthalmus youngi Krekeler, 1958 Young's Cave Beetle Coleoptera Troglobiont G3G4 IN ○ 3 ○
Zosteractis interminata Loomis, 1943 Zosteractis Cave Millipede Julida Troglobiont G1 IL, MO ○ 2 ○

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Species accounts, distributions, and conservation status for the subterranean fauna of North America and around the globe, with resources for the cave and karst research community.

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